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Chapter 5 - Concepts of Metabolism

Published online by Cambridge University Press:  12 May 2020

S. L. Kochhar
Affiliation:
University of Delhi
Sukhbir Kaur Gujral
Affiliation:
University of Delhi
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Summary

Metabolism

Plants can synthesize most of the organic matter on earth by assimilating inorganic elements (carbon and nitrogen), from the environment into organic molecules, driven by energy from sunlight. Every year, plants incorporate through photosynthesis nearly 100 billion metric tons of carbon, which is approximately 15 per cent of the total carbon dioxide (CO2) present in the atmosphere. Respiration by plants and heterotrophic organisms converts the same amount of carbon present in organic compounds back into CO2. These processes are important not only for the plants, but also for animals, fungi, and bacteria, which receive their carbon and nitrogen solely from organic compounds and thus completely rely on plant sources to obtain nutrition. Plants acquire the major elements (C, H, O and N) chiefly as carbon dioxide, water, and nitrate along with other minerals elements in lesser amounts from the soil. This mode of nutrition, as obtained from inorganic compounds, is called autotrophy and photosynthetic bacteria, green algae, and vascular plants are examples of autotrophs. On the other hand, for those organisms that get their carbon and nitrogen exclusively from organic compounds, their mode of nutrition is called heterotrophy. Multicellular animals, carnivorous plants, fungi, and most of the microorganisms are heterotrophs. Carbon, oxygen, and nitrogen are continuously recycled between the heterotrophs and autotrophs present in the biosphere, using the sun's energy as the driving force and this recycling process depends on a proper balance between their activites. These cycles of matter are controlled by huge energy flows within the biosphere, starting with the harvesting of sun's energy by photoautotrophs and its utilization in producing organic compounds including carbohydrates, which are then consumed as sources of energy by heterotrophs. Although carbon, oxygen, and nitrogen keep on recycling constantly, a part of the energy involved is being continuously changed into non-usable form such as heat.

Metabolism can be defined as ‘the sum total of all the chemical reactions occurring in a cell or organism’, and takes place through a sequence of reactions catalyzed by enzymes, thus constituting metabolic pathways and the metabolic intermediates are known as metabolites. Every successive step of the metabolic pathway results in a small and specific chemical change (elimination/inclusion/shifting of a specific molecule or the functional group).

Type
Chapter
Information
Plant Physiology
Theory and Applications
, pp. 153 - 167
Publisher: Cambridge University Press
Print publication year: 2020

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