1.1 Outline of Sections

In this Element, we investigate human well-being through the evolutionary lens and assess the potential of evolutionary insights to inform policy and institutions designed to maximize human flourishing. We critically review influential literature that highlights what some scholars consider to be both biological and cultural universals – such as collaboration and altruism – and their relation to well-being (Reference Bowles and GintisBowles & Gintis, 2013; Reference HaidtHaidt, 2006; Reference Sober and WilsonSober & Wilson, 2013; Reference WelzelWelzel, 2013; Reference WilsonWilson, 2015, Reference Wilson2019).

In Section 2, we explore a strain in research on happiness that highlights cultural differences with regard to the ideals of good life and developmental paths (Reference Krys, Capaldi and ZelenskiKrys et al., 2021a, Reference Krys, Park and Kocimska-Zych2021b). Large efforts are underway to develop quantitative surveys that – unlike the World Happiness Report – privilege culturally sensitive approaches to human well-being, studying the uniqueness of diverse normative patterns and ideas of social development. The question of whether the culturally sensitive and universalist approaches are at loggerheads is perhaps spurious. We argue that these perspectives complement one another by shedding light both on the distinctive traditions within, and on meeting points between, cultures.

In Section 3, we discuss the relationship between well-being and the ideas of good life as they evolved over millennia and across cultures. In spite of the cultural variety of eudaemonic ideals, there are striking parallels between various traditions that associate the good life with balance, working for the good of others, and the importance of compassion. These similarities spring from cultural learning and cross-pollination, but they also have a source in the evolutionary sciences’ idea of a shared human nature. The increasing evidence for the commonalities of human nature calls for revisiting the philosophical stance of cultural relativism, which undergirds the constructivist approaches of social anthropology and the humanities. We highlight the work of David Sloan Wilson’s ProSocial agenda, as he comes closest to crafting a comprehensive vision of well-being as anchored in the ethos of work for the betterment of humanity and the planet.

In Section 3.2, we discuss an intriguing reversal in the dynamic relationship between the social and evolutionary sciences. Well into the twenty-first century, the agenda of the humanities and the social sciences rested on the idea of social improvement. Evolutionary biology was largely identified as the study of selfish genes that were taken to stand in the way of human moral advancement. In the last decades – while social scientists plunged into declinism –evolutionary thought moved to a hopeful history of humankind based on the salience of altruism, prosociality, and cooperation. What are the implications of this intellectual transition?

In Section 4, we present a novel, multilevel selection (MLS) model for well-being, which emphasizes how human evolution has occurred under two often conflicting pressures: individual and group selection. Individuals are incentivized to be selfish to outcompete in-group members. At the same time, individuals are compelled to cooperate with in-group members in order to strengthen the group for competition against other groups. We propose that it can be profitable to think of human well-being as having evolved to motivate individuals affectively to contribute both to their own and their group’s success. Our contention in this Element is that approaching happiness and meaning as connected rather than disjointed involves a double-fold dividend. First, it counteracts a conceptual overabundance in the field of positive psychology. Second, it draws attention to how experiences of well-being help individuals manoeuvre between the potentially conflicting pressures of individual and group selection. We sum up these insights with the equation: ‘Happiness + Meaning = Well-Being’.

Section 4 is supplemented with insights from seminal works of narrative psychology relevant for our focus on well-being and meaning. Jerome Reference BrunerBruner (1990) adds valuable narrative perspectives on the human search for meaning, as does Reference FranklViktor Frankl (1946) with his groundbreaking work on logotherapy in the treatment of concentration camp survivors.

In Section 5, we draw attention to the Nordic countries – with Norway in the spotlight. We explore challenges to these nations’ transition from being welfare states to becoming well-being societies. In the twenty-first century, Norway, Denmark, Sweden, Finland, and Iceland have increasingly let their policies be guided by a shifting emphasis from welfare, whose basis is largely socio-economic, to highlighting human well-being – that is, creating institutional structures for helping citizens to create better lives for themselves. An important part of Nordic policy has been to socialize citizens into engaging in altruistic activities. A shift from economic growth to concentrating on citizens’ well-being is not without frictions but it is increasingly part of the Nordic governments’ agenda.

In Section 6, we sum up our findings and reinspect the policy implications of applying the evolutionary lens to well-being. Both insights from the literature we scrutinized and our multilevel selection model of human well-being inform an integrative approach to human flourishing, one that draws attention to its non-zero-sum sources. Our purpose is to offer a cross-culturally applicable framework for well-being that can be used to identify which policies are most likely to create happier lives for more people.

2.1 Culturally Sensitive Approaches to Human Flourishing

There is a current within positive psychology which emphasizes the cultural relativity of well-being and insists on respecting cultural particulars. In WEIRD countries (Western, Educated, Industrialized, Rich, Democratic), it is taken for granted that individuals should pursue happiness. The proponents of the culturally sensitive approach argue that universalizing Western traits simplifies human diversity. Desiring a good life may be universal, but cultures understand happiness differently, emphasize distinct aspects of well-being, and have conflicting ideals as to what level of happiness is desirable. To what extent well-being is viewed as an individual or social phenomenon varies from culture to culture (Reference Krys, Capaldi and ZelenskiKrys et al., 2021a, Reference Krys, Park and Kocimska-Zych2021b; Reference Rappleye, Komatsu, Uchida, Krys and MarkusRappleye et al., 2020; Reference Uchida and KitayamaUchida & Kitayama, 2009; Reference Uchida, Townsend, Markus and BergsiekerUchida et al., 2009).

Positive psychologists representing the relativist position work on developing quantitative surveys that assess well-being across dozens of nations. Respondents’ answers with regard to different aspects of well-being are weighted on the basis of their culture’s specific preferences for what constitutes a good life. This allows for a ranking of nations more in line with cultural experiences, redolent of how purchasing power parity (PPP) functions for economic comparisons. An alternative World Happiness Report could see less of a Western dominance across its headline findings.

In our view, while cultural sensitivity is valuable in that it requires an eye for difference, it is also important to identify transcultural commonalities. Evolutionary perspectives can help uncover the predispositions that cultural differences build on, as well as make sense of seeming paradoxes. The fact that some cultures favour independent, individualistic happiness pursuits while others have a preference for interdependent, communal practices, is not a random result of history. What is adaptive varies with the cultural, political, and economic aspects of the environment.

Our modern environment consists of various levels of organization – from the interpersonal social group (e.g., family) to the impersonal moral group (e.g., nation) – often with overlapping constellations that we belong to on each level. We have family groups, kin groups, social groups, professional groups, interest groups, voluntary groups, geographically and politically delineated groups, and many others. These groups can have shared and conflicting interests. The level of individual independence within these collectives varies from culture to culture.

At the conceptual level, we can still draw a clear distinction between pressures of individual and group selection, even in those environments in which individuals are more dependent on their social collective to succeed. In kinship societies, the well-being of the kin group is of such importance to each member’s fitness that interdependent, collectivistic concerns take precedence (Reference HenrichHenrich, 2020). In most Western cultures, individual strategies are paramount. A culturally sensitive approach to well-being could still define happiness as related to individual selection yet recognize that distinct cultural values incentivize diverse strategies in terms of social organization when individuals solve adaptively relevant challenges. We suggest how to investigate these dynamics in Section 6.2.

In environments that make social groups especially important for individual flourishing, our evolutionary perspective predicts that members of a group benefit when each individual’s well-being system is synchronized with those of the other members. This perspective illuminates why interdependent happiness assessments focus on how individuals feel in relation to other group members (Reference Hitokoto and UchidaHitokoto & Uchida, 2015). To respond to signals of opportunity or threat, groups form collectivist cultures so that they can feel well or ill together – at least to a greater extent than in individualistic cultures. Happiness still relates to individual selection, but must be shared among more people – one’s in-group – to optimize for adaptivity. The quantitative challenge is to operationalize such insights in cross-cultural surveys.

3.1 Enter Evolutionary Science

In the 2000s, David Buss, a pioneer of evolutionary psychology, and Randolph Nesse, a pioneer of evolutionary medicine, made valuable contributions to positive psychology. Reference BussBuss (2000) concluded that modernity detracts from people’s happiness primarily due to environmental mismatch and larger comparison groups. We live in a more prosperous environment, but this fact does not undermine that there can be a benefit to feeling distress when our adaptive strategies fail, as such emotions signal that we should change our strategies.

Reference Nesse, Huppert, Baylis and KeverneNesse (2005) elaborated on the adaptive functions of positive and negative emotions, which explains why these affects evolved. He proposed that an evolutionary approach could offer a theoretical bridge from which we can better understand human emotions as they relate to goal pursuit, as well as suggest policies that align with our desires and predispositions. Reference Hill, Buss, Eid and LarsenHill and Buss (2008) noted that with the field’s high stakes, it was ‘surprising that few researchers have yet to explore subjective well-being from an evolutionary perspective’. Only sporadic contributions have since been made. Evolutionary scholars have researched the drivers of human flourishing and discontent – for instance Gluckman and Hanson’s Mismatch (Reference Gluckman and Hanson2006) – but rarely within the field of positive psychology. A notable exception is Positive Evolutionary Psychology (Reference Geher and WedbergGeher & Wedberg, 2019), which offers an overview of psychological predispositions and their relevance to human thriving. In Introduction to Positive Evolutionary Psychology – which is part of the same Element series as our Element – Reference Geher, Fritche, Goodwine, Lombard, Longo and MontanaGeher et al. (2023) illustrate the utility of applying evolutionary thinking to positive psychology.

Many evolutionary scholars’ focus on the largely negative sources of human well-being – such as selfishness and self-interest – is something of a puzzle. By way of speculation, the hegemony of the selfish gene theory could be explained, at least in part, through the strength of logos, ethos, and pathos in the narratives emphasizing selfishness. The logos had to do with Richard Dawkins’ (Reference Dawkins1976) masterly, scientifically grounded narrative, anchored in the dominant behaviourist theories and pointing to selfishness as the ultimate motif all human pursuits, including acts of altruism. But the pathos and ethos played a possible role as well; in the era when Dawkins’ book was published, the promotion of individual, egoistic ends coincided with the rise of neoliberal ideals marshalled by influential thinkers such as Friedrich Hayek, Milton Friedman, and Ayn Rand. There were, of course, alternative views: the great philosopher Mary Reference MidgleyMidgley (2010), we recall, criticized the selfish gene theory for presenting a one-sided, reductionist view of humans as ruled by self-interest alone, and for threatening to legitimate, even fetishize, neoliberal individualism as the basis of human well-being.

When seen in this context, evolutionary science’s interest in cooperation and prosociality has marked a seeming paradigm shift. We write ‘seeming’ because the shift has been less a dramatic breakthrough and more a scholarly ricorso: a return to strains and ideas that had been subjects of earlier scholarly studies. Illuminating examples are Lee Dugatkin’s Cooperation Among Animals: An Evolutionary Perspective (Reference Dugatkin1977) and Robert Axelrod’s The Evolution of Cooperation (Reference Axelrod1984). More importantly, many ideas that emerge from modern labs working on animal cooperation gesture towards evolutionary theories by Peter Kropotkin. When Kropotkin undertook an epic journey through Siberia in the 1860s, the evolutionary orthodoxy advanced that the natural world was a brutal place. And yet, instead of nature red in tooth and claw, Kropotkin found countless examples of mutual aid within species, as well as evidence of the superiority of groups that practiced altruism and cooperation. A close observation of the behaviour of migrating birds, mammals, fish schools, and insect societies made him conclude that a driving evolutionary force behind all social life – be it in microbes, animals, or humans – was the law of mutual aid (Reference Woodcock and AvakumovićWoodcock & Avakumović, 1950).

From our present-day perspective, Kropotkin’s findings were of course incomplete – and as one-sided as Dawkins’ theory of selfish genes. But they paved the way for a careful rereading of Darwin’s ideas and rebalancing the studies of evolutionary biology, anthropology, and psychology through complementing the focus on ruthless competition with studies of empathy and altruism as sources of human well-being.

Buss and Nesse limited themselves to studying well-being mostly from a perspective that connects happiness to in-group competition – that is, individual selection. When we solve adaptively relevant challenges – those that ultimately promote our survival and reproduction – we are rewarded by positive feelings that motivate us to repeat such behaviour. Unhappiness evolved to tell us that our strategies are failing and that we need to change course. Feelings of happiness can be viewed as a compass that steers us towards successes that exceed those of our comparison group. Yet this is only one part of the equation. Considerable well-being can be derived from altruistic behaviour that does not directly enhance fitness for the altruist (Reference Dolan, Peasgood and WhiteDolan et al., 2008; Reference Meier and StutzerMeier & Stutzer, 2008; Reference Musick and WilsonMusick & Wilson, 2003; Reference Piliavin, Keyes and HaidtPiliavin, 2003; Reference Thoits and HewittThoits & Hewitt, 2001).

Roy Baumeister’s work on meaning illuminates how positive affects evolved to reward people for contributing to their community (Reference BaumeisterBaumeister, 2005; Reference Baumeister, Vohs, Aaker and GarbinskyBaumeister et al., 2013). Acting in line with cultural ideals – such as helping others – offers the altruist a sense of meaning that can have a more lasting effect on their well-being than the happiness rush that is typically derived from winning in-group competitions. The term hedonistic treadmill refers to how successful happiness pursuits often only lead to a temporary increase in well-being (Reference Diener, Lucas, Scollon and DienerDiener et al., 2009b). This is another apparent paradox of positive psychology, which an evolutionary perspective can help illuminate.

The pursuit of meaning can – in contrast to happiness quests – engender a sense of self-esteem and social belonging. It can also transform individual identity, elevating the altruist’s quality of life for months or years. Success in such pursuits also tends to be more available, as helping others is a less competitive field. Our drive for meaning, which is uniquely human, offers a win-win source of well-being, one that benefits the altruists, those they help, and the community they all belong to. Cultures able to compel individuals to aid in-group members gain advantages in terms of group selection.Footnote ⁷ A community in which people take care of each other will have more resourceful members and greater cohesion than one dominated by selfishness (Reference Wilson, Hessen, Witoszek and MidttunWilson & Hessen, 2018).

3.2 Evolutionary Thinkers as World Improvers

The idea that anything altruistic is cultural and anything selfish is biological would be a gross simplification. Empathy is not just a cultural construct; it is certainly useful at the biological level in the sense that those who are driven to care for their family and friends assist with the survival of those family and friends and thus increase their own fitness and survival when that altruism is reciprocated. Similarly, an excessive focus on the self, together with the promotion of selfish behaviour, does not exclusively spring from selfish genes. In a country like the United States, selfishness has become a cultural hallmark, an ostensible condition of a successful and happy life.

In this context, the agenda of social improvement highlighted by the prosocial wave of evolutionary thought marks an intriguing turn.Footnote ⁸ Evolutionary theory as preached and practiced by such thinkers as Frans De Waal or David S. Wilson is no longer associated with legitimizing the ‘demonic’ side of modernity. Exit imperialism, racism, and colonial might-is-right. Enter the survival of the friendliest and kindest. The synergy between biology and historical-cultural perspectives has produced a new narrative: evolution is not just a parade of selfish genes, but also progress towards altruism and prosociality. This is an interesting trend because it runs counter to a deluge of social science studies focused on the multiple, and seemingly irresolvable, crises of our time such as the apparent death of democracy, and even a descent to a New Middle Ages (Reference WooldridgeWooldridge, 2020). In defiance of these apocalyptic scenarios, Rutger Bregman argues in Humanity: A Hopeful History, ‘It’s when crisis hits when the bombs fall or the floodwaters rise – that we humans become our best selves’ (2019: 4).

Bregman contributes to a growing body of literature which offers a positively charged, optimistic view of human potential. The idea that humans are hard-wired primarily for selfishness and ruthless competition has been challenged by Frans de Reference WaalWaal (2010). He argues that the success of Homo sapiens rests to a significant extent on our capacity for empathy and our urge to understand and appreciate others. While being competitive, humans also possess an innate sensitivity to the emotional status of other members of our species. De Waal cites the consensus among biologists that empathy arrived with the evolution of maternal care in mammals. Crucial to this process was the controlling role of the hormone oxytocin, which induces empathy in males and females alike.Footnote ⁹ Chimpanzees care for mates who are wounded by leopards. Elephants offer ‘reassuring rumbles’ to youngsters in distress. Dolphins guide sick companions to near the water’s surface to prevent them from drowning.

The well-being of many animals and humans is the result of an emotional contagion: a happy and joyful environment is likely to generate happy and joyful individuals. Empathy holds our societies together, drives us to look after the sick, and makes us more flexible under stress. Through a better understanding of empathy’s survival value in evolution, de Waal suggests, we advance a less simplistic and more accurate view of the complexity of human nature that allows us to work together towards a more just and flourishing society.

A similarly upbeat view of human development is championed by evolutionary psychologist Steven Reference PinkerPinker (2012, Reference Pinker2019). Probing into the adaptive functions of our cognitive and emotional systems, Pinker documents how parts of humanity have progressed from violent tribalism to modern human-rights societies. He uses a telescopic, historical perspective to argue that the images of violence and misery peddled by contemporary media do not give an accurate picture of human development. Using empirical evidence and ample statistics, he substantiates how the present era is less violent, less cruel, and more conducive to nurturing human well-being than any previous period. Most people today are not only less likely to meet a violent death; they are increasingly able to seek happiness and self-realization.Footnote ¹⁰

Pinker reveres the Enlightenment, which he defines as the triumph of humanism and scientific breakthroughs. It was in the time of the Lumières that people gradually began to question forms of violence that had previously been taken for granted: slavery, torture, despotism, duelling, and extreme forms of cruel punishment. The ensuing decrease in suffering contributed to greater well-being. Countless individuals from across the world escaped honour killings, retaliations, and the traumas of religious persecution, setting out to search for happiness. The American dream was born.

Needless to say, Pinker’s one-eyed, Cyclopean vision of the Enlightenment has been the subject of many critical assaults for its inaccurate account of inequality and a cavalier attitude to the dark sides of modern globalization (Reference GoldinGoldin, 2018). The eighteenth century’s progress was not just about reason and science. It was inseparably connected to Western empire-building, and the selected horrors of the Industrial Revolution, whose advocates preached and practiced slavery, genocide, exploitation, and racism. Ironically, this murky Enlightenment – or Endarkenment – sought legitimation for its inhumanity by quoting evolutionary science, in its Spencerian version, and the laws of nature.Footnote ¹¹

Despite this massive criticism, Pinker’s stance on the advancement towards a kinder world has remained heroically unflinching. It is hard to disagree, he argues, that – in a relatively short time – many societies have come far from the days when lynchings were commonplace, torture was ubiquitous, women were treated as cattle, and children were put to hard work at the age of seven. What about the civilizational collapse caused by the unspeakable horrors of Nazi and Bolshevik death factories? Even here, Pinker insists, violence was ‘no more extreme’ than the massive genocide and mayhem caused, say, by thirteenth-century Mongol conquests that led to the deaths of an estimated forty million people – not far from the more than fifty-five million who died in World War II; and this was in a world with only one-seventh of the population of the mid-twentieth century.

Pinker’s lofty argument is that after World War II, most Western countries experienced the ‘long peace’, marked by the ‘rights revolution’, and growing revulsion against violence inflicted on ethnic minorities, women, children, sexual minorities, and even nature. Despite the ongoing conflicts, the ideal of eudaimonic happiness has taken hold of the imagination of people from all over the world; witness the narrative about the ‘slumdog millionaire’ in an award-winning Indian blockbuster. Its message of hope and redemption sent to the wretched inhabitants of Indian slums invokes a classic American narrative of the rise from rags to riches. But the film is not, as some critics contend, a re-branding of the American dream as Indian and sending it back to the West. Rather, in our view, it is about the universal human longing for happiness, prosperity, freedom, and recognition – a quest whose modern version was codified in a multiethnic society like the United States.

Genetic evolution can hardly explain these rapid changes, reasons Pinker. Evolution shaped the basic design of our brain, but it also equipped us with our cognitive and emotional faculties, which have been invested into cultural norms and values. The result is that propensities for violence – our ‘inner demons’ – exist side by side with ‘the better angels of our nature’ (Abraham Lincoln’s words) that incline us to be peaceful and cooperative. It is our material circumstances and existential constraints, along with the working of morally charged cultural values and practices, that determine whether the demons or the angels gain the upper hand.

3.3 Cooperative and Reciprocal Humans on a Path to Eudaimonia

A preoccupation with the pivotal importance of altruism and cooperation in human evolution is increasingly penetrating interdisciplinary scholarship. Evolution may have created ‘selfish genes’, as Richard Reference DawkinsDawkins (1976) would have it, but this is only one part of a more complex picture. Citing Adam Smith’s Theory of Moral Sentiments, Reference DarwinDarwin (1874) argued that the

Brian Hare and Vanessa Woods (2020), who examined the meaning of Spencer’s ‘survival of the fittest’, argue that this term cannot be reduced to ‘dog eats dog’. Darwin’s theory of evolution is as much about individual competition as it is about cooperation, symbiosis, and reciprocity. Friendliness and cooperation rather than dominance, Hare and Woods insist are the keys to evolutionary survival. Modern evolutionary biologists have entered into a fruitful cooperation with economists, anthropologists, archaeologists, and psychologists who provide empirical evidence for social cooperation as one of the core conditions for the survival of the species (Reference Buchanan and PowellBuchanan & Powell, 2018; Reference CorningCorning, 2012; Reference HaidtHaidt, 2006; Reference HenrichHenrich, 2017; Reference Richerson and BoydRicherson & Boyd, 2005; Sober & Wilson, 2013; Reference WelzelWelzel, 2013).

Insights from evolutionary psychology (Reference Cosmides and ToobyCosmides & Tooby, 2013), cross-cultural psychology (Reference Ryan and DeciRyan & Deci, 2017), evolutionary anthropology (Reference Richerson and BoydRicherson & Boyd, 2005), clinical psychology (Reference Hayes, Hofmann and WilsonHayes et al., 2020), and experimental philosophy (Reference Guglielmo, Monroe and MalleGuglielmo et al., 2009) indicate that certain human proclivities, such as a thirst for freedom and fairness, derive from evolutionary root principles that relate to all cultures. Christian Reference WelzelWelzel (2013) contends that an emancipative drive is universal, constituting the motivational source of human empowerment, which – in the last instance – increases people’s well-being.

This emancipative trajectory does not imply a linear process. Our urge for freedom is countered by a desire for security. Our quest for justice is offset by the human penchant for ruthless domination. Our ability to cooperate is countered by our compulsion to compete. The works we reviewed in the previous paragraphs argue that these opposing drives are evolutionary universals. The prominence of one or the other trait in dominant cultural narratives has a bearing on societies’ predicament and the well-being of their members. Cultures that advocate tolerance and social inclusion have high levels of human flourishing. Those that promote a competitive-sectarian ethos often succumb to social polarization that can make nations, tribes, and empires crumble (Reference TurchinTurchin, 2007).

Cultural evolution is a zig-zag process replete with advancements and regressions. Sophisticated Etruscan and Harappan civilizations were vanquished by muscular barbarians. The tolerant European Enlightenment was for decades eclipsed by Fascism and Bolshevism. Yet in each case, cooperative and prosocial societies resurfaced and rebuilt themselves after multiple traumas. There are several explanations for these resurrections. One is the evolutionary pull of prosocial moral sentiments, meaning that groups predisposed to cooperate and uphold ethical norms tend to survive and expand relative to more selfish groups.

Numerous studies contend that despite setbacks, the advancement towards modern, happier societies has been dependent on cultural values that foreground cooperation, altruism, fairness, and emancipative aspirations (Reference CorningCorning, 2011; Reference HaidtHaidt, 2006; Reference RicardRicard, 2015; Reference WelzelWelzel, 2013; Reference WilsonWilson, 2016; Reference WilsonWilson & Hessen, 2015). Research on 170 societies shows that feelings of freedom enhance a population’s life satisfaction. In Freedom Rising, Reference WelzelWelzel (2013) portrays happiness as being founded on a ‘ladder of freedoms’, which is highly dependent on people’s level of existential stress. If evolution has instilled in humans an adaptive quest for liberty, this drive can hibernate under authoritarian pressures but awaken when existential opportunities widen.

Welzel makes a strong case for the connection between emancipative aspirations and well-being. The human search for freedom is universal, he insists, and this drive becomes stronger the more prosperous a population becomes. The theoretical foundation of his approach –and its evolutionary roots – is described as ‘the utility ladder of freedoms’. To support his claims, Welzel quotes data from World Values Survey that show that societies with the highest levels of social emancipation are also the top scorers on global well-being indexes.

Rich, first-hand accounts mapping human behaviour in extreme situations concur that cooperative, rather than selfish, practices were crucial in the struggle for survival (Reference FranklFrankl, 1946; Reference LeviLevi, 1996; Reference PawełczyńskaPawełczyńska, 1979). These accounts lend support to David Sloan Wilson’s argument (Reference Wilson2016) to the effect that people cooperate not only for self-interest, but because they are genuinely concerned about the well-being of others. Many value behaving ethically for its own sake. Contributing to the success of a joint project, which benefits one’s group, even at a personal cost, is tied to stronger feelings of self-respect and personal integrity. Altruism can evoke a sense of pride and even elation, as the account of our qualitative study in Section 4.4 shows.

In Altruism: The Power of Compassion to Change Yourself and the World, Matthieu Reference RicardRicard (2015) traces the evolution of altruism across six million years. He argues against Dawkins who insisted that ‘universal love and the welfare of the species as a whole are concepts that simply do not make evolutionary sense. [We must therefore] teach our children altruism, for we cannot expect it to be part of their biological nature’ (1976: 2, 150). According to Ricard, Dawkins overlooks that humans are so driven to help others that they often do so even when altruistic activities generate distressing emotions.

Reference Bowles and GintisSamuel Bowles and Herbert Gintis (2013) pursue a similar line of reasoning in A Cooperative Species: Human Reciprocity and Its Evolution. Since cooperation was so crucial for survival, humans evolved the capacity for suppressing short-sighted selfish interest through developing ethical norms. Communities created institutions to protect the generous from being exploited by the selfish. The evolution of feelings such as guilt and shame underpinned our capacity for turning social norms into personal ideals. Groups with culture effective at promoting such processes would typically win in contests against groups marked by greater selfishness.

There are other strains of coevolution-inspired studies that link human well-being to our innate desire for fairness and freedom. In The Fair Society (2011) and Superorganism (2023), Peter Corning argues that we need what he calls ‘a new bio-social contract’ that assures human survival in the age of socio-environmental crises. In order to flourish, humanity needs to reduce inequality via redistribution of resources and creating fair forms of social organization which rest on the principles of equality, equity, and reciprocity.

There is one more determinant of well-being – elaborated in Section 4 – which has been illuminated in the work of the Harvard psychologist Jerome Reference BrunerBruner (1990, Reference Bruner1996). Bruner’s cultural psychology challenged the behaviourist paradigm and biological determinism dominant in cognitive psychology of the 1950s and early 1960s. More importantly in the context of our study, it points to a close relationship between human self-realization, cultural narratives, and the search for meaning.Footnote ¹² Culture is ‘the way of life and through that we construct, negotiate, institutionalize, and finally (after it’s all settled) end up calling “reality” to comfort ourselves’ (Reference BrunerBruner, 1996: 87). Narratives that we spin to order and interpret the confusing world out there do not emerge in a vacuum. Tales are powerful psychological tools that come from our cultures and create paths to our well-being or to desolation and distress. Our well-being is thus a function of culturally entrenched stories we tell about ourselves and about others. According to Bruner, our immediate experience – and the meaning of our life – is framed in a narrative way. Negatively charged or constraining stories that we repeat to ourselves can make us feel lost and helpless. Our unhappiness may thus stem from a narrative ‘neurosis’ – a reflection of an insufficient, incomplete, or inappropriate story about ourselves. Empowering stories turn us into agents of our fate, guide us on a path to meaning and direction of our lives that, in turn, make us prosper and flourish.

The interaction between of a meaning-saturated culture and individual well-being is a complex process. On the one hand, the meaning we derive from our shared stories ties our groups together (Reference LarsenLarsen, 2020). Thus, the narratives that guide us need to be shared to offer adaptive advantage. On the other hand, meaning is as much a part of the collective commons as it is shaped by unique individuals. The two do not always chime. When what we do or experience deviates from dominant cultural norms or stories, our well-being may suffer. Individuals who derive meaning from narratives and practices that cannot be synchronized with the worldview of their community, risk becoming branded as outcasts or heretics (it is enough to think of such famous cases as Socrates, Giordano Bruno, or Spinoza). That said, cultural norms and values are hardly ever switched off. On the contrary, even those individuals who defy traditions testify to the potency of entrenched habits of the heart and mind. The narrative construction of human well-being as intricately tied to meaning presents an ongoing and understudied challenge to positive psychology.

To sum up: Human biology makes up the foundation for how we experience well-being, but affects that are triggered by our prosocial behaviour are mediated through culture. Most individuals, who live up to the ideals of the cultural scripts into which we have been culturized, feel that their lives have meaning. The question is: What if stories cherished by the community makes it passive or reinforce disempowerment, since happiness and well-being are determined by God, emperor, or vengeful spirits? How to reconcile mobilizing stories that impel people to actively improve their lives with the entrenched, value-charged narratives that offer security in a chaotic world, even at the cost of individual happiness or freedom?

At the beginning of the new millennium, there has been a wealth of studies attempting to answer this question. In Born to Be Good: The Science of a Meaningful Life (Reference KeltnerKeltner, 2009), kindness to others and to oneself is the key to a happy life. In Spiritual Evolution: How We Are Wired for Faith, Hope, and Love (Reference VaillantVaillant, 2009), the cultivation of hope and love ensures that humans will evolve into more caring and spiritual beings. This is not just empty rhetoric.Footnote ¹³ In Together: The Rituals, Pleasures, and Politics of Cooperation, Richard Reference SennettSennett (2013) urges that we reclaim and redesign our cooperative skills. Today, we share space and institutions with people of different cultures, ethnicities, classes, and religions. Such diversity requires novel approaches to the challenges of prosociality. To minimize the risk of tribalism undermining everyone’s well-being, we must learn to listen and debate rather than fight and squabble. Perfecting the art of cooperation to meet the challenges of our time, Sennett argues, is a crucial condition not only for human well-being but also for saving the planet.

The human search for happiness is like a river that never dries out. Even in situations that Hannah Reference ArendtArendt (1951) defined as the fruit of ‘radical evil’, dreams of well-being, however unattainable, do not die. Similarly, Reference FranklViktor Frankl (1946), an Auschwitz survivor, and the author of Man’s Search for Meaning, argued that self-transcendence in the act of helping others was often crucial for survival. In the most gruesome of circumstances, Frankl witnessed how people derived a minuscule amount of well-being from religion or brave altruistic acts that kept them going and endowed their life-in-death with meaning. As Nietzsche wrote, ‘He who has a why to live can bear almost any how.’

In 2022, we interviewed a North Korean dissident who told us that, as a child, he had been brainwashed into believing that Kim Il Sung was God and that ‘it was completely wrong and punishable to help one another’. After his parents had eloped to China, abandoning him to a life on the street, our North Korean informant still experienced occasional acts of kindness and compassion, such as a piece of bread being thrown to him by a fellow brother under the cover of night.

Psychologists who study patients with profound traumas argue that meaning cannot be generated from talking or argumentative thinking, only from actions and strong beliefs. In times of unspeakable horror, humans search for a modicum of well-being in selfless acts, turning to religion, or confirming one’s humanity via mental or spiritual exercises (Reference HillmanHillman, 1994). After the war, Frankl used his experiences in the extermination camps to create a meaning-making logotherapy, a method for helping severely traumatized patients reclaim the purpose of their lives and which could be used to imagine and build a livable, post-apocalyptic future.

3.4 David Sloan Wilson’s Prosociality: A Key to Planetary Well-Being

One of the most interesting, if controversial, contributions of evolutionary science to the study of human well-being has been made by David Sloan Wilson’s ‘ProSocial World’ programme, a bold, research-based and practice-oriented agenda that promotes ‘productive, equitable, and collaborative groups’.Footnote ¹⁴ The ProSocial programme is based on countless publications, conversations with world sages such as His Holiness the Dalai Lama, and concrete educational projects at local schools, playgrounds, and parks. The programme is as adventurous as it is breathtaking. Although Wilson prefers to talk about prosociality rather than well-being, it is clear that he treats the collaborative and altruistic ethos as a proxy of – and the path to – human flourishing. Genetics may have revolutionized our understanding of evolution in the twentieth century, but in our crisis-ridden times, Wilson contends, we need to push evolution beyond its genetic basis. We must reinforce evolutionary insights with historical and cultural research to find new ways of social improvement at both the local and global levels.

Relying on a perspective of multilevel selection, Wilson acknowledges that evolution on the individual level favours self-preservatory selfishness. At the same time, he shows that in the broader context, investing only in competitive winners can be counterproductive. Prioritizing exceptional individuals could undermine productivity and well-being in a variety of spheres, from animal breeding to the functioning of complex societies. Both in animal and human environments, having too many rapacious and self-centred individuals tears apart the social fabric upon which everyone relies.

Predictably, Wilson rejects Homo economicus as hardly conducive to fostering human flourishing. While not denying that happiness may spring from the fulfilment of one’s selfish desires, Wilson’s ProSocial framework is based on a narrative about human well-being as anchored in altruist and collaborative work, as well as in innovation and creativity. More importantly, in terms of policy implications, evolution is not top-down design but bottom-up adaptation and experimentation, as is the norm in biological systems. In Wilson’s amplified biocultural perspective, important drivers of social change and advancement are small businesses, civic associations, localities, and nonprofits. In contrast to big, overly bureaucratized structures, small groups incentivize groundbreaking innovation and promote what works.

Wilson’s biocultural perspective has been the subject of a lively controversy. It has been assaulted as being utopian, based on incomplete evidence, and involving methodological hazards in transmuting evolutionary biology into social science. Yet historical evidence – especially that furnished with Peter Turchin’s data collected by the Seshat programmeFootnote ¹⁵ – shows how the drive towards selfishness and sectarian tribalism in history has often resulted in a war of all against all, produced traumas, and, ultimately, social collapse. Similarly, Homo economicus may win against other members of their own tribe, but the selfish tribe mostly loses to cooperative groups which are more open to creativity and altruism.

In This View of Life: Completing the Darwinian Evolution, Reference WilsonWilson (2019) presents a progressive vision of human complexity increasing over the aeons, as bands become clans, clans become tribes, and tribes become nations. Inspired by the now largely forgotten ideas of Pierre Teilhard de Chardin, the twentieth-century evolutionary biologist, Jesuit priest, and philosopher, Wilson contends that evolution includes expansion of human consciousness towards an ‘Omega Point’. The omega point of human consciousness gestures towards a moral community of people working for the good of the planet. That is the ultimate goal of the ProSocial programme: to create and refine a compelling, science-based vision – complete with cultural and educational tools and political strategies – to make humans better stewards of their own well-being and the well-being of Mother Earth. In short, This View of Life frames the evolutionary worldview as an ethical, even Promethean, project.

This Prometheanism is not without risk. Viewing evolutionary sciences as a key to what Wilson calls ‘managing the future’ through mastery of evolutionary processes (Reference Wilson, Hayes, Biglan and EmbryWilson et al., 2014: 416) carries a danger of making evolutionary science into a tool of social engineering. Digital modernity – with its focus on AI, the biological hacking of humans, and a line of other innovations associated with the Fourth Industrial Revolution – is crammed with ‘managers of the future’. The men-like-gods hubris is nothing new, but the tech-heroes of the twenty-first century have unprecedented power to pursue a seemingly rational and benign, yet potentially perilous, scenario of remodelling imperfect humans into happy Frankenstein monsters (Reference BostromBostrom, 2019; Reference HarariHarari, 2016).

4.1 An Evolutionary Recipe for a Fulfilling Life

Since our well-being system rewards success related to individual selection with a positional bias – that is, in a relative manner – who we view as our comparison group is of crucial importance. Our forager ancestors compared themselves to a few dozen in-group members, a couple of hundred at the most. Individuals competed for a dozen or two potential mates.

For agriculturalists, too, small comparison groups were the norm until modern times. Cities give us thousands of other individuals to compete against and select between. Mass media exposes us to the successes of people we will never meet, but whose luxury goods, high-value partners, and exciting lifestyles we can learn of in painstaking detail. Social media lets us covet the successes of the world’s most accomplished people. Our comparison group can now comprise billions.

Envy motivated our ancestors to try harder to match their neighbours’ achievements. Many succeeded. Today, 99+ per cent of people have a miniscule chance of becoming glamorous billionaires with tens of millions of social media followers. With the bar raised sky high, envy and depression can throw us into bottomless pits of despair rather than be a helpful kick in the butt. As mentioned earlier, social media – which can be an arena of compulsive comparisons and the narcissistic focus on the self – seems to explain, at least in part, the past decade’s rise in youth depression (Reference Foa and MounkFoa & Mounk, 2019; Reference Hellevik and HellevikHellevik & Hellevik, 2021).

Cultivating an attitude of gratitude can function as an antidote to comparing ourselves to more successful individuals. By being grateful for what we already have, we shift focus from our comparison group – and from what we lack – to what we can cherish and what makes us unique (Reference Emmons and McCalloughEmmons & McCallough, 2003).Footnote ¹⁷ Receiving gratitude can have a similar effect. Our inherent positional bias can be modified by engaging in altruistic activities. Volunteering often involves interaction with people who are worse off than ourselves. Research shows that being confronted with the suffering of others can make altruists recalibrate their comparison group in a manner that generates greater well-being (Reference Strack, Schwarz, Kern and WagnerStrack et al., 1990). According to Reference Hill, Buss, Eid and LarsenHill and Buss (2008), people could increase their well-being through reducing their exposure to comparison-inducing media and relaxing their fixation on high-performing individuals.

As argued earlier, many ancient sources and modern positive psychologists agree that happiness is not a destination point. It is a journey. It is an intermittent reward that tells us that we are making solid progress. Theoretically, the secret of a happy life is to make gradual progress, reaching as high as we can, maintaining modest goals. Nesse emphasizes the considerable influence people have on their own well-being through interpreting events and negotiating expectations. Psychological dispositions can drive ill-being, but so do ambitions. A key to happiness is to choose ambitions that match your talents. If you pride yourself on being overly ambitious in life, you may achieve more, but the price you are likely to pay is increased ill-being. If you adjust your goals so that they better match your talents and resources, you will probably feel better. Life is good, writes Reference Nesse, Huppert, Baylis and KeverneNesse (2005),

4.2 A Multilevel Selection Model for Well-Being

This section of the Element – inspired by works cited earlier – originated as a response to Buss and Nesse’s calls for an evolutionary approach to well-being (Reference BussBuss, 2000; Reference Hill, Buss, Eid and LarsenHill & Buss, 2008; Reference Nesse, Huppert, Baylis and KeverneNesse, 2005). Combining their studies of happiness with Baumeister’s work on meaning allows us to synthesize the insights of the well-being field under an umbrella of multilevel selection (MLS). The MLS perspective highlights how human evolution has occurred under two often-conflicting pressures: individual and group selection. Individuals are incentivized to be selfish to outcompete in-group members. Yet the fact that that group members also had to pull in the same direction was crucial throughout the genus Homo’s evolutionary past, which was marked by inter-group competition and war. These insights were summed up by David Sloan Wilson and Edward O. Wilson in a now-famous aphorism: ‘Selfishness beats altruism within groups. Altruistic groups beat selfish groups. Everything else is commentary’ (Reference Wilson2007: 46).

We propose that it can be profitable to think of human well-being as having evolved to affectively motivate individuals to contribute to their own success and that of their group. If we approached happiness and meaning as unconnected, we would overlook how sensations of well-being help individuals manoeuvre between the potentially conflicting pressures of individual and group selection. We sum up our approach with the following equation: ‘Happiness + Meaning = Well-Being’.

The terms ‘happiness’ and ‘meaning’ are used with a variety of definitions in scholarly and popular contexts. As argued earlier, as concepts, they have different connotations depending on language and culture. Within positive psychology, there has been considerable discussion around which specific affects contribute to happiness as opposed to pride, satisfaction, meaning, and other proposed components of well-being. Our model is agnostic in this regard. We do not consider it fruitful to try to compile lists of affects that should be associated with happiness and meaning, respectively. Such lists would vary between cultures. Moreover, a multitude of semantic disagreements over how certain terms for affects should be understood has little explanatory value. In line with our MLS model, ‘happiness’ can be defined as comprising the cluster of affects that promotes individual selection. ‘Meaning’, on the other hand, can be taken as referring to an amalgam of emotions and cognitions that promote group selection.

There is considerable overlap between the happiness and meaning clusters. Winning in-group contests can be experienced as living up to cultural ideals. Quite often, we solve adaptively relevant challenges for ourselves while also contributing to our group. The biocultural mechanisms of this well-being system are far too complex to untangle, as are its specific sources. Happiness has a stronger biological foundation, but it cannot be accessed without cultural mediation. Meaning is predominantly cultural, but it is made possible by universal predispositions (Reference Baumeister, Vohs, Aaker and GarbinskyBaumeister et al., 2013; Reference RicardRicard, 2015).

Precisely which affects motivate people to different actions is outside the purview of our model. To understand how sensations of well-being propel adaptation on individual and group levels, we need not philosophize with regard to what well-being should be or which affects or virtues it consists of. Nature and culture have already negotiated this content within each individual, group, and moral community – with varying degrees of functionality. Understanding how a distinct culture motivates certain behaviours with regard to meaning and happiness calls for deep insights into a community’s history. But from our MLS perspective, to comprehend well-being itself we need not opine on ontology and semantics, a practice that is inevitably culturally biased. It is sufficient to view well-being as a biocultural phenomenon that makes people feel ‘good’ – or ‘right’ – in a manner that motivates them to continue the behaviour that has generated well-being.

We cannot expect the well-being system to function precisely and consistently. It will not always trigger an adaptive response. Evolution shaped our affects so that they should, on average, motivate behaviours that tend to maximize reproductive success (Reference Nesse, Huppert, Baylis and KeverneNesse, 2005). Emotions will regularly arise from misunderstandings or motivate a dysfunctional response. In the modern world, environmental mismatch further undermines the precision of our emotional responses. Our MLS approach therefore cannot prescribe in detail how to make populations or individuals flourish, nor can any other perspective on well-being. People and their communities are far too complex and diverse. Understanding sensations of well-being as contributing to multilevel selection has theoretical value, but our model’s practical utility comes from combining the MLS perspective with Wilson’s work on prosociality (Reference Atkins, Wilson and HayesAtkins et al., 2019; Reference WilsonWilson, 2015). This fusion lets us illuminate the distinction between the two forms of social behaviour: competitive and cooperative/prosocial (Figure 1).

Figure 1 This multilevel selection model for well-being attempts to remedy positive psychology’s Western-centrism and conceptual overabundance while drawing attention to the more accessible, prosocial aspects of enhancing the quality of life. Happiness pursuits can be both competitive and prosocial. Successful meaning pursuits are prosocial, that is, beneficial for all involved.

While cooperation is a win-win behaviour, prosocial acts can also be altruistic, that is, win-intentional lose (competitive behaviour is win-lose). Oxford Reference defines ‘prosocial behaviour’ as helping others, altruism, or acts meant to promote the interests of society. Figure 1 illustrates how human well-being is derived from competitive (red) and prosocial (green) sources. Meaning quests are prosocial, as their actions are designed to directly benefit others. In spite of altruism being conceptualized as win-intentional lose, the altruist typically receives affective rewards and can make gains as an individual in an indirect way (reproductive, reputational, etc.). Happiness quests can involve both competitive and prosocial behaviours. Solving adaptively relevant challenges benefits the individual, but because humans are a social species, we regularly cooperate with allies to achieve individual goals.

The adaptive benefit of sociality (Reference Lewis, Al-Shawaf, Russell, Buss and DemirLewis et al., 2015), as well as indirect fitness concerns (centred on family and kin), allows us to reap happiness from spending time with our immediate circles of family and friends, and also helping them with and appreciating their accomplishments. Such activities are conceptualized as non-competitive sources of well-being. For many people, the most accessible happiness derives from the adaptive functions of mating, friendship, kinship, and coalition (Reference BussBuss, 2000). Compared to in-group competition, investing in these activities is more likely to pay off in terms of a higher quality of life. Naturally, individuals cannot opt out of solving important adaptive challenges. But when strategizing as to how to use one’s resources to improve personal well-being, it can be the wiser choice to focus on the prosocial mainsprings of human flourishing, such as being social and working for the benefit of others.

4.3 Problematizing Meaning

Baumeister (Reference BaumeisterBaumeister, 2005; Reference Baumeister, Vohs, Aaker and GarbinskyBaumeister et al., 2013) conceptualizes happiness and meaning in a similar way to how we do in our model, but he does not invoke the multilevel selection framework. Instead of viewing partially discordant demands for individual and group success as evolutionary pressures that must be navigated, he offers a moral distinction. He connects meaning and happiness, respectively, to giving and taking. When we compete as individuals, our gains are often at the expense of others. When we contribute altruistically, we give to members of our community, a practice which Baumeister values higher than competition.

Animals are mostly incentivized to help kin. Reference Warneken and TomaselloWarneken and Tomasello (2009) place the roots of human altruism – in the form of instrumental helping – with our common human ancestry with chimpanzees. Yet chimp prosociality mostly limits itself to close kin, small hunting groups, and boundary enlargement. Other social behaviours tend to be highly competitive; chimps seem not to care much for the well-being of non-kin (Reference Silk, Brosnan and VonkSilk et al., 2005; Reference WranghamWrangham, 2019a, Reference Wrangham2019b). With the evolution of imaginative culture – around 70,000 years ago (Reference BellahBellah, 2011; Reference Dawkins and WongDawkins & Wong, 2016; Reference HarariHarari, 2014)Footnote ¹⁸ – Homo sapiens became able to extend natural predispositions for nurture to non-kin, and even strangers.

Expanding our in-groups to include groups of non-kin was a great evolutionary leap (Reference WilsonWilson, 2007). Such prosociality, boosted by cultural norms and ideals, made large-scale sociality possible. Human groups would not have been able to grow beyond our species’ Dunbar number of around 150 interpersonal relationships if culture did not facilitate impersonal prosociality. We would have remained an insignificant ape (Reference HarariHarari, 2014). Our capacity for meaning allowed us to override self-interest, writes Reference BaumeisterBaumeister (2005: 135), as ‘nature makes each creature selfish, but culture functions best if people will sometimes set aside their selfish wishes and impulses in order to do what is best for everyone, that is, for the collective’.

Our ancestors’ newfound capacity for uniting around cultural values facilitated prosociality by turning what benefits the group into intrinsic motivation for members via our well-being system. Because we want to feel good, we are driven to do what cultural scripts compel us to do, as this is emotionally rewarding. The well-being we can derive from meaning pursuits can be much greater – and longer lasting – than happiness pursuits can offer. Individuals can be convinced to entirely disregard their own fitness. Self-sacrificing soldiers and suicide bombers exemplify how culture can trigger such intense affect that individuals become devoted actors who eagerly sacrifice their lives for their group and its abstract values (Reference Atran, Sheikh and GomezAtran et al., 2014). Their reward is being suffused by a deep sense of meaning in this act of self-immolation. The evolutionary rationale behind such overwhelming experiences seems to be to impel individuals to place group needs first.

The potency of meaning is closely related to the feeling of self-esteem, which is a crucial component of well-being (Reference Kirkpatrick and NavarreteKirkpatrick & Navarrete, 2010; Reference Solomon, Greenberg and PyszczynskiSolomon et al., 2000). Well-socialized people feel good about themselves to the extent that they live up to cultural ideals. When in-group members put us in higher esteem, we experience our elevation as gratifying. When they look less favourably on us, we are distressed. Self-esteem is adaptive because it results from, and reinforces, social belonging, which offers powerful advantages in terms of survival and reproduction (Reference BaumeisterBaumeister, 2005).

As our communities grew from dozens, to hundreds, to thousands, to millions of individuals, our ancestors had to craft new forms of cooperation that motivated large-scale buy-in. To make people tie their destiny to larger numbers, they expanded their circles of prosociality. Reference DarwinDarwin (1871) considered this to be ‘the noblest part of our nature’. He stressed Homo sapiens’ ability to extend our nurturing instinct to larger circles, from offspring and kin to social groups, and nations, and even other species. When some modern humans are able to view humanity itself as their in-group, they can derive meaning – and thus well-being – from helping faceless strangers in far-away places.

Baumeister draws a temporal axis for how meaning functions differently to happiness. Happiness relates to our wants and needs in the present. Meaning integrates our past with the present and future. We interpret our experiences in the light of cultural values. If we can understand our past in a way that empowers us with regard to the future, past distress – or even trauma – can be infused with meaning. If we anchor our present-day actions in an imagined value-charged future outcome, what we do feels meaningful. The longer the timescale, the deeper the sense of meaning. This mechanism can motivate enormous sacrifices for a utopian community that lies decades – or generations – ahead, or in the afterlife.

Meaning pursuits can also turn dysfunctional, even extremely so. It may feel deeply meaningful to submit to a radical political ideology, a religious sect, or another creed. ‘Altruistic pathologies’ underpinned many of humanity’s greatest follies and atrocities. Similar to the ways in which the pursuit of happiness through individual success can be taken too far – to individual and communal detriment – pursuing meaning through personal sacrifice has the potential to ruin lives. In their work on pathologies of altruism, Reference Oakley, Knafo, Madhavan and WilsonOakley et al. (2011) examine an array of domains in which our meaning system can become corrupted and lead the altruist astray.

Baumeister does not focus on these potential pitfalls. He takes a normative stand for meaning over happiness, concluding that

4.4 Happiness + Meaning = Well-Being

Quantitative studies establish valuable correlations between life factors but fall short of explaining these mechanisms at a deeper level (Reference Hui, Ng, Berzaghi, Cunningham-Amos and KoganHui et al., 2020; Reference Moche and VästfjällMoche & Västfjäll, 2021). Experimenting has been of limited utility in this pursuit (Reference Charness and GrosskopfCharness & Grosskopf, 2001; Reference Konow and EarleyKonow & Earley, 2002). Reference Meier and StutzerMeier and Stutzer (2008) explain the suboptimal outcome of laboratory studies by how such stimuli provide too meagre stakes to influence reported life satisfaction. Our MLS model advances an approach to the connection between prosociality and individual and social flourishing which offers several advantages. First, it makes a distinction between happiness and meaning in order to highlight different functions of individual and group selection in forging human well-being. Second, it relies on a synergy of quantitative, qualitative, and historically based studies. Third, it attempts to flesh out both cultural and universal elements of human well-being.

In this section, we will illustrate the workings of our MLS model through a study we conducted in 2022, in the aftermath of Putin’s invasion of Ukraine (Reference Larsen and WitoszekLarsen & Witoszek, 2023; Reference Larsen, Witoszek and YeungLarsen et al., 2023). The informants in our qualitative study were volunteers (n = 32) who helped Ukrainian refugees in Norway. By selecting dedicated altruists, we gained access to thick descriptions of prosocial motivation and experience. The collected insights let us examine the specifics of the human well-being system. Our main concern was to understand the ways in which altruistic activities contribute to the well-being of culturally diverse volunteers. Can we talk about cultural varieties of altruism as practiced by Polish, Russian, Ukrainian, and Norwegian helpers? What narratives, root metaphors, and images did they use to communicate their experiences?

Many Polish volunteers felt that the Ukrainian war was also their war, so that they had a particular responsibility to contribute. Their strong engagement infused their lives with meaning, guiding their actions during a difficult time, and sending them off on an exciting, though unpredictable, journey. Many informants felt empowered, expanded their social circles, experienced personal growth, and became eager to advocate the benefits of altruism. A Ukrainian woman said, ‘What is happening to me now is like a whole new life. I was someone who could not make any decisions on their own or fix things. I needed help. But not anymore. I feel like I’m bigger on the inside, that I have room for more people.’

Experiencing altruistic reward, several Slavonic informants bemoaned how volunteering had atrophied in their native countries. A Ukrainian man said, ‘Working together is the only way to build trust. For now, in Russia and Ukraine, people generally do not trust the government, police, doctors – it’s a sad life.’ A Belarusian man said, ‘We fear scams, spies, propaganda. We want to help, but we have to focus on our own house. I cannot tell my friends in Belarus that I do voluntary work now. They would think I was a political activist only out to please the party to earn money for myself.’

Meaning-generating activities do not necessarily trigger positive affect in the present (Reference Dakin, Laham, Tan and BastianDakin et al., 2021). Certain forms of altruism do produce moments of elation, but meaning seems, to a greater extent than happiness, to induce a lasting sense of well-being. Often, this is more a process than a moment. Such a process can involve a slow recognition of one’s own potential and sometimes a discovery of hidden talents. Many of our Slavonic informants reported feeling distressed and emotionally drained due to identification with the war’s victims. In the long run, however, they experienced a more persisting sense of purpose and sense to their lives. ‘Nothing is more meaningful than helping others to restore their dignity and humanity,’ as Polish volunteer Aleksandra put it.

The Belarusian informant wanted his volunteering to benefit his new and former community. Slavonic nations should copy Nordic practices to receive similar advantages, he suggested: ‘Norwegians are very naïve, thinking everyone is honest like them. But we must choose to be optimistic about the future, we must risk being naïve. We are all in the same boat. A good future requires that we cooperate. I have a 100-year perspective on helping. I want a better world for my children.’

One of the intriguing findings in our project had to do with different, culturally determined styles and strategies of altruism. The Norwegian helpers followed bureaucratic rules and acted in accordance with the official aid codex. Slavonic volunteers tended to work around the system, operated at breakneck speed, and initiated projects that defied existing protocols. It may well be that the Slavonic informants – with their history of anti-communist and anti-state ethos – were motivated by what we term anti-systemic altruism, that is, being a good person who challenges an oppressive system. This ethos highlights spontaneity, improvisation, and frequent rule-breaking. In contrast, Norwegian systemic altruism has always been based on trust, efficacy, and compliance with institutional requirements.

These cultural differences notwithstanding, for most informants, helping the Ukrainian refugees was more than a feel-good experience. It was often a transformative process which made them discover their hidden potential, even reclaim self-respect. As Polish volunteer Hanna put it:

These qualitative interviews illuminate aspects of our well-being system that an MLS perspective helps make sense of. As individuals, we are doomed to strive in status contests, but our communities are also dependent on our altruistic contributions. Experiences of happiness and meaning help us respond to these contradictory pressures and navigate between them.

One key to the relative success of Nordic prosociality is aligning what benefits the individual with what strengthens the community. Several informants stressed how Nordic egalitarianism, in combination with jobs that impart a sense of communal contribution, was important for their high quality of life. When success at work also feels like altruism, the result is an amalgam of happiness and meaning, which has an amplifying effect on long-term well-being. For those without such professional positions, and for many retirees, volunteering infuses their lives with meaning and significantly enhances their quality of life. Several informants reported developing a reservoir of meaning to draw well-being from during times when it is harder to achieve happiness.

Since the happiness side of our equation is mostly relative, policy aimed at enhancing meaning has the greatest potential for increasing societal well-being – especially in prosperous countries, in which people’s basic material needs have been covered. This conclusion is supported by studies that attest to an intriguing trade-off between happiness and meaning. Some global surveys show that when GDP per capita goes up, on average, happiness increases at almost the exact same rate that meaning diminishes (Reference Oishi and DienerOishi & Diener, 2014). One possible explanation behind this relationship could be that when nations do better economically, communal need diminishes, reducing people’s access to meaning-providing activities.

This remarkable stability speaks to the relative nature of our well-being. Still, variance between cultures attests to how some environments, like the Nordic ones, are systemically programmed to enhance human flourishing. After gaining happiness from economic growth, finding new ways to promote activities that generate meaning seems to be the most productive way forward. The central role that helping refugees now has among a growing number of Norwegian voluntary groups lends support to the claim that Norwegians draw part of their high well-being from looking after exceptionally disadvantaged groups. Research substantiates that exposure to those who are worse off makes people appreciate their own life more (Reference Strack, Schwarz, Kern and WagnerStrack et al., 1990). Our MLS perspective illuminates why such dynamics can make volunteering a win-win activity, for the beneficiary, the altruist, and the community as a whole.

5.1 Nature as a Space of Renewal

There is yet another important factor behind high levels of well-being in the Nordic countries: access to – and interaction with – pristine nature, our evolutionary habitat. Scandinavian nature is not just about the ecological environment; it is linked to an array of practices and values based on interaction between humans and the natural world. The most significant Norwegian icons, narratives, and rites – which have empowered and cemented the national community over time –include images, stories, and modes of conduct related to nature (Reference AnkerAnker, 2022; Reference LarsenLarsen, 2023; Reed & Rothenberg, 1992; Reference WitoszekWitoszek, 2011).

From Norse poetry to Næss’ eco-philosophy, nature has been deployed as a locus of belonging and an emblem of national identity. There is a remarkable consistency in the way nature-inspired tropes and images have been linked to projects which have advanced social justice, equality, peace, and cooperation. Studies on the origins of social democracy point to how the older generations that built ‘Happy Norway’ had their identity anchored in nature as an ancestral home, the sacred place to regenerate, to cultivate friendship, and to carry out projects requiring collective mobilization for the welfare of others.

In most Nordic cultures, nature use has been a social glue cementing trust between diverse social groups. A variety of activities – from the obligatory Sunday hiking tour, to long visits at summer and winter cottages, and cross-country skiing – have nourished national health, longevity, and well-being. Until recently, Norwegian happiness was intimately connected to the open air, or joys and rituals of friluftsliv. In spite of the growing indignation about the Norwegian oil industry as one of the planet’s largest polluters, most Norwegians have embraced a cognitive dissonance that allows them to combine their high ecological footprint with their ideal of a good life linked to the idyll of nature. They did so en masse until the second decade of the twenty-first century when new technology and culture brought new challenges (see Section 5.2).

Two additional elements are inherent in modern Nordic well-being societies. First, the idea of self-realization fully extends to all genders. Nordic productivity is in part due to having one of the highest female employment rates (Reference Witoszek and MidttunWitoszek & Midttun, 2018). The cultural ideal of gender partnership rather than power struggle or machismo accounts for increased well-being for women and men alike. Unlike the more liberal Anglo-Saxon feminism, which is more centred on the individual self, the Nordic model educates men and women to value partnership and teamwork, which affects relationships at home, at the workplace, and in civil society.

The Nordic well-being society is not underpinned merely by economic and political concerns. It is supported by a whole happiness lexicon consisting of words and memes like the Danish hygge, the Norwegian kos, and the Swedish lagom – to mention the most popular examples. These concepts have no exact translations in English, but evoke coziness, harmony, balance, security, and at-homeness in the world – not grandeur or splendour but kos; no drama, just de-stressing at a relaxed pace. Or Swedish lagom, meaning just the right amount, not too much or too little.

Implicit in hygge or kos is the idea that human well-being is not just about relaxing after winning a palatial place with a pool and a Jaguar parked in the garage. Kos and hygge are about relishing simple pleasures which have little or nothing to do with material gains. For instance, in Norway, the intense experience of kos can refer to touchingly trivial pursuits, like rewarding oneself with a piece of chocolate, a biscuit, or an orange after climbing to the top of a slope. Little matches the calm ecstasy of sitting atop a mountain with a magnificent view, and slowly peeling one’s well-deserved orange.

Policy-wise, to promote the community of kos, one could aim for shorter work weeks, longer vacations, and subsidized day care, as well as investment in arenas that enable people to come together to solve concrete tasks of communal importance – like building a new playground – or just to enjoy each other’s company.

The Nordics exemplify our model of well-being as an amalgam of happiness and meaning. While accepting that there are many paths to individual self-realization, Nordic culture promotes a model of education, and a Bildung, which associates well-being with humanitarian projects at home and abroad. For the regular Scandinavian, life becomes imbued with meaning when individual happiness is linked to work for the common good and helping the needy, wherever they may be.

5.2 The Nordic Model under Siege: Twenty-First-Century Challenges to Human Flourishing

Several intriguing – if not disturbing – features of the transition of Norway to a well-being society have been signalled by international indexes and national research findings. From 2018 to 2023, Norway fell from being the happiest society in the World Happiness Report to number 7. The cause seems primarily to be a marked decrease of well-being among the young.

In surveys from the 1980s on, 15- to 24-year-olds were mostly the happiest while the oldest generation were the least happy. In the period 2009–2019, this relationship was inverted (Reference Hellevik and HellevikHellevik & Hellevik, 2021). The Norwegian Monitor’s quantitative study concluded that the main sources of youth ill-being were loneliness, dissatisfaction with health and physical fitness, poor relationships with family and friends, dissatisfaction with gender equality, worries about the future, and high levels of stress related to the imperative to succeed and excel.

Our own qualitative interviews with high school students point to excessive use of social media as an important source of reduced well-being. In terms of cultural evolution, a new development seems to be underway. Sherry Reference TurkleTurkle (2012) argues that curating one’s online profiles alters the presentation of the self and, ultimately, influences individual identity. The ethos ‘I share, therefore I am’ drives people to craft their identities for others. Continuous digital performance, even if successful, often leads to disconnection anxiety or a ‘fear of missing out’, so-called FOMO.

The compulsion to compare oneself with others is a source of stress and anxiety. The have-nots suffer because they do not live up to their ideals and aspirations, while the haves suffer because of a pressure to match the have-mores. Facebook and TikTok facilitate a commodification of individual identity, incentivizing the young to present themselves as living happy, enviable lives. Paradoxically, for many young captives of the vicarious life through social media, our era’s ‘ideology of well-being’ results in greater ill-being.

Adding to the misery of the young is a growing disconnection from nature. Unlike for their parents and grandparents, friluftsliv – the ideal of regenerative communing with fjords, forests, and mountains – seems to have lost much of its appeal to the young generation. Gym workouts give quicker and more visible results than walking or skiing. Hiking in the forest or going to a summer cottage is now surveyed through the eye of Instagram and distracted by a stream of messages flowing from social media.

It is too early to conclude what consequences the ongoing digitalization and ‘nature deficit disorder’ will entail for the young members of well-being societies. What is certain is that the twenty-first-century youth will have to cope with the unpredictable sociopolitical, economic – as well as evolutionary – challenges pertaining to the age of ‘dataism’ (Reference HarariHarari, 2016).

6.1 From Play and Flow to Happiness for Sale

Competing for happiness will always be part of human life. But once the basic adaptive challenges are solved – or when the competitive paths to happiness are less attractive – individuals can be better off pursuing the prosocial sources of well-being. Alas, also these sources can be hard to access in some cultures. The practice of impersonal prosociality can be prohibitively costly in dysfunctional environments. Being overly generous to strangers in highly selfish, low-trust societies is a recipe for being taken advantage of or punished (Reference Balliet and Van LangeBalliet & Van Lange, 2013; Reference GintisGintis, 2008), which tends to reduce well-being. Any form of altruism can be risky, or impossible, in the most oppressive of regimes, such as North Korea. Tyrants and dictators fear prosociality because of its empowering, agency-building capacity. The most spectacular social transformations – from Gandhi’s peaceful resistance to the Polish Solidarity movement of the 1980s – were based on the awakening of human altruism and compassion which contributed to the collapse of empires.

There is one more source of well-being which merits attention: the human capacity for play and creativity. Homo ludens is a natural hero of the happy, joyful world whose aspects have been studied by Johan Reference HuizingaHuizinga (2016) and Mihaly Reference CsikszentmihalyiCsikszentmihalyi (2008). Huizinga points to play as both universal and autotelic; that is, it has no function beyond eliciting a spirited joy of playfulness in both human and some animal species. But, in the case of humans, the nature of play is often enriched by a creative process that defamiliarizes the familiar and leads to artistic and scientific breakthroughs. Csikszentmihalyi drew attention to the key element of play by deploying the concept of flow. He studied the mental state of ‘being in the zone’ – a peak of enchanted self-transcendence. Being in the flow is entering into a sphere of timelessness in which the ego is partly dissolved and rules feel less like a hindrance and more like an exciting challenge. Playing music, engaging in sport activities, and engaging in spiritual exercise are mentioned as mainsprings of flow. Ludic activities create a world which, at least for a moment, is immune to limitations, existential constraints, and even the trauma of war. Joys of creativity and innovation combine a sense of mastery and enchanted expectations which, in turn, increase happiness and self-realization (Reference BussBuss, 2000; Reference CsikszentmihalyiCsikszentmihalyi, 1997a, Reference Csikszentmihalyi1997b; Reference HaidtHaidt, 2006; Reference Seligman, Seligman and GillhamSeligman, 2000).

The question is: Can we learn to be more ludic and creative? Are there any steps to reaching flow – the omega point of happiness? In his bestseller Happiness: A Guide to Developing Life’s Most Important Skill (2007), Matthieu Ricard argues that happiness can be reached through a process of learning – but not without a price. That is to say, there is a ‘happiness codex’ to be followed and a skill to be honed for those who can afford to listen to Ricard, standing in his maroon and orange robes among the glitterati of the World Economic Forum in Davos. The sight of the eudaimonic guru may be inspiring to some and unsettling to others. What is inspiring is Ricard’s meteoric rise from a modest Buddhist sage to a leader of meditation and happiness sessions for industrial tycoons, Nobel-winning academics, and heads of state. What is unsettling is the realization that there is a ‘happiness market’ which commodifies our dreams and aspirations.

We live in the age in which the mining, processing, and selling of mass data is not just the fundament of ‘surveillance capitalism’ (Reference ZuboffZuboff, 2018), but the domain of marketeers. There are reasons to fear that soon neuroscience and psychology, in alliance with social economics, will have located happiness, measured it, and made it ready to be sold as part of the growing eudaimonic industry. Some critical commentators predict that the mathematical, mechanized view of the mind will triumph, simply because of the financial rewards of objectively defined happiness. ‘Positive psychologists and happiness economists make a great play of the fact that money and material possessions don’t lead to an increase in our mental wellbeing,’ declares the writer William Davies in The Guardian:

This is a disturbing reflection. While positive psychology, mindfulness, and pharmaceuticals can all potentially be helpful, a covert cultural brainwashing which fetishizes happiness as a must, whatever the cost, is dangerous both to society and to the democratic order as we know it.

6.2 Improving on the MLS Model

The MLS model of human well-being as drafted in this Element invites further work. For well-being scholars, our evolutionary approach can hopefully inspire productive lines of enquiry. To substantiate the utility of an MLS model, our empirical research mostly explored the sources of meaning and their connection to well-being among altruistic volunteers (Reference Larsen and WitoszekLarsen & Witoszek, 2023; Reference Larsen, Witoszek and YeungLarsen et al., 2023). Quantitative studies could shed further light on the array of motivations and inspirations that energize prosocial work. In what ways do other categories of altruistic actors than the ones we interviewed benefit from helping others? More in-depth qualitative studies could illuminate varieties of prosociality in other contexts than voluntary work to help refugees from countries with cultural norms in opposition to prevalent Western metrics and values.

Cross-cultural studies with a wider range than ours should offer fertile contributions to understanding the relationship between human universals and cultural and historical legacies. With regard to expanding one’s circles of prosociality, we expect there to be significant variance in scale and narrative justification of altruistic acts. Studying non-WEIRD nations and kinship societies could help us better understand the biocultural influences that inform our criteria for whom to help and what strategies are most effective.

We propose that scholars think flexibly in terms of which group levels an MLS perspective should include. Of particular interest could be the distinction between solving adaptively relevant challenges individualistically or as part of a group (Figure 2). Collaborative activities allow individuals to harvest the diverse rewards related to individual and group selection. Progressing towards goals generates happiness, while aiding other group members offers interpersonal meaning. If this were to occur under mortal threat, identity fusion can make individuals feel closer to other group members than to kin (Reference Atran, Sheikh and GomezAtran et al., 2014). Individuals who tie their destinies together – especially in challenging or perilous environments – make happiness and meaning merge in a manner that can greatly elevate well-being. There are numerous testimonies from soldiers who, after war, long to go to back to the strongly rewarding affect that their previous group activities generated in situations of intense danger.

Figure 2 When people form close-knit groups that work to solve everyone’s adaptive challenges, this can have a powerful effect on well-being. Succeeding in competition against other groups generates happiness. Reciprocal altruism between members generates meaning from interpersonal prosociality. The group can draw additional meaning from bestowing impersonal prosociality upon members of the larger community. Future research should investigate how best to facilitate activities that let people tap into multiple sources of well-being.

How a similar togetherness can be cultivated in modern, peaceful environments is not clear. Intriguingly, well-being studies from cultures that emphasize interdependent flourishing, such as the Confucian ones, report lower well-being (Reference Krys, Park and Kocimska-ZychKrys et al., 2021b; Reference Rappleye, Komatsu, Uchida, Krys and MarkusRappleye et al., 2020). Is this due to a local preference for lower emotional intensity? To what extent are these populations influenced by their existential constraints? Or does peaceful interdependence require that individuals restrain their emotions so as not to be group outliers?

We justified our equation ‘Happiness + Meaning = Well-Being’ by emphasizing the importance of viewing the human well-being system as one, affected by both individual and group selection. There is a need for further investigation of how psychological and environmental demands affect individual strategies for well-being. How people prioritize differently between sources of well-being when facing a crisis could be another productive avenue for further study. Nesse’s hypothesis, that high levels of well-being are a result of adaptive signal interpretation and goal adjustment, needs further substantiation. Qualitative research on people who have gone through crisis and goal re-evaluation could help illuminate the mechanisms of personal and social renewal.

These questions suggest some of the ways in which studies of well-being could be enriched by a biocultural MLS perspective. How does it affect the human reward system to raise the primary level of competition from us as individuals to our social group? Does it feel better to win alone or as a group? Does failing together feel less bad? What is the difference between meaning springing from altruistic activities that benefit members of our interpersonally related social group, and actions that benefit strangers? Is it more rewarding to engage in altruistic activities alone or as a group?

Figure 2 illustrates how we conceptualize these levels. Individual and social-group functionality relies on ancient cognitive mechanisms. At the level of the moral community – today, typically one’s nation – we rely on culture (Reference HenrichHenrich, 2020). With interpersonal relations, we can, to a great extent, depend on Homo sapiens’ reflex for reciprocity (Reference HaidtHaidt, 2006). To cultivate prosociality among strangers, we need well-functioning norms, values, and institutions. A deeper understanding of how interpersonal altruism, juxtaposed with impersonal altruism, affects our well-being system could be of paramount importance.

There are many themes and methodological tangles that we have only signalled in this Element, and which need further research. Methodologically, exploring the galaxy of ‘mongrel concepts’ of well-being, happiness, and meaning calls for more in-depth studies comparing what the people say in interviews or questionnaires and what they actually do. There is often a gap between words and deeds – a gap which may be a conscious attempt of redesigning the self into a more wished-for specimen, or an unconscious strategy of a ‘pilot response’ which has not been thought through. Both qualitative and quantitative studies of well-being would also profit from taking into account a dynamic, temporal, and contextual dimension of our perceptions of well-being. These perceptions do not stand still; they change over time and generate conflicting interpretations with regard to not only whether what makes us happy or unhappy in particular situations, but also who we are (or were), and where we are going with our lives.

A possible venue for more exploration has to do with the project of studying and understanding culturally sensitive definitions of well-being. Our biocultural approach is based on the assumption that human beings from all latitudes are biologically wired for harmony and well-being, just as they are wired for the search for freedom and fairness, desire for recognition or altruistic behaviour. It is cultural norms and values, which emphasize particular drives and aspirations, that make us different. The question is: do they – and to what degree?

To mention but one example of a conundrum, most Latin Americans and most Africans cherish their cultural traditions, but, when given a chance, they dream of – and vote with their feet – moving to Western countries which profess values conflicting with their indigenous legacies. Respecting the immigrants’ separate paths to development and happiness within Western societies is a huge, and unresolved, challenge in modern migration studies. Our model – emphasizing both biological commonalities and cultural differences – may be helpful in balancing more effective solutions. That said, matters are complicated by highly subjective responses to cultural challenges. Some individuals increase their well-being by embracing Western freedoms, while others refuse to be replanted and demand respect while disrespecting their host societies. It is enough to read the Somalian writer’s Ayan Hirshi Ali’s biography Infidel (2008), to see the contrasting well-being paths of two young sisters from Somalia who are replanted as immigrants in Germany. While one of them (Ayan) flourishes and spreads wings, the other, appalled and paralysed by excruciating pressures of Western freedoms, falls into deep depression and – paradoxically – dies of the excess of life choices and opportunities that life in the West has to offer.

Ayan Hirshi Ali’s story is instructive because – were their contrasting well-being trajectories transposed to the second decade of the twenty-first century – they may have taken a different turn, especially with regard to Ayan’s sister. The ongoing virtualization of Western – and immigrants’ – lifeworld encourages cultural ghettoization, for better or for worse. Not only it is now possible to happily inhabit in a different culture without speaking its language or complying with ‘oppressive’ customs of the host country; one can offer one’s offspring alternative, home-based models of remote education via internet platforms.

What also needs more research is the role of small groups as triggers or deterrents of individual well-being. On the one hand, our evolutionary past, which goes back to a life in small social groups, programs us to thrive – and effectively solve problems – in intimate settings. The hasty conclusion would be that problems that can be solved at a lower, subsidiary level should be the preferred option than involving a complex machinery. Again, matters are more complicated than that. Many individuals thrive and blossom in small, intimate communities which function as extended families and, at best, as warm circles where everybody feels safe and looked after. But, as we suggested, well-being requires more than the mere satisfying of basic needs or a handful of fleeting moments of happiness. Meaning matters in the equation. The well-being of some inhabitants of local villages is bound to be eroded by a sense of constant surveillance, by strong control mechanisms, or by group pressures on how the proper life should be lived. Cosmopolitan or urban cultures are more relaxed and tolerant of ‘deviants’, not to mention that they offer a better chance to achieve self-realization and thus increase well-being.

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Our MLS model does not offer a silver bullet to solve an array of problems bedevilling developed and developing democracies in the twenty-first century. But, as we see it, our MLS approach may have relevance in three key areas which impact social well-being and bear on the design of modern democratic institutions. The first one concerns rethinking of our caring and health services. All too often the well-being of hospital patients is misconceived as dependent on successful surgery or efficient application of medicines. All too frequently patients who have been traumatized or handicapped by serious accidents are treated as cured as soon as they seemingly recover – and left to their own devices. The system is not geared towards attending to their well-being as meaning-making and meaning-seeking individuals. The hazards of progressive bureaucratization of care services are anchored in increasing efficiency, not in helping a person to recover a meaningful life. The well-being of patients is a deep black hole in geriatric care, where many patients are treated as travellers at the end of their journey. The widespread misconception is that at this point of their lives, there is no longer any meaning to be found – apart from eating, being put to bed, and occasionally receiving visits from family or friends. If, as our model suggests, meaning is the basis of human well-being, then both the process of recovery of sick patients and the care of the elderly would need to be reimagined by health institutions, social services, and nursing homes.

Second, our biocultural approach to well-being – anchored as it is in cultural sensitivity and universal drives of human nature – has implications for current educational paradigms. Although the Council of Europe talks about ‘improving well-being at school’,Footnote ²⁵ modern educational institutions function mostly as instruments of learning and acquiring knowledge. The discussion of well-being is often reduced to health issues, and there is as yet no project of education to well-being through the pedagogy of meaning-making.

Third, individual and social well-being is often a blind zone in fragile democracies that attempt to rebuild social structures destroyed by despotic or authoritarian regimes. There are other priorities: economic growth, restoring the rule of law, ensuring a minimum welfare. It is easy to forget that stressed and distressed populations are potential victims of authoritarian populists and demagogues. To mention but one example: A country like Poland that regained its democracy in 2023 – after eight years of crypto-fascist rule that specialized in hate and social polarization – faces a demanding project of building a ‘democracy 2.0’. Such democracy cannot be reduced to restoring the rule of law and governmental welfare hand-outs that make people ‘happy’ for a day or two. It has to rely on forging new educational institutions that attend to two often-overlooked projects: (1) a broad emancipative programme that connects meaning with freedom and reclaims individual agency; and (2) a novel pedagogy boosting society’s well-being through promoting ideals of altruism and cooperative ethos. The fate of Poland – and of all ailing democracies – is as much dependent on material prosperity as on innovative, meaning-making schooling that emphasizes empowering, can-do stories rather than flaunting a passive-aggressive identity anchored in suffering and hate of the others.

To sum up: more insights into strategies to forge prosociality and cooperation as the condition of human flourishing are vital for rethinking modern democracy and its institutions. As we have indicated earlier, altruism, teamwork, as well as a sense of belonging to a caring community are dependent on close, face-to-face interactions. The age of artificial intelligence, where, increasingly, net-based transactions – from banking to education – has melted the glue that holds communities together. The functioning of e-based institutions may be cheaper, smarter, and more efficient, but the art of building cooperation, friendship, and warmth within families and between generations is eroded. The social bubbles of TikTok, Instagram, and Facebook reduce the imaginative and inspirational energy that flows from people who meet each other in the same physical space.

Although it may sound overambitious, a combination of evolutionary and cultural insights into the drivers of well-being is not just about creating more effective educational and institutional arrangements; it may contribute to humanity’s next great evolutionary leap. The twenty-first-century challenge is to craft prosocial culture to solve global problems, from the climate crisis to migration, biological hacking, nuclear proliferation, and artificial intelligence (Reference BostromBostrom, 2019; Reference HarariHarari, 2016). It has been repeated ad infinitum – by UN leaders,Footnote ²⁶ international aid organizations, and scholars – that only cooperation and prosociality can help us address the civilizational crises of our time. For people to expand their circles of empathy to the global level, transnational teamwork and solidarity have to feel good, or at least right and desirable. A synergy between positive psychology and the evolutionary sciences can help policymakers by providing insights into the relationship between impersonal altruism and our well-being system.

In the short run, interdisciplinary insights into the mechanisms of well-being could help counter the ongoing cultural and political polarization which bedevils many societies. Seen most prominently in the United States, reduced impersonal prosociality and an unwillingness to cooperate across political divides diminish political efficacy and undermine social cohesion. This cultural regression results in nationally uniting values and narratives losing their hold. As a sense of belonging to a moral community is reduced, people seem to crave stronger interpersonal relations at a smaller-group level, a process which contributes to what Reference PopperKarl Popper (2020) called the ‘retribalization of the world’.

Figure 2 shows how this process entails a devolution of moral allegiance from a higher to a lower level and an intensification of intra-group competition. Societies that turn in on themselves risk being engulfed by political chaos and self-destruction. Evolutionary well-being scholars cannot tell us which values will underpin tomorrow’s global village – if one such ‘village’ comes into being – but they can suggest which values are imperative for preventing social breakdown and ensuring our survival. The search for selfish happiness alone, without the quest for transformative meaning anchored in the prosocial ethos, is pernicious from both an evolutionary and a cultural point of view.

The stakes are significant. Combining insights from contextual, cultural studies, and evolutionary science may guide policymakers in their efforts to improve human cooperation and resilience in crisis situations. Many scientists talk about a ‘climate end game’, mapping out the potential catastrophes that could follow a ‘tipping cascade’ and trigger multiple system failures that afflict societies across the globe. Without a social mobilization based on prosociality and cooperation, humanity could face a civilizational collapse. A framework with cross-cultural predictive potential – one that helps those in charge design or reimagine their social and educational policies and evaluate outcomes – could both contribute to improving lives for more people and avert social breakdown in our challenging times.