Nations and national projects are gendered in different ways. Feminist theory has raised important questions about the conceptualisation of ‘difference’. This article develops the conceptualisation of the different ways in which nations and national projects are gendered, arguing for a mid-level conceptualisation of gender relations. It argues against, on the one hand, the excessive fragmentation of gender, and on the other, too simple dichotomies of more/less unequal gender relations. This draws on a theorisation of gender relations which connects the different dimensions into specific kinds of gender regimes, either public or domestic gender regimes. This enables us to conceptualise different national projects as having a more or less public or domestic gender project. The conflicts between different national projects and with other polities, such as states, are then conflicts between differently gendered projects. The usefulness of this mid-level conceptualisation is demonstrated through examples of the competing relations between the UK, Ireland, the EU and the Catholic Church in a global era.