During the Early–Late Cretaceous transition, marine ecosystems in Eurasia hosted a diverse set of large predatory reptiles that occupied various niches. However, most of our current knowledge of these animals is restricted to a small number of bonebed-like deposits. Little is known of the geographical and temporal extent of such associations. The middle Albian – middle Cenomanian phosphorite-bearing succession exposed at Annopol, Poland produces numerous ichthyosaurian and plesiosaurian fossils. These are mostly isolated skeletal elements (e.g. teeth, vertebrae), but disarticulated partial skeletons and an articulated, subvertically embedded ichthyosaur skull are also available. The following taxa are identified: ‘Platypterygius’ sp., cf. Ophthalmosaurinae, Ichthyosauria indet., Polyptychodon interruptus, Pliosauridae indet., Elasmosauridae indet. and Plesiosauria indet. The large-sized ichthyosaur ‘Platypterygius’ and the pliosaurid Polyptychodon interruptus predominate within the upper Albian – middle Cenomanian deposits. The Annopol record, combined with data from England, France and western Russia, suggests that ‘Platypterygius’ and Polyptychodon interruptus formed a long-term, stable ecological sympatry in marine ecosystems of the European archipelago, at least during the Albian – middle Cenomanian. In addition, the marine reptile assemblage from Annopol is distinct from other Eurasian ecosystems in containing also elasmosaurids in its Albian portion.

Guest Editors Olivier Lacombe, Jonas Ruh, Dennis Brown and Faramarz Nilforoushan are to be congratulated for their efforts in putting together this exciting and comprehensive special issue on basement-involved fold-and-thrust belts. On behalf of Geological Magazine, I also thank the numerous authors and reviewers, whose efforts have made the special issue possible.

The South Dehshir Ophiolite, exposed along the southwestern margin of the Central Iranian microcontinent, comprises a mantle and a crustal sequence. This massif contains serpentinized peridotites, gabbros and diabase sheeted dykes cutting across wherlite, pegmatitic gabbro and plagiogranite, and is tectonically covered by radiolarites, pelagic limestones with Late Cretaceous microfauna and lavas. Under a microscope, the gabbros exhibit mesocumulate, adcumulate and hetradcumulate textures presumably formed in open magma chambers. Hypabyssal and lava samples show tholeiitic to calc-alkaline affinities, and their compositions cluster in the fields of arc or supra-subduction zone magmatism. Binary diagrams display compositional trends of magmatic differentiation, plotting away from the mid-ocean-ridge basalt (MORB) (tholeiitic) trend and following the arc (calc-alkaline) trend. Rare Earth element (REE) and spider diagrams display various enrichments at different levels. Moderate enrichment of the large-ion lithophile elements (LILE; Rb, Ba, Th, U, La, Sr) relative to the high-field-strength elements (HFSE; Nb, Ta) and slight depletion of Ti and Zr in some patterns may be attributed to variable influences of subduction components over the depleted mantle wedge. Elemental ratios display evidence for the contribution of both fluids and melt released from the subducted slab. Few patterns display ocean-island basalt (OIB) characteristics, possibly attributed to the involvement of local mantle plumes on the partial melting of the mantle wedge. This evidence indicates that the crustal rocks of the South Dehshir Ophiolite formed in a supra-subduction zone.

Trilobites are widespread in Lower Devonian deposits of north Gondwana, and some have been collected from two known sections of the Saoura Valley in SW Algeria, from the ‘Chefar el Ahmar’ Formation. This formation is considered to be from late Emsian to Frasnian in age, but only the lower parts of this formation have yielded trilobites. Nevertheless, no detailed studies have focused on their biodiversity and their morphological variability. New occurrences of phacopids including Barrandeops chattertoni sp. nov., Geesops fabrei sp. nov., Austerops legrandi sp. nov. and Phacops boudjemaai sp. nov. are described from this area and comparisons are made with closely allied species. These new occurrences have been integrated into analyses of intra- and inter-specific variability and biodiversity.

This thematic volume arose from a desire to acknowledge the pioneering contributions to mass extinction studies made by Professor Tony Hallam over the past 50 years and includes papers from several theme sessions held at the annual meeting of the Geological Society of America held in Denver in October 2013 along with other solicited contributions. They provide both a breadth of temporal coverage and also reflect the broad spectrum of earth scientists currently involved in studying these ancient crises, ranging from palaeontologists to volcanologists to earth system modellers.

Defining the structural style of fold-and-thrust belts is an important step for understanding the factors that control their long- and short-term dynamics, for comprehending seismic hazard associated with them, and for assessing their economic potential. While the thin-skinned model (no basement involvement) has long been the driving methodology for cross section construction and restoration of foreland fold-and-thrust belts, a wealth of new geological and geophysical studies have shown that they are often thick-skinned, that is, basement-involved.

Detailed analysis of the distribution of archaeocyaths in five lower Cambrian sections in South China has resulted in the erection of four new archaeocyathan zones and one new archaeocyathan bed. Listed in order of ascending age, these are: the Dailycyathus xiuqiensis Zone; the Dictyocyathus shaanxiensis Zone; the Spirillicyathus duplex Range Zone; the Sibirecyathus meitanensis Range Zone; and the Archaeocyathus yanjiaoensis beds. These new subdivisions permit a correlation of lower Cambrian strata both within the area of the Yangtze Platform (South China) as well as between this region and Siberia, Australia, Western Europe and North America. Within the Yangtze Platform area, archaeocyaths of the Dailycyathus xiuqiensis and Dictyocyathus shaanxiensis zones co-occur with trilobites of the middle and upper Eoredlichia Trilobite Zone of the Qiongzhusian Stage. The Spirillicyathus duplex and the Sibirecyathus meitanensis Range Zone correspond to the Malungia, Yiliangella–Yunnanaspis and Drepanuroides Trilobite zones of the basal and middle Canglangpuan Stage, respectively. Finally, the Archaeocyathus yanjiaoensis beds belong to the Palaeolenus fengyangensis Trilobite Zone of the upper Canglangpuan Stage. Global correlations based on archaeocyath assemblage zones suggest that: (1) the middle–upper Qiongzhusian of South China correlates with the middle Atdabanian Stage of the Siberian Platform, the Warriootacyathus wilkawillinensis Archaeocyath Zone in South Australia, the middle Issendalenian Stage of Morocco and the lower Ovetian Stage in Spain; (2) the lowermost Canglangpuan Stage of South China correlates with the uppermost Atdabanian of the Siberian Platform, the Spirillicyathus tenuis – Jugalicyathus tardus archaeocyath zones in South Australia, the uppermost Issendalenian – lower Banian stages of Morocco, the middle Ovetian Stage of Spain and the middle Montezuman Stage of North America; (3) the middle Canglangpuan Stage approximates the lowermost Botoman Stage of the Siberian Platform, middle Banian Stage of Morocco, the uppermost Ovetian – lowermost Marianian stages of Spain and the upper Montezuman Stage of North America; (4) the uppermost Canglangpuan Stage is equivalent of the middle Toyonian Stage of the Siberian Platform, Archaeocyathus abacus beds in Australia, the middle Bilbilian Stage in Spain and the middle Dyeran Stage (Bolbolenellus euryparia Trilobite Zone) in North America.

In contrast to the numerous excellently preserved arthropods, vertebrates and plants from the Mesozoic lacustrine fossil lagerstätten of northeastern China, which have calcium phosphate or organic skeletons, the preservation of taxa with a calcareous skeleton is fairly poor. Here we investigate, using a scanning electron microscope and energy dispersive X-ray spectrometer, the preservational modes of bivalves from the Jurassic Daohugou Fossil Beds of Inner Mongolia and the Lower Cretaceous Yixian Formation of eastern Liaoning. The Jurassic bivalve Ferganoconcha sibirica is preserved as strongly compressed composite moulds which contain remains of the organic periostracum. In the Yixian Formation, the bivalves Sphaerium anderssoni and Arguniella ventricosa occur as compacted internal, external or composite moulds or are preserved with a silicified shell, and rarely with a shell consisting of iron hydroxides, which had replaced pyrite during late diagenesis/weathering. Silicification produced partly fabric-replacive microcrystalline quartz and partly void-filling megaquartz crystals after the carbonate shell had been dissolved. Films of authigenic aluminosilicate minerals, partly secondarily silicified, cover the exterior and interior shell surfaces. Occasionally, early diagenetic pyrite crystals, now oxidized to iron hydroxides, filled shell cavities forming internal moulds and rarely replaced the bivalve shell. The poor preservation of the bivalves reflects the environment and water chemistry of these lakes, which were heavily influenced by volcanic processes. Frequent ash deposition and decomposition of volcanic glass particles created acidic and alkaline lake and interstitial waters, which led to early diagenetic formation of authigenic aluminosilicate minerals, ferruginous internal moulds, dissolution of shell carbonate and silicification of shells.

Tony Hallam's contributions to mass extinction studies span more than 50 years and this thematic issue provides an opportunity to pay tribute to the many pioneering contributions he has made to this field. Early work (1961) on the Jurassic in Europe revealed a link, during the Toarcian Stage, between extinction and the spread of anoxic waters during transgression – the first time such a common leitmotif had been identified. He also identified substantial sea-level changes during other mass extinction intervals with either regression (end-Triassic) or early transgression (end-Permian) coinciding with the extinction phases. Hallam's (1981) study on bivalves was also the first to elevate the status of the end-Triassic crisis and place it amongst true mass extinctions, changing previous perceptions that it was a part of a protracted period of turnover, although debates on the duration of this crisis continue (Hallam, 2002). Conflicting views on the nature of recovery from mass extinctions have also developed, especially for the aftermath of the end-Permian mass extinction. These discussions can be traced to Hallam's seminal 1991 paper that noted the considerable delay in benthic recovery during Early Triassic time and attributed it to the persistence of the harmful, high-stress conditions responsible for the extinction itself. This idea now forms the cornerstone of one of the more favoured explanations for this ultra-low diversity interval.

Recent finds of tetrapod ichnites in the red-bed and volcaniclastic succession of the Iberian Pyrenean Basin permits an assessment of the faunal diversity and palaeoenvironment of a late early Permian setting. The tetrapod ichnoassemblage is inferred with the aid of photogrammetry and constituted by Batrachichnus salamandroides, Limnopus isp., cf. Amphisauropus (these three ichnotaxa present associated swimming traces, assigned to Characichnos), cf. Ichniotherium, Dromopus isp., cf. Varanopus, Hyloidichnus isp. and Dimetropus leisnerianus. These ichnotaxa suggest the presence of temnospondyls, seymouriamorphs, diadectomorphs, araeoscelids, captorhinids and synapsid pelycosaurs as potential trackmakers. These faunas correlate to the late early Permian. Two ichnoassociations correspond to two different palaeoenvironments that were permanently or occasionally aquatic (meandering fluvial systems and unconfined runoff surfaces, respectively). Ichnotaxa in the fluvial system is more diverse and abundant than in the runoff surfaces system. The Iberian Pyrenean ichnoassemblage reveals the faunistic connection and similarities among nearing basins (Spain, southern France and Morocco) differing from the Central European basins (i.e. German Tambach Formation). Based on the palaeogeography and the climate models of the early Permian, we suggest the correlation of ichnofaunal composition with different palaeoclimate biomes. This results in a diffuse boundary of Gondwana–Laurasia land masses, indicating no geographic barriers but a possible climate control on the faunal distribution. Further studies, integrating data from distant tracksites, should refine these biome boundaries.

The Fishguard Volcanic Group represents an excellently preserved example of a volcanic sequence linked to the closure of the Iapetus Ocean. This study re-examines the petrogenesis and proposed tectonic setting for the Llanvirn (467–458 Ma) Fishguard Volcanic Group, South Wales, UK. New major and trace element geochemical data and petrographic observations are used to re-evaluate the magma chamber processes, mantle melting and source region. The new data reveal that the Fishguard Volcanic Group represents a closely related series of basalts, basaltic andesites, dacites and rhyolites originating from a spinel lherzolite source which had been modified by subduction components. The rocks of the Fishguard Volcanic Group are co-genetic and the felsic members are related to the more primitive basalts mainly by low-pressure fractional crystallization. The geochemistry of the lavas was significantly influenced by subduction processes associated with a coeval arc, while significant amounts of assimilation of continental crust along with fractional crystallization appear to have contributed to the compositions of the most evolved lavas. The Fishguard Volcanic Group was erupted into a back-arc basin where extensive rifting but no true seafloor spreading had occurred.

Flysch-type, syn-orogenic deposits of Carboniferous age occur in relation to the emplacement of a large allochthonous nappe stack in the Variscan belt of NW Iberia. New U–Pb age populations of detrital zircons obtained using laser ablation inductively coupled plasma mass spectrometry (LA-ICP-MS) are considered together with others from previously dated samples to establish the relationships between sedimentation and thrusting. The age populations of four syn-orogenic formations are compared with those of the pre-orogenic sequence in the Autochthon and Parautochthon, representing the Gondwanan passive margin, and in the Allochthon, formed by peri-Gondwanan and oceanic terranes. In addition, a new structural study has been carried out to understand the relationships between the syn-orogenic deposits and the development of Variscan structures. The aims are to identify the sources of sediments and to establish the relationship between Variscan structural evolution and syn-orogenic sedimentation. Development of a forebulge outwards from the allochthonous front, deduced from the structural study, suggests the existence of depocentres that hosted the syn-orogenic sediments. Together with the trend shown by the more recent zircons in each formation, that are younger towards the external zones, the data suggest that sedimentation occurred in progressively migrating depocentres formed in front of the allochthonous wedge during its emplacement. The zircon age populations point to the Allochthon as the main source of detritus for the syn-orogenic basins, with perhaps a limited participation of the Parautochthon and Autochthon in the younger formations.

The links between the Siberian Traps and the end-Permian mass extinction, and between the North Atlantic igneous province (NAIP) and the Paleocene–Eocene thermal maximum (PETM), demonstrate a critical role for large igneous provinces (LIPs) in the disruption of the Earth-surface carbon cycle (ESCC). High-precision ages for both volcanic provinces and the associated environmental crises show that, in both cases, the crisis was contemporaneous with the volcanism. The NAIP comprises two phases: the earlier Phase 1 (c. 61 Ma) and the much more voluminous Phase 2 (c. 56 Ma), linked to the opening of the NE Atlantic. The latter triggered the PETM, the largest Cenozoic hyperthermal. The Siberian Traps are significantly more voluminous than the NAIP, and triggered the end-Permian mass extinction. The masses of volcanic CO2 emitted from these provinces may have been much greater than previously suggested as substantial gas may come from intrusive bodies deep within the crust. Precursory warming due to the accumulation of volcanic CO2 in the atmosphere likely triggered the release of low-δ13C methane hydrate, although the masses of methane hydrate alone may have been insufficient to account for the observed temperature rises. The organic C was likely strongly supplemented by magmatically derived carbon and thermogenic carbon released during emplacement of sills and dykes into C-rich sedimentary units. More data are required on the volcanic flux rates in order to refine the cause–effect relationships between LIPs and the ESCC.

A new plant of Late Devonian (Famennian) age, Melvillipteris quadriseriata gen. et sp. nov., is reported from Melville Island, Nunavut, Canadian Arctic Archipelago. Main axes and two orders of lateral branches are recognized. The main axes are monopodial in habit, with a zigzag and/or upright appearance and dense adventitious roots attached on some internodes. First-order branches are inserted on the main axes in distichous pairs, showing a quadriseriate pattern (i.e. alternate pairs); they bear one or two sterile ultimate appendages on the proximal portions, and then distally bear alternate second-order branches. Second-order branches bear alternate ultimate appendages which may be fertile or sterile; the sterile ultimate appendages are composed of one or two successive dichotomies, terminating in oppositely recurved tips, and fertile appendages show an elaborate branching system. Each fertile appendage has an initial dichotomy forming two sister branches, each of which further divides three or four times to produce ultimate divisions terminated by sporangia. Two to four small fusiform sporangia are grouped in a truss which is supplied by penultimate division of the fertile appendage. The sporangia have a pointed tip and a longitudinal dehiscence. This new plant resembles other plants with a quadriseriate branching pattern (e.g. Rhacophyton Crépin, Cephalopteris Nathorst, Protocephalopteris Ananiev, Ellesmeris Hill, Scheckler & Basinger and Protopteridophyton Li & Hsü), and is assigned to the Rhacophytales. Its similarity to and divergence from the rhacophytaleans, cladoxylopsids and other related plants are discussed.

Defining the structural style of fold–thrust belts and understanding the controlling factors are necessary steps towards prediction of their long-term and short-term dynamics, including seismic hazard, and to assess their potential in terms of hydrocarbon exploration. While the thin-skinned structural style has long been a fashionable view for outer parts of orogens worldwide, a wealth of new geological and geophysical studies has pointed out that a description in terms of thick-skinned deformation is, in many cases, more appropriate. This paper aims at providing a review of what we know about basement-involved shortening in foreland fold–thrust belts on the basis of the examination of selected Cenozoic orogens. After describing how structural interpretations of fold–thrust belts have evolved through time, this paper addresses how and the extent to which basement tectonics influence their geometry and their kinematics, and emphasizes the key control exerted by lithosphere rheology, including structural and thermal inheritance, and local/regional boundary conditions on the occurrence of thick-skinned tectonics in the outer parts of orogens.

The style of inversion of inherited extensional basins in the Western Alps is investigated through thermo-mechanical modelling. Two-dimensional models consist of a half-graben embedded in a relatively strong crust (basement) and filled with weak syn-rift sediments (cover). We investigate the relative influence of the internal friction (µ) of the basin-bounding normal fault, tectonic burial (h) under an overlying nappe and the geothermal gradient. We use a viscoplastic model with symmetrical shortening. The inherited normal fault is implemented as a curved thin body with a variable friction coefficient (µ) ranging from 0.1 to 0.6. The style of basin inversion is controlled at shallow depth by the internal friction coefficient, whose influence decreases with the increase of both burial depth and geothermal gradient. With increasing burial and/or geothermal gradient, fault reactivation is inhibited and distributed deformation in the basement induces the vertical extrusion of the cover. The basin inversion is accompanied by distributed deformation in the cover and by the shearing of the basin and basement interface. The results are consistent with the style of inversion of inherited half-grabens in the external Western Alps, where no significant fault reactivation occurred owing to tectonic burial underneath the Alpine internal units during the early Alpine collision.

A review of the palynofloral succession at the well-documented Triassic–Jurassic boundary sites – Kuhjoch (Austria), St Audrie's Bay (UK), Stenlille (Denmark), Astartekløft (Greenland), Sverdrup Basin (Arctic Canada), Northern Carnarvon Basin (Western Australia), Southeast Queensland (eastern Australia) and New Zealand – show all sites experienced major to moderate re-organization of the terrestrial vegetation during the end-Triassic event. The changes led to subsequent taxonomic losses of between 17% and 73% of the Rhaetian pre-extinction palynoflora. The majority of the typical Rhaetian taxa that disappear are so far not known from in situ occurrences in reproductive structures of macrofossil plant taxa. From an ecological perspective, the most dramatic changes occurred in the Sverdrup Basin, Stenlille, Kuhjoch and Carnarvon Basin, where the pre- and post-extinction palynofloras were fundamentally different in both composition and dominance. These changes correspond to ecological severity Category I of McGhee et al. (2004), while the remaining sites are placed in their Subcategory IIa because there the pre-extinction ecosystems are disrupted, but recover and are not replaced post-extinction. Increased total abundances of spores on both hemispheres during the extinction and recovery intervals may indicate that environmental and/or climatic conditions became less favourable for seed plants. Such conditions may include expected effects of volcanism in the Central Atlantic Magmatic Province, such as acid rain, terrestrial soil and freshwater acidification due to volcanic sulfur dioxide emissions, fluctuating ultraviolet flux due to ozone depletion caused by halogens and halocarbon compounds, and drastic changes in climatic conditions due to greenhouse gas emissions.

Diverse, abundant and usually well-preserved communities of skeletal organisms occur in the lower Pleistocene (Gelasian) siliciclastic deposits of the Greek island of Rhodes. Benthic foraminifers, molluscs and bryozoans have been studied in four measured and sampled sections located in the northern part of the island. Among these bottom-dwelling organisms, numerous extant taxa are good environmental indicators and, combined with field observations and sedimentological data, they provide information on the probable conditions in which they developed. The siliciclastic deposits of the Kritika Formation have been divided into 14 different bio- and lithofacies, which have been further grouped into four facies associations corresponding to four different environmental settings: (1) continental to fluviatile; (2) brackish-water (lagoonal/deltaic); (3) infralittoral (0–20 m); and (4) upper circalittoral (depths of 20–40 m, but also down to c. 50–60 m). Among the marine facies associations, several characteristic biocoenoses have been recognized: soft-bottoms (fine to coarse sands and gravels); seagrass meadows; biogenic calcareous crusts on drowned beachrock slabs; red algal rhodoliths; and bivalve shell beds. In the studied sections, 13 superimposed genetic sequences have been documented. The repetition of similar facies associations within each sequence suggests: (1) a possibly eustasy-controlled, cyclic sedimentation; (2) a general subsidence of Rhodes during the deposition of the studied facies associations; and (3) a mostly constant range of environmental conditions (i.e. sedimentation rates and temperature) throughout the Gelasian.

The sediments of organic-rich oil shales in the Bilong Co. area can be correlated with those of the early Toarcian anoxic black-shale events in Europe. The Bilong Co. sediments are rich in trace elements Se, Mo, Cd, As and Ni, and, to a lesser extent, Li, F, V, Co, Cu, Cs, Hg and Bi, in comparison to the upper continental crust. Thirty-two oil shale samples were collected from the Bilong Co. oil shale to evaluate the controlling factors of trace-element enrichment in the lower Toarcian anoxic sediments. Minerals identified in the Bilong Co. oil shale include calcite, quartz, illite, feldspar and dolomite, and trace amounts of siderite, magnesite, halite, haematite, zeolite, amphibole, gypsum, anhydrite, apatite, pyrite, sphalerite, barite and mixed-layer illite/smectite. Mineralogical and geochemical data show that seawater and hydrothermal activities are the dominant influences on the mineralogical composition and elevated trace-element concentrations in the oil shale. The clay minerals, quartz and feldspar in the Bilong Co. oil shale were derived from the Nadi Kangri volcanic rocks. Input of sediment from this source may have led to enrichment of trace elements Li, Cr and Cs in the oil shale. Carbonate minerals and nodular- and framboidal-pyrite are authigenic phases formed from seawater. The enrichment of V, Co, Ni, Cu, Mo, As, Se, Bi and U in the oil shale was owing to marine influence. Barite, sphalerite and fracture-filling pyrites were derived from hydrothermal solutions. High concentrations of F, Zn and Cd were probably derived from hydrothermal fluids.

A large number of naked, fossil spore tetrads assignable to the dispersed microspore genera Indotriradites, Microbaculispora and Microfoveolatispora are reported for the first time from an early Permian stratum (Lower Barakar Formation) of Singrauli Coalfield, Son-Mahanadi Basin, Central India. This is also the first record of tetrads from any Artinskian strata in the world. There is no evidence of any kind of sporangia or related plant parts in the present investigation that could ascertain the affinity of these tetrads; however, the presence of a trilete mark in the spores of the tetrads demonstrates their alliance at least with the pteridophyte group. The present study suggests possible factors affecting the sporogenesis process in the past, considering other available global records pertaining to fossil spore tetrads. The results of significant physiological and biochemical analyses performed on the anthers of modern plants related to reproductive biology, in order to understand the conditions and changes responsible for the formation of tetrads, are also considered. We analysed the globally occurring fossil tetrads and the palaeoclimates prevailing during their deposition. A correlation between extreme climatic conditions, specific pH values inside microsporangium and the formation of tetrad is envisaged. It is deduced that extreme climatic conditions (extreme cold/extreme hot) might have triggered some sort of malfunctioning in the sporogenesis process that altered the specific pH values inside the microsporangium. Any restraint of the activity of the callase enzyme, responsible for dissolution of callose walls laid between the individual spores, may therefore have apprehended the dissociation of tetrads into individual spores.