One hundred and twenty-eight years ago in the historic city of Oxford a relatively brief impromptu verbal exchange at a scientific convention occurred. It is still vividly remembered in and out of academia. This so-called ‘debate’ between the Bishop of Oxford, Samuel Wilberforce, and the young scientist, Thomas Henry Huxley, a simple and concrete episode, has continued to symbolize dramatically the complex and abstract phenomenon of the conflict between science and religion in the late nineteenth century. while that symbol may be somewhat inaccurate, or its relevance may have shifted from a century ago, it still is a powerful image, one which continues to be an important part of the religious, scientific and rhetorical history of the late Victorian era. Moore recently wrote: ‘No battle of the nineteenth century, save Waterloo, is better known.’ It is, as Altholz put it, ‘one of those historical events the substance and significance of which are clear, but whose specifics are decidedly fuzzy around the edges.’ It is the purpose of this paper to present a full and balanced view of the specific ingredients, permitting a better insight into the event's symbolism and significance.