New material of the pareiasaur Deltavjatia rossicus from the Kotel'nich locality, Kirov Province, Russia, is described in detail. The taxon is characterised by a distinctive pattern of dermal sculpture and the exaggerated embayment of the posterior skull roof, resulting in the dorsal exposure of the braincase. Postcranially, Deltavjatia shares some aspects of its morphology with basal pareiasaurs, including the osteoderm pattern. Features such as the forward-slanting and pointed iliac blade are shared with stratigraphically younger, more derived forms. Well-preserved material of the taxon spans a wide size-range, allowing an assessment of ontogenetic trends. A geometric morphometric analysis of the skull roof of Deltavjatia reveals an allometric increase in snout length and postorbital area, a result that can serve as a basis for examining morphological trends within pareiasaurs. A reassessment of pareiasauromorph relationships, using both parsimony and Bayesian methods of phylogenetic inference, recovers similar topologies in both cases. Four Bayesian analyses were completed, with and without a gamma-shaped parameter and with and without the inclusion of autapomorphies. Despite differing taxon and outgroup selection, the recovered topologies are similar to previous phylogenies of pareiasaurian relationships, with Deltavjatia appearing as a relatively basal taxon.

The Insect Limestone has long been known as a source of exquisitely preserved insects and other arthropods. It occurs on the north side of the Isle of Wight and is latest Eocene in age. A summary of the history of the study of the Insect Limestone is given, along with detailed stratigraphical and sedimentological information. This paper is the first in a thematic set of papers.

Antarctica has been known as the “keypiece” of the Gondwana supercontinent since publication of Du Toit's 1937 classic book Our Wandering Continents. It is also important to reconstruction of the early Neoproterozoic supercontinent Rodinia. Laurentia, with its circumferential late Precambrian rifted margins, can be regarded as the ‘keypiece’ of Rodinia. The Southwest US–East Antarctica (SWEAT) hypothesis suggested former juxtaposition of the Pacific margins of Laurentia and East Antarctica. Several new lines of evidence support this hypothesis in a revised form, but must be reconciled with opening of the Pacific Ocean basin predating amalgamation, not only of Gondwana, but even of today's East Antarctic craton. The sequence of events is envisaged to have been: (1) formation prior to 1·6 Ga of a craton, including Laurentia and the Mawson craton, that extended from South Australia along the present Transantarctic margin to the Shackleton Range; (2) suturing of southernmost Laurentia to the Kalahari craton along the Grenville, Namaqua–Natal–Maud orogenic belt ca. 1·0 Ga; (3) rifting of the East Antarctic margin (Mawson craton) from western Laurentia ca. 0·7 Ga; (4) pan-African suturing of the Mawson craton to southernmost Laurentia as Gondwana amalgamated, forming the ephemeral Pannotia supercontinent; and (5) end-Precambrian separation of Laurentia as Iapetus opened.

New fossil vertebrate microremains from the Lower Silurian of NW Mongolia, Tuva and S Siberia have been discovered, and previous collections of thelodonts (Vertebrata: Thelodonti) from this region re-studied, figured and described, following recent advances in morphology and systematics of thelodont scales. As a result, six thelodont species are described here and attributed to two families and three genera. An emended diagnosis is given for each species. Morpohological scale varieties of each species are revised, and the squamation types are introduced to the species descriptions, resulting in newly grouped morphological sets of scales. The previously monotypic genus Talimaalepis Žigaitė, 2004, is proposed to contain two different species; therefore, two new combinations of morphological scale sets of two species are described for the first time. Both of them show transitional scale structure between the genera Loganellia (family Loganellidae) and Helenolepis (family Phlebolepididae). Two of the thelodont genera, Angaralepis and Talimaalepis, as well as all the six species, are endemic and not known anywhere else in the world. They are accompanied by a variety of other peculiar early vertebrates, such as mongolepids, two endemic genera of acanthodians, and putative galeaspids. This work enhances our knowledge of early Silurian vertebrate diversity, and provides evidence of regional palaeoenvironmental conditions and palaeogeographical relationships of the Siberia and Tuva terranes.

Earth history is punctuated by events during which large volumes of predominantly mafic magmas were generated and emplaced by processes that are generally accepted as being, unrelated to ‘normal’ sea-floor spreading and subduction processes. These events form large igneous provinces (LIPs) which are best preserved in the Mesozoic and Cenozoic where they occur as continental and ocean basin flood basalts, giant radiating dyke swarms, volcanic rifted margins, oceanic plateaus, submarine ridges, and seamount chains. The Mesozoic history of Antarctica is no exception in that a number of different igneous provinces were emplaced during the initial break-up and continued disintegration of Gondwana, leading to the isolation of Antarctica in a polar position. The link between the emplacement of the igneous rocks and continental break-up processes remains controversial. The environmental impact of large igneous province formation on the Earth System is equally debated. Large igneous province eruptions are coeval with, and may drive environmental and climatic effects including global warming, oceanic anoxia and/or increased oceanic fertilisation, calcification crises, mass extinction and release of gas hydrates.

This review explores the links between the emplacement of large igneous provinces in Antarctica, the isolation of Antarctica from other Gondwana continents, and possibly related environmental and climatic changes during the Mesozoic and Cenozoic.

The morphologies of the hyobranchial apparatus in early tetrapods are reviewed, based primarily on first-hand examination and supplemented by published descriptions. The basic arrangement of the “aquatic” hyobranchium, with four pairs of branchial arches and internal gills, was conserved to a remarkable degree across the fish–to–tetrapod transition and was retained in further evolution in adults of several tetrapod lineages. Thus, a fish-like hyobranchium in basal tetrapods does not necessarily represent a larval or paedomorphic character, respectively, as was often suggested in analogy to extant salamanders. Rather, it represents the plesiomorphic state of the adult hyobranchium in tetrapods. The changes in the hyobranchium during the fish–to–tetrapod transition include the reduction of the number of skeletal elements and their morphological simplification. In all three presently discussed scenarios of lissamphibian origin, the temnospondyl, lepospondyl and diphyly hypotheses, the internal gills were reduced independently within temnospondyls and on the amniote stem below seymouriamorphs. Evidence of remodelling into a true “terrestrial” hyobranchium, with reduction of the posterior branchial arches and modification to support terrestrial tongue feeding, is scarce in early tetrapods. It evolved within temnospondyls in zatracheids, amphibamids and lissamphibians, as well as once or several times in early amniotes or in their immediate stem-forms.

Latest Eocene fossil plant remains occur in concentrations within blue-grey micrite known as Insect Limestone near the base of the Bembridge Marls Member (Bouldnor Formation, Solent Group), Isle of Wight, southern England. Some of the previously reported taxa (collections in the Natural History Museum, London) are not preserved within the Insect Limestone. These (e.g., all Arecaceae (palms)) are excluded from the floral list. New non-destructive techniques have yielded additional taxonomic information. Leaves previously assigned to Ficus and Fagus are now incertae sedis. Wetland elements are abundant, particularly Typha, but also Acrostichum, Azolla, Potamogeton, Sparganium and others. Non-wetland elements are rare. Trees and shrubs included representatives of Betulaceae, Caprifoliaceae, Juglandaceae, Lauraceae, Rhamnaceae (the sclerophyllous Zizyphus), other flowering plants and several genera of conifers. There are rare specimens of possible herbaceous plants and propagules with plumes or awns, the latter possibly an early fossil record of Clematis. The common plant remains were probably derived from vegetation near a freshwater body, sometimes with slight brackish influence, whilst rarer elements were probably blown in from a greater distance. There is little evidence of plant–insect interaction; one leaf with small galls, a stem containing an insect larva and a possible association between stratiomyid flies and Typha.

The discovery of marine Pliocene diatoms in warm-based glacial deposits (now termed the Sirius Group) high in the Transantarctic Mountains in the 1980s began a three-decade-long controversy over the stability of the East Antarctic Ice Sheet. Their presence implied that this ice sheet had collapsed as recently as three million years ago to allow their deposition in shallow interior seas, followed by transport and deposition from an expanded over-riding ice sheet. Though the glacial deposits included clasts with older diatoms, no evidence of clasts with Pliocene diatoms was published, but the hypothesis gained wide acceptance. Increasing knowledge of ice sheet behaviour and the antiquity and stability of the Transantarctic Mountains, along with new techniques for dating age and denudation rates for landscapes, has led to a more likely alternative hypothesis – that the high-level Sirius Group deposits pre-date Transantarctic Mountains uplift and their Pliocene diatoms are atmospheric contaminants. Surveys have shown that marine diatoms from the Antarctic margin and the Southern Ocean are indeed reaching the surface of the ice sheet and blowing through the mountains, with permafrost processes providing opportunities for contamination. Modelling and geological evidence is now consistent with a stable East Antarctic Ice Sheet in the interior for the last 14 Ma, with some retreat around the margins and periodic collapse of the West Antarctic ice sheet in Pliocene times.

A revision of the Carboniferous lungfish genus Ctenodus is undertaken. Ctenodus is the longest-surviving genus of lungfish from the Carboniferous, and is the only one to cross the Lower–Upper Carboniferous boundary. Five species span the period, with species correlated to stage. Species are distinguished by differences in tooth plate morphology. Ridge number increases from the Tournaisian to the Moscovian, and the length to width ratio decreases. A new, almost complete articulated individual of C. interruptus allows identification of this species from pterygoid tooth plates and other characteristics. These, and new features of the postcranial skeleton, are described for the first time. Ctenodus retains primitive features of the skull roof, such as a pair of C-bones and retention of a D-bone; its contemporaries Conchopoma, Sagenodus and Straitonia show the derived condition of these features. Ctenodus retains primitive features of the lepidotrichia, which are distally segmented and bifurcated. However, it also shows the derived condition of the tail region in having a combined first and second dorsal fin: whether there was a separate caudal fin remains unknown.

Molluscs from the Insect Limestone of the Bembridge Marls Member are described. The gastropods Lymnaea (Galba) longiscata (Brongniart 1810), Planorbarius discus (Edwards 1852), Gyraulus similis Férussac, 1814, Hippeutis headonensis Jodot, 1942, Viviparus lentus (Solander 1766) and Viviparus angulosus (J. Sowerby 1817) indicate freshwater alkaline conditions. The cerithioid gastropod Tarebia acuta (J. Sowerby 1822) probably represent more brackish conditions. Polymesoda (Pseudocyrena) obovata (J. Sowerby 1817) is the only bivalve present. All were potentially washed into the depocentre. Despite a wide outcrop of the Insect Limestone on the Isle of Wight's northwest and northeast coasts, there are no records of molluscs other than from the Gurnard/Thorness Bay area on the northwest coast of the island.

The twin summits of Preshal More and Preshal Beg, near Talisker, Isle of Skye, comprise the erosional remnants of a thick (at least 120 m) compound olivine tholeiite lava, or flow field, that ponded in palaeo-valleys within the Palaeocene lava field of west-central Skye. This unique flow field constitutes the Talisker Formation and is the youngest preserved extrusive unit of the Skye Lava Field. The lava inundated a complex of palaeo-valleys incised into the higher stratigraphical levels of the existing lava field, and remnants of the original sedimentary fill of these valleys still exist, the Preshal Beg Conglomerate Formation. The lava displays spectacularly well-developed two-tier (colonnade-entablature) columnar joint sets that formed as a consequence of slow, uninterrupted cooling through its base and sidewalls, aided by groundwater circulation and water ingress (from displaced drainage) directed into the lava's interior by master-joint systems. Intrusive phenomena developed at both the base and the top of the lava and there is evidence for the existence of subsurface feeder tubes. The tholeiitic composition of the Talisker Formation lava contrasts with the transitional, mildly alkaline characteristics of the remainder of the (older) lavas of Skye Lava Field. In broad terms, the Talisker Formation lava is compositionally very similar to the suite of cone-sheets emplaced into the oldest of the four intrusive centres that comprise the Skye Central Complex – the Cuillin Intrusive Centre – together with a high proportion of the Skye regional dyke swarm. The stratigraphical position, field relationships and compositional characteristics of the lava indicate that it was erupted and emplaced as an intracanyon-style flow field during the early shield-building stage in the growth of the (tholeiitic) Cuillin Volcano, which post-dates the main Skye ‘plateau’ Lava Field. Although the remnant outcrops are detached from their likely source area through erosion, this tholeiitic lava provides the first direct evidence linking the central complexes of the British Palaeogene Igneous Province and their eruptive products.

Once thought to be devoid of life, the Antarctic Ice Sheet is now known to be a dynamic reservoir of organic carbon and metabolically active microbial cells. At the ice-bed interface, subglacial lake and sedimentary environments support low diversity microbial populations, adapted to perennial cold, anoxia and lack of light. The dynamic exchange of water between these shallow environments conveys meltwaters and associated sediments into the coastal ocean. This, together with the release of iceberg-rafted debris to more distal coastal environments, could be important for sustaining primary productivity in the iron-limited Southern Ocean, via the release of associated nutrients and bioavailable iron. We estimate the magnitude and review the wider impacts of the potential export of nutrients (N, P, C, Si and bioavailable Fe) dissolved and associated with suspended sediments in Antarctic runoff and entombed in iceberg rafted debris. Located beneath subglacial lakes and the subglacial till complex are deep sedimentary basins up to 14 km thick, located largely around the Antarctic periphery. These sedimentary basins are largely hydrologically decoupled from shallower lake and till environments by the presence of highly consolidated sediments which limit the penetration of glacial meltwaters to depth. They provide extensive habitats for sustained microbial activity over Ma timescales, and are likely to be a focal point for the anaerobic cycling of organic carbon and other elements in the deep sub-surface. Organic carbon buried in these basins during ice sheet formation is thought to be microbially cycled to methane gas, and the methane largely stored as hydrate within sediments, stabilised by the high pressure/low temperature conditions. We conclude that the export of nutrients and biogenic gases from deep and shallow subglacial Antarctic environments designates Antarctica as a potentially important component of the Earth's carbon cycle, and highlight the importance of evaluating these potential impacts further via global and regional-scale biogeochemical modelling.

The arachnids from the late Eocene Insect Limestone of the Isle of Wight are reviewed. Only spiders (Araneae) have been recorded, of which several dozen are known, mostly juveniles. Nearly all are araneomorphs; the putative mygalomorph Eoatypus woodwardii McCook, 1888 was removed from that infraorder by Selden (2001), but new specimens, yet to be described, could belong in this group. Among the adult specimens, which are thus identifiable to family level, most belong to the argyronetid water-spider Vectaraneus yulei Selden, 2001; some resemble araneids, and a small number can be referred to other families. New morphological information on the holotype of Vectaraneus is presented, and three other specimens are described in more detail here, in Segestriidae, Araneidae and Salticidae. Segestriids are nocturnal sedentary hunters living in tubular retreats in rock or bark crevices. Araneids are the archetypal weavers of orb webs. Salticids are the familiar jumping spiders; the one described here is an ant mimic. Their presence suggests a drier habitat was present somewhere in the source environment for the fossil biota, and they were probably inhabitants of the raised hammocks suggested for the Insect Bed palaeoenvironment.

The odonatan fauna of the Late Eocene of the Isle of Wight is revised. The following taxa are revised or described: the gomphaeschnids Oligoaeschna? anglica Cockerell & Andrews, 1916 and Anglogomphaeschna eocenica gen. et sp. nov.; the aeshnids ‘Oplonaeschna’ vectensis Cockerell & Andrews, 1916, Aeschnophlebia andreasi Nel et al., 2005, Oligaeschna wedmanni sp. nov., and a ‘Gynacanthinae’ species; Neophya legrandi sp. nov., first fossil representative of the Cordulephyidae; three undescribed ‘Corduliidae’; Eomacrodiplax incompleta gen. et sp. nov., first fossil representative of the Urothemistidae; the second representative of the Palaeogene family Bolcathoridae; a Thaumatoneuridae Dysagrionini species A; the megapodagrionid Oligoargiolestes oligocenum Kennedy, 1925; the two hypolestids Anglohypolestes fasciata gen. et sp. nov. and Eohypolestes hooleyi gen. et sp. nov.; the coenagrionid ‘Enallagma' oligocena Cockerell & Andrews, 1916, and three other undescribed species; Angloprotoneura emilielacroixi gen. et sp. nov., first fossil European representative of the damselfly family Protoneuridae; and the lestid Lestes aff. regina Théobald, 1937. This fauna has strong similarities with the Recent Afrotropical and Indo-Malayan Odonata, suggesting a warm palaeoclimate for the Late Eocene of the Isle of Wight. ‘Megalestes’ anglicus Cockerell, 1915 is a Zygoptera Lestiformia or Coenagrionomorpha of uncertain affinities.

Data are the foundation of modern observational science. High-quality science relies on high quality data. In Antarctica, unlike elsewhere, researchers must disperse data and conduct science differently. They must work within the laws enacted under Antarctic Treaty that defines Antarctica as a continent for peace and science, where data sharing and international collaboration are requisite keystones. Scientists also work under oversight guidance of the Scientific Committee on Antarctic Research (SCAR). In the last decade, rapid technological advances and vast increase in digital data volumes have changed the ways data are acquired, communicated, analysed, displayed and reported. Yet, the underlying science culture in which data are funded, utilised and cared for has changed little. Science-culture changes are needed for greater progress in Antarctic science.

We briefly summarise and discuss aspects of Antarctic ‘data care’, which is a subset of data management. We offer perceptions on how changes to some aspects of current science-culture could inspire greater data sharing and international collaboration, to achieve greater success. The changes would place greater emphasis on data visualisation, higher national priority on data care, implementation of a data-library concept for data sharing, greater individual responsibility for data care, and further integration of cultural arts into data and science presentations.

Much effort has gone into data management in the international community, and there are many excellent examples of successful collaborative Antarctic science programs within SCAR built on existing data sets. Yet, challenges in data care remain and specific suggestions we make deserve attention by the science community, to further promote peace and progress in Antarctic science.