Chinese tallowtree [Triadica sebifera (L.) Small] has reached unprecedented prevalence in coastal landscapes in the Gulf of Mexico, especially along edge habitat with low competition and abundant resource (e.g., light) availability. This study investigated the spatiotemporal patterns and mechanisms of T. sebifera spread along roadways and fire lines. Triadica sebifera individuals and landscape and community features were surveyed in equally spaced, spatially mapped plots. All T. sebifera individuals were felled to determine tree age and status (seed trees or non-seed bearing trees), and T. sebifera seed and seedling (≤2 yr old) densities and community and landscape features (over- and understory conditions, distance to seed trees) were measured. A zero-inflated negative binomial model was used to evaluate factors affecting T. sebifera seed dispersal and seedling recruitment contributing to the observed spatiotemporal patterns. Introduced into the Grand Bay National Wildlife Refuge around 30 yr ago, T. sebifera trees distribute in clustered patterns along roadways and fire lines and exhibit an exponential growth in density. High T. sebifera seed and seedling densities mainly occurred in sites that are ≤250 m from seed trees or have sparse overstory and high understory grass/herb coverage. With respect to the avian seed dispersal mechanism, the spatiotemporal patterns of T. sebifera spread along roadways and fire lines could be simply characterized by using landscape and community features that influence avian behaviors, including distance to seed trees, overstory tree density, and ground grass/herb coverage.

Japanese stiltgrass is one of the most aggressive, rapidly spreading invasive plants in the eastern United States. Management guidelines state that mowing can help manage Japanese stiltgrass but that mowing is most effective when done late in the season after the plants begin to flower and before they set seed. In this study, I tested the effectiveness of mowing at three different times between mid-June and early September in 2009 and 2010, as well as mowing twice in 1 yr and for two consecutive years. The effectiveness of mowing Japanese stiltgrass was determined by measuring percentage of cover, biomass, seed production, and the number of stems in the summer following mowing. All mowing treatments significantly reduced percentage of cover, biomass, seed production, and the number of Japanese stiltgrass stems the following year. In 2009, all of the mowing treatments significantly reduced biomass, percentage of cover, and seed production. The latest mow, at the end of August, resulted in a slightly greater reduction of cleistogamous seeds. In 2010, the earliest mowing treatment, in mid-June, did not reduce cover and biomass as much as the other mowing treatments. Overall, these results suggest that mowing can be an effective control method for Japanese stiltgrass and that mowing any time after June should be effectively equivalent, although later mowing may provide some marginal advantage.

White sweetclover is invading the Alaska glacial river floodplains and roadsides adjacent to natural areas, and control methods are needed. Chlorsulfuron, 2,4-DB, clopyralid, triclopyr, and 2,4-D controlled white sweetclover seedlings below recommended rates in the greenhouse. Biomass of established plants in the field was reduced by chlorsulfuron at recommended (17.6 g ai/ha), 1/2, and 1/4 rates and was reduced by triclopyr and 2,4-D at recommended rates (1,260 and 1,600 g ai/ha). Herbicides were more effective at reducing white sweetclover viable seed production in 2007 than in 2006. Only chlorsulfuron at 17.6 g ai/ha (recommended rate) eliminated seed production in both years. Flaming killed first-year plants, but some second-year plants resprouted and produced viable seed. Cutting at the 2.5 or 10 cm height did not control first-year plants because of regrowth, and second-year plant density and seed production was reduced by cutting at 2.5 cm but not by cutting at 10 cm.

Downy brome inhibits revegetation efforts following ecosystem disturbance. Imazapic is a commonly used herbicide for downy brome management, but more information is needed regarding effective application timing for restoration efforts. We wished to determine (1) if native species establishment exhibited a tradeoff between downy brome competition and injury from herbicide and (2) if this differed between pre- and postemergent applications of imazapic. We used a standard replacement series design and overlaid herbicide treatments. Nine weeks after planting, aboveground biomass was harvested and relative yield (RY) indices calculated. Both imazapic applications reduced downy brome biomass by 91% or more (P < 0.05). Imazapic caused drastic reductions in native biomass but less than what was caused by downy brome competition (P < 0.05). Natives were less injured by a pre- than postemergent application (P < 0.05). In situations where downy brome may impact restoration efforts, pre-emergent applications of imazapic at 70 g ai ha−1 (0.06 lb ai ac−1) may reduce downy brome with less negative impacts on newly-seeded native grasses than post-emergent applications. Ensuring sufficient proportions of native species seeds on restoration sites may reduce downy brome.

Common reed (Phragmites australis) is an invasive perennial grass in aquatic and riparian environments across the United States, forming monotypic stands that displace native vegetation that provides food and cover for wildlife. Genetic variation in global populations of common reed has given rise to two invasive haplotypes, I and M, in the United States. Our objectives were to (1) determine if any differences in herbicide efficacy exist with respect to common reed haplotypes I and M and (2) screen for other labeled aquatic herbicides that may have activity on common reed haplotypes I and M, most notably imazamox and diquat. A replicated outdoor mesocosm study was conducted in 1,136-L (300-gal) tanks using haplotypes I and M of common reed. Restriction fragment length polymorphism methodologies were used to verify the identification of I and M haplotypes used in this study. Diquat at 2.2 (1.9) and 4.5 (4.0) kg ai ha−1 (lb ai ac−1), glyphosate at 2.1 (1.8) and 4.2 (3.7) kg ae ha−1 (lb ae ac−1), imazamox at 0.6 (0.5) and 1.1 (0.9) kg ai ha−1 (lb ai ac−1), imazapyr at 0.8 (0.7) and 1.7 (1.5) kg ai ha−1 (lb ai ac−1), and triclopyr at 3.4 (3.0) and 6.7 (5.9) kg ae ha−1 (lb ae ac−1) were applied to the foliage of common reed. After 12 wk, no difference (P = 0.28) in herbicide tolerance was seen between the two haplotypes with respect to biomass. The 4.2-kg ae ha−1 rate of glyphosate and the 0.8- and 1.7 kg ai ha−1 rates of imazapyr reduced common reed by > 90% at 12 wk after treatment (WAT). Imazamox at 0.6 and 1.1 kg ai ha−1, and triclopyr at 3.4 and 6.7 kg ae ha−1 reduced common reed biomass (62–86%) at 12 WAT, though regrowth occurred. Diquat did not significantly reduce biomass by 12 wk. Glyphosate and imazapyr were the only herbicides that resulted in > 90% biomass reduction and corroborate control from previous studies.

Barb goatgrass is an invasive annual grass from the Mediterranean region that negatively affects both native plant biodiversity and the forage quality of grasslands. Prescribed burning may be the best landscape-level tool available to manage invasive species like barb goatgrass while also enhancing biodiversity, but few studies have quantified the long-term effects of fire on goatgrass and the rest of the plant community. We assessed the effects of fire on an invading front of barb goatgrass on a private ranch in Sacramento County, CA. We established burned and unburned treatment plots within the goatgrass-infested area and used prescribed fire to burn the treatment plots in June 2005. We monitored plant-community composition before burning and for 7 consecutive yr following the burn. Additionally, we tested the viability of goatgrass seeds in both burned and unburned plots. One year after the burn, goatgrass cover in burned plots was 3% compared with 21% in unburned plots. This reduction in goatgrass cover was still strong 2 yr after the burn (burned, 6%; unburned, 27%) and weaker but still statistically significant for 4 of the next 5 yr. The burn also reduced germination of goatgrass seed by 99% as indicated by seed-viability tests conducted in the laboratory. The native plant community responded positively to the burn treatment in the first year following the burn with an increase in native diversity in burned plots vs. unburned plots, but the effect was not detectable in subsequent years. Nonnative annual forb species cover also increased in the first year following the burn. Our study shows that a single springtime burn can result in a short-term boost in native species diversity, reduced seed germination of barb goatgrass to near zero, and reduced cover of barb goatgrass for at least 7 yr after the burn.

Greenhouse experiments were conducted to evaluate mile-a-minute response to selected herbicides. In the first experiment, mile-a-minute response was evaluated following the application of aminocyclopyrachlor, aminopyralid, fluroxypyr, glufosinate, glyphosate, imazamox, imazapic, metsulfuron, penoxsulam, and triclopyr. Applications of aminocyclopyrachlor, aminopyralid, fluroxypyr, glufosinate, glyphosate, and triclopyr all resulted in mile-a-minute control 70% or greater 8 wk after treatment (WAT). Mile-a-minute sensitivity to these six herbicides was further evaluated in a dose-response study. Although 90% growth reduction (GR90) values were determined for aminopyralid and glyphosate, statistically significant results were not obtained for aminocyclopyrachlor, fluroxypyr, glufosinate, and triclopyr. The results of these experiments suggest that glyphosate, as well as the synthetic auxin herbicides aminocyclopyrachlor, aminopyralid, fluroxypyr, and triclopyr, should be further evaluated for mile-a-minute control in south Florida.

One of Australia's most serious weeds, Chrysanthemoides monilifera subsp. rotundata (bitou bush) was recently found for the first time in Western Australia as a well established population in Kwinana, a major port and industrial area south of Perth, the State's capital. This population is remote from other bitou bush infestations in Australia and had escaped detection despite extensive surveys in the same State for the other subspecies that is present in Australia, Chrysanthemoides monilifera subsp. monilifera (boneseed). The main reasons it went undetected are thought to be the tightly controlled access to this area because of mineral processing and port activities, the unusual invasion route via a heavy industrial area and the morphological similarity to a native species when it is not flowering. Two surveys defined the core population of 1038 plants that are spread along the coast over a 25-ha semi-circle with about a 500-m (1640 ft) diameter. Subsequent surveys of first a 500 m buffer zone and later a 1-km (0.621 mi) buffer found four additional plants, indicating that there is considerable potential for dispersal. We concluded that the survey has not delimited the distribution because of the potential and evidence for long distance dispersal. Cooperation by the various land managers has led to all plants being killed, as an initial step to management of this species. Other steps to be undertaken include an awareness campaign in the area that would need to be surveyed for delimitation of the spatial distribution and seed bank assessment to measure potential dispersal both in space and through time. It remains to be determined what is the best strategic response: eradication or containment.

Many restoration projects rely on invasive plant removal to restore ecosystems. However, success of restoration efforts relying on invasive removal can be jeopardized, because in addition to displacing native plants, invasives can also dramatically impact soils. Many studies have documented invasives’ effects on soil chemistry and microbiota. While European beachgrass [Ammophila arenaria (L.) Link] is a worldwide invasive problem in coastal dunes outside northern Europe, little attention has been paid to effects of this species on soil chemistry following invasion, even though it establishes persistent, dense monocultures. In our study, we evaluated effects of A. arenaria invasion on soil chemistry of coastal dunes at Point Reyes National Seashore (PRNS); persistence of effects following removal by mechanical or herbicide treatment (legacy effects); and effects of treatment independent of invasion. Dune restoration efforts at PRNS have met with mixed success, especially in herbicide-treated backdunes, where decomposition of dead A. arenaria has been greatly delayed. Based on results, invasion impacted 74% of 19 variables assessed, although there was a significant interaction in many cases with successional status (earlier vs. later). Almost 60% of invasion effects persisted after restoration, with legacy effects prevalent in herbicide-treated backdunes where sand deposition from adjacent beaches could not mitigate effects as it could in herbicide-treated foredunes. Mechanical removal—or inversion of invaded surface soils with less-contaminated subsoils—resulted in fewer legacy effects, but more treatment effects, primarily in backdunes. Soil chemistry may decelerate decomposition of A. arenaria due to the limited nitrogen (N) available to enable microbial breakdown of the high carbon(C):N (70.8:1) material, but microbial factors probably play a more important role. Success of restoration at PRNS may not be fully realized until legacy effects are resolved through additional actions such as inoculation with healthy microbiomes or necromass reduction through controlled burning.

Chloroplast DNA sequences and recently established morphological characters were used to confirm the widespread invasion of California's vernal pools by European low mannagrass. Morphological similarities between low mannagrass and western mannagrass have led to different taxonomic treatments, depending on the geographical extent of a particular flora. When California's flora was last revised, the two species were combined as western mannagrass, which was then considered to be a native species. Unfortunately, the revised flora was published just as low mannagrass began to rapidly expand its range within the state and, because it was considered to be a native species in the new flora, no actions were initiated to limit the invasion. Our data show that low mannagrass was present at all localities in the Central Valley of California that were investigated, indicating a widespread and undetected invasion. The invasion has led to a degradation of the vernal pool ecosystems, which are the habitat of many federal and state protected endangered and threatened species.

Cabomba is a submersed aquatic plant native to the southeastern United States that is commonly sold worldwide through the aquarium trade. While infrequently managed in its native range, cabomba has recently been reported as invasive and tolerant to management efforts in the northern areas of the United States and in other countries. Invasive populations of cabomba are characterized by a phenotype that is bright green. In contrast, cabomba native to the southeastern United States is characterized by a red phenotype, while plants sold through the aquarium trade have intermediate characteristics of both the green and red phenotypes. The response of the three cabomba phenotypes to selected herbicides was evaluated by measuring photosynthetic response over the course of a static 144-hr exposure. Plants were exposed to the maximum recommended use-rates of 2,4-D, carfentrazone, copper, diquat, endothall (amine and dipotasium salt formulation), flumioxazin, quinclorac, triclopyr, and a combination of diquat and copper. A submersed plant species known to be sensitive to each of these herbicides was also included to compare photosynthetic response of the cabomba to a susceptible plant. The photosynthetic response of the red and green phenotypes differed following exposure to carfentrazone, diquat, 2,4-D, triclopyr, and flumioxazin. Diquat, diquat plus copper, endothall (amine salt), and flumioxazin were the only products that resulted in a greater than 50% reduction of photosynthesis in all three phenotypes of cabomba. A second experiment was conducted where all three phenotypes of cabomba were exposed to these four herbicides for 24 hr, and photosynthesis was evaluated. Following the 24-hr exposure, results further documented distinct response differences between the green and red phenotypes, with the green phenotype demonstrating a reduced sensitivity to the herbicides evaluated. Results demonstrate clear phenotypic differences in response to herbicide treatments and lack of susceptibility of cabomba to most herbicides.

Extensive areas in the upper Midwest have been invaded by spotted knapweed, and effective management strategies are required to reestablish native plant communities. We examined effects of mowing, mowing plus clopyralid, or mowing plus glyphosate in factorial combination with hand pulling and burning on knapweed abundances on a knapweed-infested site in western Michigan. We applied mowing and herbicide treatments in summer 2008, and seeded all plots with native grasses and forbs in spring 2009. We conducted the knapweed pulling treatment from 2009 to 2012 in July. The prescribed burn was conducted in April 2012. By 2012, hand pulling reduced adult knapweed densities to 0.57 ± 0.12 m−2 (0.053 ± 0.011 ft−2) (mean ± SE), which was 5.8% of nonpulled treatments, juvenile densities to 0.29 ± 0.07 m−2 (2.1% of nonpulled treatments), and seedling densities to 0.07 ± 0.06 m−2 (2.6% of nonpulled treatments). After 3 yr, hand pulling reduced seed bank densities to 68 ± 26 m−2 as compared to 524 ± 254 m−2 in nonpulled treatments and 369 ± 66 m−2 in adjacent untreated areas of the study site. Without hand pulling, effects of mowing or mowing plus glyphosate were short-lived and allowed knapweed to rapidly resurge. In comparison, although a single mowing plus clopyralid treatment maintained significantly reduced densities of knapweed for 4 yr, by 2012 knapweed biomass in the nonpulled clopyralid treatment was approximately 60% of that in the other nonpulled treatments. Burning had minimal impacts on knapweed densities regardless of treatment combination, probably as a result of low fire intensity. Results demonstrated that persistent hand pulling used as a follow-up to single mowing or mowing plus herbicide treatments can be an effective practice for treating isolated spotted knapweed infestations or for removing small numbers of knapweed that survive herbicide applications.

The macrophyte swamp stonecrop has invaded a wide range of wetland habitats across Europe. An experiment was conducted within an invaded fen habitat, which tested whether the presence of grazing disturbance affected the relative abundance of swamp stonecrop, and whether any detected effect was suppressive or facilitative. The abundance of swamp stonecrop and co-occurring resident plants was monitored within fenced grazing exclosures and in adjacent unfenced plots. Swamp stonecrop abundance was higher in the unfenced plots compared to the fenced exclosures (t(87) = 28.974, P < 0.001), whereas the abundance of co-occurring plants was higher in the fenced exclosures compared to the unfenced plots (t(87) = 6.264, P < 0.001). These results indicate that the presence of large vertebrates could facilitate a higher abundance of swamp stonecrop in situations where competitive resident plant species were selectively removed by these grazing animals.

Two field experiments were conducted at the Agronomy Research Station near Stillwater, OK in 2004 and 2005 to identify and evaluate herbicides applied preplant incorporated, preemergence, early postemergence, or late postemergence for the control of seedling sericea lespedeza. Trifluralin, applied preplant incorporated, controlled seedlings 77 (15 wk after emergence [WAE]) and 63% (16 WAE) in 2004 and 2005, respectively. Flumioxazin, imazapic, fluometuron, diuron, sulfentrazone, atrazine, metribuzin, and metolachlor applied preemergence all provided greater than 86% seedling control at 15 and 16 WAE in both years. Diclosulam, applied preemergent, controlled seedlings 47% at 15 WAE in 2004 and 91% control at 16 WAE in 2005. In 2004, triclopyr, metsulfuron-methyl, glyphosate, picloram, dicamba, and 2,4-D amine plus picloram (tank-mix) applied early postemergence controlled 90 to 100% of the sericea lespedeza seedlings at 15 WAE. However, in 2005, only triclopyr, metsulfuron-methyl, glyphosate, and 2,4-D amine plus picloram (tank-mix) showed greater than 80% control at 16 WAE. Triclopyr, applied late postemergence, controlled seedlings 100% at 15 and 16 WAE, in both years. In 2004, dicamba plus 2,4-D (premix) and glyphosate were the only other herbicides which provided greater than 75% control at 15 WAE. These data suggest that there were preemergence-applied herbicides that were effective for the control of seedling sericea lespedeza. These data also suggest that triclopyr was the most effective postemergence-applied herbicide for the control of seedling sericea lespedeza. The data also showed that the overall level of control of seedling sericea lespedeza decreased as the season progressed and the plant matured.

Downy brome (Bromus tectorum) is an invasive, annual grass that has spread through much of the Great Basin desert but remains patchy at high elevation. This study evaluates control options for outlier infestations in the eastern Sierra Nevada, CA, based on their ecological effectiveness and their economic and practical feasibility. I tested the efficacy of hand-pulling, sheet-mulching, and soil solarization followed by broadcast and seedball seeding of native forbs and grasses. Downy brome cover, density, and dominance in the seed bank decreased with all removal treatments. Soil solarization and sheet mulching were most successful at eliminating downy brome (decreasing density by 99% after just 1 yr of treatment in both cases), but they had negative nontarget impacts on other herbaceous species. Germination of native seeds was low with both broadcast and seedball seeding, probably because of dry conditions. Each of the methods tested has potential for decreasing or eliminating small-scale, outlier infestations of downy brome along roadsides and in disturbed sites and thereby helping to contain the invasion.

Miconia (Miconia calvescens DC) was introduced to the East Maui Watershed (EMW) a half-century ago with more than 25 yr of management recorded. Using a historical spatiotemporal data set, we constructed a leptokurtic dispersal kernel with 99% of progeny confined to within 549 m of the nearest maternal source and the remaining 1% dispersed out to 1,636 m. Seedbank persistence, based on postdated recruitment, displayed an exponential decay projecting extinction beyond 20 yr. These parameters are highly congruent to independent interpretations of M. calvescens in Australia and Tahiti. In a simulated stage matrix model, we projected management efforts to locally eradicate a small incipient propagule bank wherein optimal management was achieved with an annual harvest rate that eliminated all juvenile recruits before reaching maturity, until extinction. Based on current pricing for helicopter herbicide ballistic technology (HBT) operations, the optimal, variable cost to locally eradicate this incipient propagule bank was estimated to be less than US$42,000, with ~90% of the effort searching for the most distant 1% of the progeny expended within the first 9 yr after the mature discovery. This variable cost was sensitive to seedbank size, recruitment rate, and dispersal range, but was most sensitive to harvest rates between suboptimal and excess. In a scenario prioritizing the upper region of EMW, we retroactively analyzed past HBT efforts eliminating satellite M. calvescens and determined that 27% of the total effort resulted in 87% of the total protection to this priority asset, with every US$1 invested potentially avoiding US$184 in future costs. Management outside the priority area was less economical, with returns in protection diminishing with distance from the priority upper region. Miconia calvescens is currently not eradicable in the EMW, and full containment of the invasion would require a substantial increase in stable, long-term funding. With limited resources, local eradication of satellite M. calvescens could be the most cost-effective alternative to protecting uninvaded areas prioritized for critical ecosystem functions.

Vegetation that becomes overtopped usually experiences a decrease in abundance or species richness. When an overtopping plant alters the physiognomy of the existing vegetation (e.g., trees invading a shrubland), ecosystem processes can also be dramatically altered. Worldwide, Monterey pine (Pinus radiata) cultivars have been planted in Mediterranean-like climates and are known to invade surrounding natural communities. Ecological impacts resulting from these invasions have been widely investigated; however, the effects from solitary pine trees on the vegetation they overtop are lacking. Furthermore, studies on the impact of P. radiata cultivars from the California floristic province, where P. radiata is native, do not exist. In coastal California, north of the present-day range of native P. radiata stands, cultivars of this species have invaded northern coastal scrub vegetation. To determine the impact of pine invasion on species richness and structure in this habitat, floristic surveys were conducted in 20 blocks that consisted of invaded and uninvaded plots. An invaded plot contained two subplots located under the canopy of an isolated pine tree, whereas a paired, uninvaded plot contained two subplots located in coastal scrub adjacent to each pine. Pine trees selected ranged in size from 2.8 to 119 cm (1.1 to 46.9 in) basal diameter. Our results demonstrate that understory native cover and species richness are negatively correlated with tree size. Understory exotic plant cover and richness of species other than P. radiata did not show any correlation with tree size, mainly because exotic plants had a very low abundance overall.

Several studies have been carried out to evaluate the main drivers behind biological invasions and their ecological consequences. Nevertheless, it is still extremely difficult to acquire a full understanding of the invasion process due to its high level of complexity. The problems that complicate invasion studies are low detection during the early stages of invasion, and the high survey cost of working over large, inaccessible, and rugged areas. The studies that develop efficient tools to reduce costs and time will help to control or mitigate the invaders' damaging effects. Hieracium pilosella is an aggressive invader of grasslands worldwide. The aim of this work was to conduct a regional assessment for the suitability of H. pilosella invasion in the Fuegian Steppe by combining field surveys, spatial modeling, and geographic information system tools. We recorded the invader cover in 167 sample sites and we determined the relationship between environmental variables and the probability of invasion. This was carried out by the selection of alternative generalized linear models. The best model indicates that disturbance and plant community were the main drivers to predict invasion suitability on a regional scale. Therefore, these two variables were used as main inputs to construct a regional invasion suitability map, which identifies the most critical areas for prevention, control, and further monitoring. This approach corresponds to a tool that permits us to evaluate the suitability of invasion even over large and inaccessible areas. The results especially warn about soil disturbance associated with usual management practices in Fuegian rangelands.

Where the nonnative annual grass downy brome proliferates, it has changed ecosystem processes, such as nutrient, energy, and water cycles; successional pathways; and fire regimes. The objective of this study was to develop a model that predicts the presence of downy brome in Central Oregon and to test whether high presence correlates with greater cover. Understory data from the U.S. Department of Agriculture (USDA) Forest Service's Current Vegetation Survey (CVS) database for the Deschutes National Forest, the Ochoco National Forest, and the Crooked River National Grassland were compiled, and the presence of downy brome was determined for 1,092 systematically located plots. Logistic regression techniques were used to develop models for predicting downy brome populations. For the landscape including the eastside of the Cascade Mountains to the northwestern edge of the Great Basin, the following were selected as the best predictors of downy brome: low average March precipitation, warm minimum May temperature, few total trees per acre, many western junipers per acre, and a short distance to nearest road. The concordance index = 0.92. Using the equation from logistic regression, a probability for downy brome infestation was calculated for each CVS plot. The plots were assigned to a plant association group (PAG), and the average probability was calculated for the PAGs in which the CVS plots were located. This method could be duplicated in other areas where vegetation inventories take place.

Studies were conducted to document that European starlings consume Russian olive fruits and determine subsequent effects on seed germination. In the first study, avian feeding patterns at Russian olive trees were monitored over a 1-yr period using motion activated digital photography. Starlings fed on Russian olive fruits with highest activity occurring in November and December. In a second study, 20 captive European starlings were fed Russian olive fruits and seed germination rates were determined for three categories: consumed by starlings, hulled fruits (pericarp removed), and whole fruits. Starlings readily consumed Russian olive fruits and most seeds were regurgitated 30 min after consumption. Germination rates of ingested/regurgitated seeds (57%) and pericarp-removed seeds (40%) were greater than whole fruits (0%). Viability tests confirmed that 85% of starling ingested seeds remained viable after consumption. Our data suggest that Russian olive dispersal may be dependent upon animals for effective spread.