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Large predatory marine reptiles from the Albian–Cenomanian of Annopol, Poland

Published online by Cambridge University Press:  03 June 2015

NATHALIE BARDET
Affiliation:
Sorbonne Universités, CR2P CNRS-MNHN-UPMC Paris 6, Département Histoire de la Terre, Muséum National d'Histoire Naturelle, CP 38, 8 rue Buffon, 75005 Paris, France
VALENTIN FISCHER
Affiliation:
Earth Sciences Department, University of Oxford, South Parks Road, OX1 3AN, Oxford, UK Département de Géologie, Université de Liège, Place du 20-Août, 4000 Liège, Belgium OD Earth and History of Life, Royal Belgian Institute of Natural Sciences, Rue Vautier 29, 1000 Brussels, Belgium
MARCIN MACHALSKI*
Affiliation:
Institute of Paleobiology, Polish Academy of Sciences, ul. Twarda 51/55, 00–818 Warszawa, Poland
*
Author for correspondence: [email protected]

Abstract

During the Early–Late Cretaceous transition, marine ecosystems in Eurasia hosted a diverse set of large predatory reptiles that occupied various niches. However, most of our current knowledge of these animals is restricted to a small number of bonebed-like deposits. Little is known of the geographical and temporal extent of such associations. The middle Albian – middle Cenomanian phosphorite-bearing succession exposed at Annopol, Poland produces numerous ichthyosaurian and plesiosaurian fossils. These are mostly isolated skeletal elements (e.g. teeth, vertebrae), but disarticulated partial skeletons and an articulated, subvertically embedded ichthyosaur skull are also available. The following taxa are identified: ‘Platypterygius’ sp., cf. Ophthalmosaurinae, Ichthyosauria indet., Polyptychodon interruptus, Pliosauridae indet., Elasmosauridae indet. and Plesiosauria indet. The large-sized ichthyosaur ‘Platypterygius’ and the pliosaurid Polyptychodon interruptus predominate within the upper Albian – middle Cenomanian deposits. The Annopol record, combined with data from England, France and western Russia, suggests that ‘Platypterygius’ and Polyptychodon interruptus formed a long-term, stable ecological sympatry in marine ecosystems of the European archipelago, at least during the Albian – middle Cenomanian. In addition, the marine reptile assemblage from Annopol is distinct from other Eurasian ecosystems in containing also elasmosaurids in its Albian portion.

Type
Original Articles
Copyright
Copyright © Cambridge University Press 2015 

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References

IIIAlbright, B. L., Gillette, D. D. & Titus, A. L. 2007. Plesiosaurs from the Upper Cretaceous (Cenomanian-Turonian) Tropic Shale of southern Utah, Part 1: New records of the pliosaur Brachauchenius lucasi . Journal of Vertebrate Paleontology 27, 3140.Google Scholar
Amédro, F. 2002. Plaidoyer pour un étage Vraconnien entre l'Albien sensu stricto et le Cénomanien (système Crétacé). Academie Royale de Belgique, Classe des Sciences 4, 1128.Google Scholar
Arkhangelsky, M. S. 2001. On a new ichthyosaur of the genus Otschevia from the Volgian Stage of the Volga region near Ulyanovsk. Paleontological Journal 35, 629–34.Google Scholar
Bardet, N. 1989. Un crâne d'Ichthyopterygia dans le Cénomanien du Boulonnais. Mémoires de la Société Académique du Boulonnais 6, 131.Google Scholar
Bardet, N. 1992. Stratigraphic evidence for the extinction of the ichthyosaurs. Terra Nova 4, 649–56.Google Scholar
Bardet, N., Duffaud, S., Martin, M., Mazin, J.-M., Pereda Suberbiola, X. & Vidier, J.-P. 1997. Découverte de l'ichthyosaure Ophthalmosaurus dans le Tithonien (Jurassique supérieur) du Boulonnais, Nord de la France. Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 205, 339–54.Google Scholar
Bardet, N. & Godefroit, P. 1995. Plesiosaurus houzeaui Dollo, 1909 from the Upper Campanian of Ciply (Belgium) and a review of the Upper Cretaceous plesiosaurs from Europe. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre 65, 179–86.Google Scholar
Bardet, N., Godefroit, P. & Sciau, J. 1999. A new elasmosaurid plesiosaur from the Lower Jurassic of southern France. Palaeontology 42, 927–52.Google Scholar
Bardet, N., Mazin, J.-M., Azéma, C., Bégouen, V. & Masure, E. 1991. “L'ichthyosaure de Bedeille” (Ariège, France): examen palynologique de la gangue et mise au point stratigraphique. Bulletin de la Société Géologique de France 162, 897903.CrossRefGoogle Scholar
Bardet, N., Wellnhofer, P. & Herm, D. 1994. Discovery of ichthyosaur remains (Reptilia) in the upper Cenomanian of Bavaria. Mitteilungen aus der Bayerischen Staatssammlung für Paläontologie und Historische Geologie 34, 213–20.Google Scholar
Barrois, C. 1875. Les reptiles du terrain crétacé du nord-est du Bassin de Paris. Bulletin Scientifque, Historique et Littéraire du Nord 6, 11.Google Scholar
Baur, G. 1887. On the morphology and origin of the Ichthyopterygia. American Naturalist 21, 837–40.Google Scholar
Blainville, H. M. D., de. 1835. Description de quelques espèces de reptiles de la Californie, précédée de l'analyse d'un système général d'érpetologie et d'amphibiologie. Nouvelles Annales du Muséum d'Histoire Naturelle Paris 4, 233–96.Google Scholar
Buffetaut, E. 1977. Remarques préliminaires sur l'ichthyosaure de Saint-Jouin (76). Bulletin de la Société Géologique de Normandie et Amis du Muséum du Havre 64, 17–9.Google Scholar
Buffetaut, E., Colletė, C., Dubus, B. & Petit, J.-L. 2005. The “sauropod” from the Albian of Mesnil-Saint-Père (Aube, France): a pliosaur, not a dinosaur. Notebooks on Geology, Letter 2005/01, 5.Google Scholar
Buffetaut, E., Tomasson, R. & Tong, H. 2003. Restes fossiles de grands reptiles jurassiques et crétacés dans l'Aube (France). Bulletin d'Information des Géologues du Bassin de Paris 40, 3343.Google Scholar
Capellini, G. 1890. Ichthyosaurus campylodon e tronchi di cicadee nelle argille scagliose dell'Emilia. Memorie delle Reale Accademia delle Scienze di Bologna 4 (10), 431–50.Google Scholar
Cieśliński, S. 1959. The Albian and Cenomanian in the northern periphery of the Holy Cross Mountains (stratigraphy based on cephalopods). Prace Instytutu Geologicznego 28, 195 (in Polish, English summary).Google Scholar
Cieśliński, S. 1976. Development of the Danish-Polish furrow in the Góry Świętokrzyskie region in the Albian, Cenomanian and Lower Turonian. Biuletyn Instytutu Geologicznego 295, 249–71 (in Polish, English summary).Google Scholar
Cieśliński, S. 1987. Albian and Cenomanian inoceramids in Poland and their stratigraphic significance. Biuletyn Instytutu Geologicznego 354, 1162 (in Polish, English summary).Google Scholar
Cieśliński, S. & Milakovič, B. 1962. Kręgowce i flora kredowa z obrzeżenia Gór Świętokrzyskich. Biuletyn Instytutu Geologicznego 174, 245–66.Google Scholar
Colleté, C. 2010. Stratotype Albien. Paris: Muséum National d'Histoire Naturelle, Mèze: Biotope, Orléans: BRGM, 332 pp.Google Scholar
Cookson, I. C. & Hughes, N. F. 1964. Microplankton from the Cambridge Greensand (mid-Cretaceous). Palaeontology 7, 3759.Google Scholar
Cope, E. D. 1869. Synopsis of the Extinct Batrachia and Reptilia of North America, Part I. Transactions American Philadelphia Society New Series 14, 1235.Google Scholar
Debris, J.-P. 1977. Découverte d'un crâne d'ichthyosaurien dans l'Albien de Saint-Jouin. Bulletin de la Société Géologique de Normandie et Amis du Muséum du Havre 64, 13–6.Google Scholar
Debris, J.-P. 1978. Données nouvelles sur les Ichthyosauriens de l'Albien de Saint Jouin (76). Bulletin Trimestriel de la Société Géologique de Normandie et Amis du Muséum du Havre 65, 25–8.Google Scholar
Druckenmiller, P. S. & Russell, A. P. 2008. A phylogeny of Plesiosauria (Sauropterygia) and its bearing on the systematic status of Leptocleidus Andrews 1922. Zootaxa 1863, 1120.CrossRefGoogle Scholar
Eichwald, K. E. 1853. Einige paläontologische Bemerkungen über den Eisensand von Kursk. Bulletin de la Société Impériale des Naturalistes de Moscou 2, 209–31.Google Scholar
Eichwald, K. E. 1865–1868. Lethaea Rossica ou Paléontologie de la Russie. Second Volume. Période Moyenne. Stuttgart: E. Schweizbart (E. Koch), 1304 pp.Google Scholar
Fischer, V. 2012. New data on the ichthyosaur Platypterygius hercynicus and its implications for the validity of the genus. Acta Palaeontologica Polonica 57, 123–34.Google Scholar
Fischer, V., Arkhangelsky, M. S., Naish, D., Stenshin, I. M., Uspensky, G. N. & Godefroit, P. 2014a. Simbirskiasaurus and Pervushovisaurus reassessed: implications for the taxonomy and cranial osteology of Cretaceous platypterygiine ichthyosaurs. Zoological Journal of the Linnean Society 171, 822–41.Google Scholar
Fischer, V., Arkhangelsky, M. S., Uspensky, G. N., Stenshin, I. M. & Godefroit, P. 2014b. A new Lower Cretaceous ichthyosaur from Russia reveals skull shape conservatism within Ophthalmosaurinae. Geological Magazine 151, 6070.Google Scholar
Fischer, V., Bardet, N., Guiomar, M. & Godefroit, P. 2014c. High diversity in Cretaceous ichthyosaurs from Europe prior to their extinction. PLoS ONE 9, e84709.Google Scholar
Fischer, V., Clément, A., Guiomar, M. & Godefroit, P. 2011a. The first definite record of a Valanginian ichthyosaur and its implication for the evolution of post-Liassic Ichthyosauria. Cretaceous Research 32, 155–63.Google Scholar
Fischer, V., Maisch, M. W., Naish, D., Liston, J., Kosma, R., Joger, U., Krüger, F. J., Pardo-Pérez, J., Tainsh, J. & Appleby, R. M. 2012. New ophthalmosaurids from the Early Cretaceous of Europe demonstrate extensive ichthyosaur survival across the Jurassic–Cretaceous boundary. PLoS ONE 7, e29234.Google Scholar
Fischer, V., Masure, E., Arkhangelsky, M. S. & Godefroit, P. 2011b. A new Barremian (Early Cretaceous) ichthyosaur from western Russia. Journal of Vertebrate Paleontology 31, 1010–25.Google Scholar
Foote, A. D., Morin, P. A., Durban, J. W., Willerslev, E., Orlando, L. & Gilbert, M. T. P. 2011. Out of the Pacific and back again: insights into the matrilineal history of Pacific killer whale ecotypes. PLoS ONE 6, e24980.Google Scholar
Godefroit, P. 1993. Les grands ichthyosaures sinémuriens d'Arlon. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre 63, 2571.Google Scholar
Gray, J. E. 1825. A synopsis of the genera of reptiles and Amphibia, with a description of some new species. Annals of Philosophy 26, 193217.Google Scholar
Hancock, J. M. 1990. Sea-level changes in the British region during the Late Cretaceous. Proceedings of the Geologists' Association 100 (for 1989), 565–94.Google Scholar
Hopson, P. M. 2005. A stratigraphical framework for the Upper Cretaceous Chalk of England and Scotland with statements on the Chalk of Northern Ireland and the UK Offshore Sector. British Geological Survey Research Reports RR/05/01, 1102.Google Scholar
Hopson, P. M., Wilkinson, I. P. & Woods, M. A. 2008. A stratigraphical framework for the Lower Cretaceous of England. British Geological Survey Research Reports RR/08/03, 187.Google Scholar
Huene, F. von. 1922. Die Ichthyosaurier des Lias und ihre Zusammenhänge. Berlin: Verlag von Gebrüdern Borntraeger, 114 pp.Google Scholar
Juignet, P. 1980. Transgressions-régressions, variations eustatiques et influences tectoniques de l'Aptien au Maastrichtien dans le Bassin de Paris occidental et sur la Bordure du Massif Armoricain. Cretaceous Research 1, 341–57.Google Scholar
Kear, B. P. 2005. Cranial morphology of Platypterygius longmani Wade, 1990 (Reptilia: Ichthyosauria) from the Lower Cretaceous of Australia. Zoological Journal of the Linnean Society 145, 583622.Google Scholar
Kiprijanoff, W. 1881. Studien über die fossilen Reptilien Russlands. Theil 1. Gattung Ichthyosaurus König aus dem severischen Sandstein oder Osteolith der Kreide-Gruppe. Mémoires de l'Académie Impériale des Sciences de St.-Pétersbourg 28 (7), 1103.Google Scholar
Kiprijanoff, W. 1882. Studien über die fossilen Reptilien Russlands. Theil 2. Gattung Plesiosaurus Conybeare aus dem severischen Sandstein oder Osteolith der Kreidegruppe. Mémoires de l'Académie Impériale des Sciences de St.-Pétersbourg 30 (7), 155.Google Scholar
Kiprijanoff, W. 1883a. Studien über die fossilen Reptilien Russlands. Theil 3. Gruppe Thaumatosauria n. aus der Kreide-Formation und dem Moskauer Jura. Mémoires de l'Académie impériale des Sciences de St.-Pétersbourg 31 (7), 157.Google Scholar
Kiprijanoff, W. 1883b. Studien über die fossilen Reptilien Russlands. Theil 4. Ordung Crocodilia Oppel. Indeterminirte fossile Reptilien. Mémoires de l'Académie impériale des Sciences de St.-Pétersbourg 31 (7), 129.Google Scholar
Lambert, O., Bianucci, G., Post, K., de Muizon, C., Salas-Gismondi, R., Urbina, M. & Reumer, J. 2010. The giant bite of a new raptorial sperm whale from the Miocene epoch of Peru. Nature 466, 105–8.Google Scholar
Lydekker, R. 1889. Catalogue of the fossil Reptilia and Amphibia in British Museum (Natural History). Part II. Containing the orders Ichthyopterygia and Sauropterygia. London: Printed by Orders of the Trustees of the British Museum, 307 pp.Google Scholar
Machalski, M. & Kennedy, W. J. 2013. Oyster-bioimmured ammonites from the Upper Albian of Annopol, Poland: stratigraphic and palaeobiogeographic implications. Acta Geologica Polonica 63, 545–54.Google Scholar
Machalski, M., Komorowski, A. & Harasimiuk, M. 2009. New chances in the quest for Cretaceous marine vertebrates in abandoned phosphate mine at Annopol on Vistula River. Przegląd Geologiczny 57, 638–41 (in Polish, English summary).Google Scholar
Machalski, M. & Martill, D. M. 2013. First pterosaur remains from the Cretaceous of Poland. Annales Societatis Geologorum Poloniae 83, 99104.Google Scholar
Marcinowski, R. 1980. Cenomanian ammonites from German Democratic Republic, Poland, and the Soviet Union. Acta Geologica Polonica 30, 215325.Google Scholar
Marcinowski, R. & Radwański, A. 1983. The mid-Cretaceous transgression onto the Central Polish Uplands (marginal part of the Central European Basin). Zitteliana 10, 6596.Google Scholar
Marcinowski, R. & Radwański, A. 1989. Stratigraphic approach to the mid-Cretaceous transgressive sequence of the Central Polish Uplands. Cretaceous Research 10, 153–72.Google Scholar
Marcinowski, R. & Walaszczyk, I. 1985. Middle Cretaceous deposits and biostratigraphy of the Annopol section, Central Polish Uplands. Österreichische Akademie der Wissenschaften, Schriftenreiche der Erdwissenschaftlichen Kommissionen 7, 2741.Google Scholar
Marcinowski, R. & Wiedmann, J. 1985. The Albian ammonite fauna of Poland and its palaeogeographical significance. Acta Geologica Polonica 35, 199219.Google Scholar
Marcinowski, R. & Wiedmann, J. 1990. The Albian ammonites of Poland. Palaeontologia Polonica 50, 194.Google Scholar
Martill, D. M. 1993. Soupy substrates: a medium for the exceptional preservation of ichthyosaurs of the Posidonia Shale (Lower Jurassic) of Germany. Kaupia – Darmstädter Beiträge zur Naturgeschichte 2, 7797.Google Scholar
Massare, J. A. 1987. Tooth morphology and prey preference of Mesozoic marine reptiles. Journal of Vertebrate Paleontology 7, 121–37.Google Scholar
Maxwell, E. E., Caldwell, M. W. & Lamoureux, D. O. 2011. Tooth histology in the Cretaceous ichthyosaur Platypterygius australis, and its significance for the conservation and divergence of mineralized tooth tissues in amniotes. Journal of Morphology 272, 129–35.CrossRefGoogle ScholarPubMed
O'keefe, F. R. 2001. A cladistic analysis and taxonomic revision of the Plesiosauria (Reptilia: Sauropterygia). Acta Zoologica Fennica 213, 163.Google Scholar
Owen, R. 1840. Report on British fossil reptiles. Reports of the British Association for the Advancement of Science 9, 43126.Google Scholar
Owen, R. 1841. Odontography; or, a Treatise on the Comparative Anatomy of the Teeth; their Physiological Relations, Mode of Development, and Microscopic Structure, in the Vertebrate Animals. London: Hippolyte Baillière, 655 pp.Google Scholar
Owen, R. 1851–1864. A Monograph on the Fossil Reptilia of the Cretaceous Formations. London: The Palæontographical Society, pp. 118.Google Scholar
Owen, R. 1860. Note on some remains of Polyptychodon from Dorking. Quarterly Journal of the Geological Society London 16, 262–3.Google Scholar
Owen, H. G. 2012. The Gault Group (Early Cretaceous, Albian), in East Kent, S.E. England; its lithology and ammonite biozonation. Proceedings of the Geologists' Association 123, 742–65.Google Scholar
Peryt, D. 1983. Planktonic foraminiferal zonation of Mid-Cretaceous of the Annopol Anticline (Central Poland). Zitteliana 10, 575–83.Google Scholar
Popov, E. V. & Machalski, M. 2014. Late Albian chimaeroid fishes (Holocephali, Chimaeroidei) from Annopol, Poland. Cretaceous Research 47, 118.Google Scholar
Pouech, J. J. 1881. Note sur un fragment de mâchoire d'un grand Saurien trouvé à Bedeille (Ariège). Bulletin de la Société Géologique de France 10 (3), 7987.Google Scholar
Pożaryski, W. 1947. A phosphate deposit of the north-eastern margin of the Holy Cross Mountains. Biuletyn Państwowego Instytutu Geologicznego 27, 156 (in Polish, English summary).Google Scholar
Radwański, A. 1968. Ischyodus thurmanni Pictet & Campiche and other chimaeroid fishes from the Albian–Cenomanian of the Holy Cross Mountains (Poland). Acta Palaeontologica Polonica 13, 315–22 (in Polish, English summary).Google Scholar
Radwański, A., Wysocka, A. & Górka, M. 2012. Miocene burrows of the ghost crab Ocypode and their environmental significance (Mykolaiv Sands, Fore-Carpathian Basin, Ukraine). Acta Geologica Polonica 62, 217–29.Google Scholar
Roberts, A. J., Druckenmiller, P. S., Sætre, G.-P. & Hurum, J. H. 2014. A new Upper Jurassic ophthalmosaurid ichthyosaur from the Slottsmøya Member, Agardhfjellet Formation of central Spitsbergen. PLoS ONE 9, e103152.Google Scholar
Rozhdestvenskiy, A. K. 1973. The study of Cretaceous reptiles in Russia. Paleontological Journal 2, 206–14.Google Scholar
Sachs, S. & Kear, B. P. 2014. Postcranium of the paradigm elasmosaurid plesiosaurian Libonectes morgani (Welles, 1949). Geological Magazine, published online 20 November 2014. doi:10.1017/S0016756814000636.Google Scholar
Samsonowicz, J. 1925. Esquisse géologique des environs de Rachów sur la Vistule et les transgressions de l'Albien et du Cénomanien dans les sillon nord-européen. Sprawozdania Państwowego Instytutu Geologicznego 3, 45118 (in Polish, French summary).Google Scholar
Samsonowicz, J. 1934. Explication de la feuille Opatów (zone 45, colonne 33). Service géologique de Pologne, Carte Géologique Générale de la Pologne au 100.000-e. Warszawa: Państwowy Instytut Geologiczny, 97 pp.Google Scholar
Sauvage, H. E. 1878. Prodrome des Plésiosauriens et des Elasmosauriens des formations Jurassiques supérieures de Boulogne-sur-Mer. Annales des Sciences Naturelles, Zoologie 13, 138.Google Scholar
Sauvage, H. E. 1882. Recherches sur les reptiles trouvées dans le Gault de l'Est du Bassin de Paris. Mémoires de la Société Géologique de France 2 (3), 21–4.Google Scholar
Schumacher, B. A. 2008. On the skull of a pliosaur (Plesiosauria; Pliosauridae) from the Upper Cretaceous (Early Turonian) of the North American Western Interior. Transactions of the Kansas Academy of Science 111, 203–18.Google Scholar
Seeley, H. G. 1869. Index of the Fossil Remains of Aves, Ornithosauria and Reptilia, from the Secondary System of Strata arranged in the Woodward Museum of the University of Cambridge. Cambridge: Deighton, Bell and Co., 143 pp.Google Scholar
Seeley, H. G. 1874. Note on some of the generic modifications of the plesiosaurian pectoral arch. Quarterly Journal of the Geological Society London 30, 436–49.Google Scholar
Seeley, H. G. 1876. On an associated series of cervical and dorsal vertebræ of Polyptychodon, from the Cambridge Upper Greensand, in the Woodwardian Museum of the University of Cambridge. Quarterly Journal of the Geological Society London 32, 433–6.Google Scholar
Sirotti, A. & Papazzoni, C. 2002. On the Cretaceous ichthyosaur remains from the Northern Apennines (Italy). Bollettino della Società Paleontologica Italiana 41, 237–48.Google Scholar
Vincent, P., Bardet, N., Houssaye, A., Amaghzaz, M. & Meslouh, S. 2013. New plesiosaur specimens from the Maastrichtian Phosphates of Morocco and their implications for the ecology of the latest Cretaceous marine apex predators. Gondwana Research 24, 796805.Google Scholar
Wahl, W. 2009. Taphonomy of a nose-dive: bone and tooth displacement and mineral accretion in an ichthyosaur skull. Paludicola 7, 107–16.Google Scholar
Walaszczyk, I. 1987. Mid-Cretaceous events at the marginal part of the Central European Basin (Annopol-on-Vistula section, Central Poland). Acta Geologica Polonica 37, 6174.Google Scholar
Walaszczyk, I. 1992. Turonian through Santonian deposits of the Central Polish Upland; their facies development, inoceramid paleontology and stratigraphy. Acta Geologica Polonica 42, 1122.Google Scholar
Welles, S. P. 1943. Elasmosaurid plesiosaurs with description of new material from California and Colorado. Memoirs of the University of California 13, 125254.Google Scholar
Welles, S. P. 1962. A new species of elasmosaur from the Aptian of Colombia and a review of the Cretaceous plesiosaurs. University of California Publications in Geological Sciences 44, 196.Google Scholar
Welles, S. P. & Slaughter, B. H. 1963. The first record of the Plesiosaurian genus Polyptychodon (Pliosauridae) from the New World. Journal of Paleontology 37, 131–3.Google Scholar
Wetzel, A. & Reisdorf, A. G. 2007. Ichnofabrics elucidate the accumulation history of a condensed interval containing a vertically emplaced ichthyosaur skull. In: Ichnology at the Crossroads: A Multidimensional Approach to the Science of Organism-Substrate Interactions (eds Bromley, R. G., L. Buatois, A, G. Mangano, Genise, J. F. & Melchor, R. N.), pp. 241–51. SEPM, Special Publication no. 88.Google Scholar
Woods, M. A., Wilkinson, G. K., Booth, K. A., Farrant, A. R., Hopson, P. M. & Newell, A. J. 2008. A reappraisal of the stratigraphy and depositional development of the Upper Greensand (Late Albian) of the Devizes district, southern England. Proceedings of the Geologists' Association 119, 229–44.Google Scholar
Young, M. T., Brusatte, S. L., de Andrade, M. B., Desojo, J. B., Beatty, B. L., Steel, L., Fernández, M. S., Sakamoto, M., Ruiz-Omeñaca, J. I. & Schoch, R. R. 2012. The cranial osteology and feeding ecology of the metriorhynchid crocodylomorph genera Dakosaurus and Plesiosuchus from the Late Jurassic of Europe. PLoS ONE 7, e44985.Google Scholar