Incomplete knowledge of the drivers of local productivity may contribute to the on-going decline of farmland bird populations despite conservation efforts. We therefore investigated spatial and temporal variation in breeding phenology, nest habitat, and nest survival in the Corn Bunting Emberiza calandra, a species of conservation concern in central and western Europe. Among 225 nests from seven study areas in cropland-dominated, mixed, and grassland-dominated landscapes, nesting phenology spread between April and August, started earlier at lower altitudes, and progressed from grassland to arable crop and agri-environment scheme flower field habitats. Nest habitats varied substantially between years, but most nests were placed in cultivated land where they were prone to fail due to land-use operations. Nest survival differed markedly between habitats, being lowest in second-year flower fields and highest in fallow grassland. Since we protected known nests, raw survival estimates only account for “natural” nest losses, e.g. due to adverse weather or predation. To estimate additional “anthropogenic” nest losses, we calculated probabilities of nesting schedules to conflict with patch-specific land-use dates. After taking into account the estimated conflict probabilities of 0.56 for mowing, 0.64 for clover harvest, and 0.38 for grazing, habitat-specific “total Mayfield nest survival” estimates declined to 21%, 13%, and 20% for meadows, alfalfa/clover-grass leys, and pastures, respectively. These habitats held about two thirds of nests in mixed landscapes, highlighting the relevance of land use-driven nest losses. To enhance productivity beyond the thresholds required for local population persistence, we propose refined conservation schemes that improve nest survival within production farmland, best coupled with the development of prolific “Corn Bunting landscapes”.