The main subject of this paper is the way in which patterns of settlement and modes of working and holding land are transformed and adapted to diverse political and economic relations. The changes discussed occurred in adjustment, under British Imperial rule, between a large-scale, commercial landlord—initially a mining and trading company—and local rulers and residents in the area of the landlord's monopoly in north-eastern Botswana. Ecological conditions in the middle and high veld of Botswana are of crucial importance for the discussion, since these posed various risks and costs for the landlord, the local rulers, and the residents. I examine primarily a sequence of decisions and the locally crucial constraints that prevailed and set a course for further choices. ‘Tribal ruler’ is the generic term which I use for such political offices as ‘paramount chief’, ‘chieftain’, and also the Protectorate Administration's grade of Sub-Chief. A paramount chief or, simply, chief, is a ruler of the highest rank, having a major territory, whose authority extends over a series of subordinates, including some chieftains. Chieftains, in turn, are petty rulers over lesser territories, and in the grades of the Administration, never higher than Sub-Chief, though they may be independent and not under a chief as an administrative superior.