We analyzed the recent (< 25 yr) spread in New Hampshire, USA, of the exotic tree Kalopanax septemlobus, native to Asia. The invasion was likely initiated by a single tree planted ca. 1972. Our objective was to assess the viability of the invasion, especially in light of the small propagule size. We tallied, mapped, aged, and measured the height and growth of K. septemlobus individuals at two sites, the University of New Hampshire campus (UC) and Thompson Farm (TF), both in Durham. We found over 3,800 plants at UC and 270 at TF in < 120 ha (296 ac) total area. Plant age ranged from 0 to 22 yr, and UC plants were as far as 775 m (2,543 ft) from the purported parent tree. Annual height growth was comparable to midtolerant native trees. Plants occurred in both open and forested habitats, and the mean level of photosynthetically active radiation incident on understory plants was 4 to 6% of full sun. The large population size, shade tolerance, rapid height growth, and ability to sprout from damaged stems suggest potential for K. septemlobus to invade and persist in forests, the most common natural ecosystem in the northeastern United States. We further suggest that small propagule size, likely a single tree, has not prevented K. septemlobus from initiating a spatially extensive and vigorous population. Kalopanax septemlobus has been planted as an ornamental in the northeastern United States, and prevention of region-wide invasion might depend on removal of these trees, even when they occur as single individuals.

Brachypodium sylvaticum, a shade-tolerant, forest dwelling, and aggressive invasive grass native to Eurasia, is a noxious weed in California, Oregon, and Washington. This species could cause ecosystem collapse by altering forest fire regimes. To examine interactions with fire, we divided two Willamette National Forest sites into eight units and randomly selected half for treatment with prescribed fire in spring 2011. We assessed the effect of B. sylvaticum on fire (severity and intensity) as well as the effect of fire on B. sylvaticum (cover, seedling emergence, and dispersal). We found that B. sylvaticum cover decreased fire severity but had no effect on intensity. Furthermore, fire severity influenced B. sylvaticum cover; in areas receiving low-severity fire, the grass increased from 21 ± 15.05 to 34 ± 15.81%, but in areas of high-severity fire, cover remained consistently around 0% (0 ± 0% cover in yr 1 to 0.2 5± 0.25% in yr 3). In the field, prescribed fire decreased seedling emergence by 32% compared to controls, but not in an associated greenhouse experiment. However, in the greenhouse, severely burned plots had zero emergence, compared to 0.29 ± 0.14 seedlings low-severity m−2 plot. Fire severity also influenced dispersal in the field; we monitored plots with < 0.5% cover B. sylvaticum initially; when these plots experienced low severity fire, they had greater B. sylvaticum cover (increasing 1,200%), suggesting increased dispersal with less severe fires. High-severity dispersal plots did not experience increased cover. High severity fires have the potential to control the grass, but low-severity fires will likely increase its cover.

The macrophyte swamp stonecrop has invaded a wide range of wetland habitats across Europe. An experiment was conducted within an invaded fen habitat, which tested whether the presence of grazing disturbance affected the relative abundance of swamp stonecrop, and whether any detected effect was suppressive or facilitative. The abundance of swamp stonecrop and co-occurring resident plants was monitored within fenced grazing exclosures and in adjacent unfenced plots. Swamp stonecrop abundance was higher in the unfenced plots compared to the fenced exclosures (t(87) = 28.974, P < 0.001), whereas the abundance of co-occurring plants was higher in the fenced exclosures compared to the unfenced plots (t(87) = 6.264, P < 0.001). These results indicate that the presence of large vertebrates could facilitate a higher abundance of swamp stonecrop in situations where competitive resident plant species were selectively removed by these grazing animals.

Southern California grasslands have largely been type-converted to dominance by exotic annual grasses, leading to displacement of many native grass and forb species. Crimson fountaingrass, Pennisetum setaceum, an exotic perennial C4 species and a relatively new invader to California, is expanding to areas currently occupied by purple needlegrass, Stipa pulchra, a C3 native. We predicted that fountaingrass seedlings might withstand cool season competition in California's Mediterranean-type climate and establish in Stipa pulchra grasslands due to less competition during the warm, dry summer season, and that interactions might be influenced by density. A field experiment was conducted to examine competitive interactions of the two species from the cool winter season to the warm summer season. As predicted, Stipa produced greater aboveground biomass in the cool season and showed strong intraspecific competition, as well as interspecific suppression of Pennisetum growth, whereas Pennisetum showed no suppression of Stipa. In the warm season, Stipa showed relatively less suppression of Pennisetum, erasing significant differences, and Pennisetum showed increased growth. Results of this study show that C3Stipa can suppress initial growth of C4Pennisetum in the cool season, but in warmer months, Pennisetum can overcome this initial suppression at both low and high densities, even within a Mediterranean-type climate with little to no summer rainfall. Thus, in southern California, temporal niche partitioning due to photosynthetic pathway in these two species can allow Pennisetum invasion. Given the similarity in life history and growth form of Stipa and Pennisetum, few options exist for controlling Pennisetum in habitats where Stipa occurs. In these cases, restoration plantings of desirable species are essential in order to reestablish competitive vegetation that will be more resistant to invasion.

Grazing represents a sustainable alternative for the control of medusahead infestations but intake of medusahead by ruminants is typically low. We determined in sheep whether (1) high-energy supplements enhance medusahead intake and preference relative to a treatment without supplementation and (2) individual differences in medusahead intake transfer to individual differences in the ingestion of a silica-containing ration. Groups of lambs (n = 10) were individually penned and randomly assigned to three supplementation treatments: (1) Control (no supplement); (2) Calcium (Ca) Propionate (beet pulp : barley : Ca propionate, 67 : 30 : 3); or (3) Yeast (beet pulp : barley : yeast culture, 65 : 30 : 5). After supplementation, all animals had ad libitum access to medusahead in late vegetative to mid-reproductive (Trial 1), and late-reproductive phenological stages (Trial 2). Medusahead preferences were assessed by offering sheep a choice between medusahead and tall fescue hay. Plant part preferences were assessed by offering a choice between medusahead tops (mostly seedheads and awns) and mid-plant parts (mostly stems + leaves). Intake of medusahead was low and cyclic, declining towards the end of each trial (P < 0.0001) and there were no treatment differences (P > 0.10). Lambs preferred tall fescue hay to medusahead and medusahead tops to mid-plant parts (P < 0.0001). Supplemented lambs gained more weight than control lambs (P < 0.10). Thus, supplemented lambs performed better than nonsupplemented controls without reducing their intake of medusahead. In Trial 3, two new groups of lambs were formed based on their intake of medusahead during Trials 1 and 2 (n = 10). One group consistently ate more medusahead and more of a ration containing silica (alfalfa : silica, 97 : 3) than the other (P < 0.10). A significant and consistent degree of individual variation was measured among lambs—irrespective of treatment—regarding their ability to ingest medusahead and silica. This variation represents a promising option for maximizing use of medusahead by livestock in rangelands.

Chinese tallowtree is an invasive tree found throughout the southeastern United States and in California. Its negative effects can be seen in numerous natural and managed ecosystems, including bottomland hardwood forests, pastures, pine plantations, and along lakes, ponds, streams, and rivers. Despite its troublesome presence for many decades, relatively few effective control strategies are available. Root sprouting following management efforts is a major impediment to successful control. Studies were conducted in Alabama and Louisiana at three locations to test several herbicides for cut stump, basal bark, and foliar individual plant treatment (IPT) methods. Herbicide treatments included triclopyr amine and ester formulations, imazamox, aminopyralid, aminocyclopyrachlor, and fluroxypyr. Data were collected just before leaf senescence at one and two growing seasons after treatment and included Chinese tallowtree foliar cover, number of stump or root collar sprouts, and number of sprouts originating from lateral roots within a 1-m radius of each tree. For the cut stump and basal bark studies, most herbicide treatments prevented sprouting from the stump or root collar region better than they did from the lateral roots. Aminopyralid reduced total sprouting better than all other treatments in the cut stump study. The high rates of aminocyclopyrachlor and fluroxypyr resulted in the highest mortality in the basal bark study. Aminocyclopyrachlor reduced total sprouting better than all other herbicides in the foliar treatment study. Triclopyr amine and ester formulations, which are commercial standards, did not consistently control Chinese tallowtree across these IPT studies. These studies provide some promising treatments to increase the number of effective tools that can be used to manage Chinese tallowtree. Additional research is needed to address the prolific nature of lateral root sprouting following any of these treatment methods.

An ecosystem's invasibility is influenced by changes in biotic and abiotic resistances, which often occur due to shifts in the prevailing disturbance regime. The susceptibility of a community to intrusion by nonnative species may interact with propagule pressure to determine the extent of a biological invasion. We examined how propagule pressure, forest community structure, and disturbance interact to influence the invasibility of temperate Pacific Northwest forests by the newly invasive grass, perennial false-brome (Brachypodium sylvaticum). Our goal was to identify factors enabling shifts from establishment to population growth in B. sylvaticum populations at the edge of its expanding range. Ecological sampling methods were used to identify patterns in B. sylvaticum habitat. An inverse relationship between the amount of B. sylvaticum and all perennial vegetation types and soil litter depth was found, suggesting that disturbance might play a role in B. sylvaticum population establishment or growth. An experimental study was then performed to test the effects of disturbance, propagule pressure, and habitat on B. sylvaticum seedling establishment in sites where B. sylvaticum was already naturalized. We found evidence that disturbance of the soil and vegetation led to increased B. sylvaticum seedling recruitment within naturalized sites, especially where conditions of high propagule pressure and deciduous forest canopy existed. In contrast, B. sylvaticum populations dominated by coniferous forest canopy were much more invasible than deciduous forests and did not show increased seedling recruitment in response to our disturbance treatments. Our study shows how propagule pressure and plant community dynamics interact to alter the invasibility of Pacific Northwest forests allowing B. sylvaticum to transition from establishment to population growth thus allowing this weed to cause greater negative impacts on the ecosystem.

Energy-extraction disturbances entail soil handling and often have large edge-to-area ratios. These characteristics should be considered when designing weed-control strategies. In western North America, many energy developments coincide with infestations of downy brome, an annual grass that severely curtails productivity, diversity, and habitat value of invaded areas. Downy brome is sensitive to soil compaction and seed burial, both of which may occur when soil is handled. In this study, I examined the effect of soil-density manipulations and herbicide application (105 g ai ha−1 imazapic with 280 g ai ha−1 glyphosate) on six simulated pipeline disturbances in a Wyoming big sagebrush ecosystem invaded by downy brome. Disturbances occurred at the end of the growing season, after ambient downy brome seed rain in the study areas had abated. Treatments and seeding occurred shortly after disturbances. The following spring, downy brome seedling density was 10-fold lower within disturbances than in control areas, but seedling density quickly rebounded in disturbed areas where no herbicide had been applied. In herbicide plots, downy brome seedling density remained low during the first growing season, and shrub cover after 3 yr was eight times higher than in no-herbicide plots. Soil density manipulations via disking and rolling treatments had little effect on downy brome. Prior research has shown that imazapic is more effective when combined with disturbances, such as fire. This study demonstrates that imazapic may also be effective in combination with a disturbance that is timed to bury downy brome seeds.

Rural landscapes are increasingly diverse and heterogeneous, involving a mix of small and large parcels, amenity and agricultural properties, and resident and absentee owners. Managing invasive plants in landscapes with changing ownership requires understanding the views and practices of different landowners. We surveyed landowners in two rural valleys with 26% absentee ownership and a large number of small parcels in Missoula County, Montana. Landowners indicated a high level of awareness and concern about weeds; more than 80% agreed that weeds are a problem in their valley. Seventy-eight percent of landowners managed weeds, but only 63% were effective at weed management. Absentee owners were far less likely to manage weeds on their properties and less likely to utilize herbicides, as compared with resident landowners. Landowners reported that seeds coming from adjacent properties were the most significant barrier to effective weed control. Many landowners manage weeds to be a good neighbor and believe that cooperation between neighbors is critical to weed management.

Waterhyacinth is a free-floating aquatic weed that is considered a nuisance worldwide. Excessive growth of waterhyacinth limits recreational use of water bodies as well as interferes with many ecological processes. Accurate estimates of biomass are useful to assess the effectiveness of control methods to manage this aquatic weed. While large water bodies require significant labor inputs with respect to ground-truth surveys, available technology like remote sensing could be capable of providing temporal and spatial information from a target area at a much reduced cost. Studies were conducted at Lakes Columbus and Aberdeen (Mississippi) during the growing seasons of 2005 and 2006 over established populations of waterhyacinth. The objective was to estimate biomass based on nondestructive methods using the normalized difference vegetation index (NDVI) derived from Landsat 5 TM simulated data. Biomass was collected monthly using a 0.10m2 quadrat at 25 randomly-located locations at each site. Morphometric plant parameters were also collected to enhance the use of NDVI for biomass estimation. Reflectance measurements using a hyperspectral sensor were taken every month at each site during biomass collection. These spectral signatures were then transformed into a Landsat 5 TM simulated data set using MatLab® software. A positive linear relationship (r2 = 0.28) was found between measured biomass of waterhyacinth and NDVI values from the simulated dataset. While this relationship appears weak, the addition of morphological parameters such as leaf area index (LAI) and leaf length enhanced the relationship yielding an r2 = 0.66. Empirically, NDVI saturates at high LAI, which may limit its use to estimate the biomass in very dense vegetation. Further studies using NDVI calculated from narrower spectral bands than those contained in Landsat 5 TM are recommended.

The invasive aquatic plant hydrilla rapidly spread through the 28,500 ha Kissimmee Chain of Lakes (KCOL) system in Florida in the late 1980s and early 1990s. Large-scale herbicide treatments with fluridone were initiated in 1993 and resulted in widespread reduction in hydrilla; however, by 2000, sustained use of fluridone resulted in dominance of fluridone-resistant strains of hydrilla throughout these lakes. The last large-scale fluridone applications on the KCOL were conducted in 2004, and in 2012, a sampling effort was initiated to determine the status of fluridone-resistant strains of hydrilla given an 8-yr period with no further selection pressure from fluridone. A total of 260 sites were sampled on the lakes during March, May, September, and December 2012. Plants were returned to the lab and exposed to fluridone at concentrations of 5, 10, and 20 μg L−1 and a pulse-amplitude-modulated (PAM) fluorometer was utilized to measure fluorescence yield of new shoot tissue growth following a 14-d exposure period. Results indicate that 80 to 90% of the sites sampled on the four lakes of the Kissimmee Chain remain resistant to fluridone. Three distinct patterns of response to fluridone were noted, suggesting that susceptible, moderately tolerant, and highly tolerant strains of hydrilla currently coexist on these lakes. Although fluridone-susceptible plants were present on the KCOL, this study clearly demonstrates that most of the hydrilla remained resistant despite an 8-yr period with no fluridone selection pressure.

Lake Pend Oreille is the largest (36,000 ha or 91,000 ac) freshwater lake in Idaho. Approximately 27% or 10,000 ha of the lake is littoral zone habitat supporting aquatic macrophyte growth. Eurasian watermilfoil has invaded large areas of this littoral zone habitat, with early estimates suggesting approximately 2,000 ha by the mid 2000s. Idaho State Department of Agriculture developed a state-wide eradication program in response to the threats posed by Eurasian watermilfoil, which attempts to quantify Eurasian watermilfoil infestations and its effects on the native plant community. Littoral zone point intercept surveys were conducted in 2007 and 2008 to monitor the trends in aquatic macrophyte community structure and assess management strategies against Eurasian watermilfoil. Lake Pend Oreille has a species-rich aquatic macrophyte community of more than 50 species. Lake-wide, the presence of Eurasian watermilfoil significantly decreased from 2007 (12.5%) to 2008 (7.9%). The native plant community has remained stable from 2007 to 2008 despite lake-wide management activities. In managed areas, the frequency of Eurasian watermilfoil during the 2008 assessment was 23.6% after herbicide applications. This represents a 63% reduction in Eurasian watermilfoil presence from the 2007 (64.5%) survey. When 2,4-D was combined with endothall, the presence of Eurasian watermilfoil declined from 63% (2007) to 36.5% in 2008. Eurasian watermilfoil treated with triclopyr also declined significantly, 64% to 18.2%. When all treatment methods were pooled and compared with areas that were not treated, the presence of Eurasian watermilfoil was significantly greater (52.5%) in untreated areas as opposed to treated areas (23%). The removal of Eurasian watermilfoil resulted in an increase in native species in most areas. Currently, there is as little as 200 ha of Eurasian watermilfoil remaining, which represents an overall reduction of 90% in approximately 7 yr of management.

Weed prevention is recognized as one of the most cost-effective management strategies for invasive plants. In the field of invasive plant management increasing emphasis is being directed toward proactive management. However, land managers are still somewhat reluctant to aggressively employ prevention programs. Part of this reluctance could be due to lack of understanding of what a comprehensive prevention program entails. The purpose of this paper is to improve strategic decision-making for site-specific prevention programs, such as those on ranches or in watersheds. Our interest is in advancing prevention planning for land managers—the people who are faced with the constant pressure of potential invasive species infestations on a day-to-day basis. To facilitate more widespread use of prevention programs we are proposing definitions for key terminology to standardize and facilitate communication about prevention programs. Additionally, we present a flow model with the steps necessary to successfully implement such programs. The model has three categories from which specific prevention planning occurs: (1) education, (2) early detection and eradication, and (3) interruption of movement. The flow model directs users through a series of interlinked steps. Finally, we provide a case study in which a ranch manager implemented a prevention program using this framework. By using this model, managers are poised to conduct more strategic planning. This model also has applications in outreach and education programs to assist land managers in prevention planning.

Green spaces such as golf courses that intermingle within or exist on fringes of urban landscapes can provide opportunities for increasing the ecological value of urban areas. To that end, more naturalistic and less input-intensive “links”-style golf courses have recently gained favor over input-intensive parkland courses. The Osgood Public Golf Course in Fargo, ND is a links-style golf course set adjacent to suburban housing developments. This course incorporated large areas of prairie plantings, or “constructed grasslands,” which over time became dominated by fescue species and infested with Canada thistle. Our objective was to explore the efficacy of using prescribed fire combined with aminopyralid herbicide to control Canada thistle and promote a more diverse mix of warm-season C4 and cool-season C3 grasses. Aminopyralid was applied during fall 2010 and prescribed fire was applied during spring 2011. We found that aminopyralid provided excellent control of Canada thistle 1 and 2 yr post-treatment. Open niches created from Canada thistle control were readily filled by C3 grasses, primarily fescue species, which were the dominant species on the constructed grasslands prior to treatment. Fire intensity was variable within and across plots and was associated with reductions of litter and C3 grasses, but was not associated with increases of C4 grasses within the time frame of this study. Our results demonstrate the effectiveness of aminopyralid for Canada thistle control in constructed grasslands. Prescribed fire maintained C3 grass dominance while removing litter, but C4 grass response was variable and appeared dependent on pretreatment C4 species abundance. Reduction of litter in constructed grasslands dominated by fescue could potentially lead to microsite conditions that would favor C4 and other C3 species, especially if short-term management promoted additional facilitation efforts, such as repeat spring fire treatments and seeding.

The objective of this article is to extend the reported period in which flood-distributed knotweed propagules may be successfully managed using only manual labor. During a second round of early detection and rapid response (EDRR) management along the Green River in Guilford, VT, we collected and measured all Japanese knotweed propagules that had been distributed by flooding approximately 21 mo earlier, in August 2011, at a single site. Our data suggest that knotweed s.l. prioritizes the growth of new stems over new rhizomes at the start of a growing season. Because the limiting factor for successful removal of new knotweed s.l. plants by hand is the size of the rhizome system, our findings support extending the time frame for EDRR management of flood-distributed knotweed s.l. into the second spring after its initial dispersal. Additionally, in November 2013, surveys of our work sites found no new knotweed s.l. plants in locations accessible to work crews. In addition to validating our EDRR management techniques, this implies that knotweed s.l. fragment viability does not extend past the second spring following its dispersal.