The plant root is a complex system that has evolved under the constraints of a number of functions. It is a pressure-probe that can penetrate the soil; it is a scavenger of nutrients that may be either tightly bound to soil particles or in low concentrations in the soil solution; it is an absorber of water from the soil. The tip of the root contains the region of dividing cells from which the root is formed, and is pushed through the soil by the extension of these cells some 10–20 times as they develop and differentiate.

The sensitivity of biological systems to changes in environmental stimuli is connected with their regulatory properties. In order to achieve efficient control, these systems must respond to minute environmental variations by amplifying external stimuli to yield a significant response. To that end, biochemical systems have often evolved to a cascade organization in which the product of the nth reaction in a chain acts as a catalyst in subsequent transformations. The amplification properties of such cascades were first noticed in the process of blood clotting (MacFarlane, 1964, 1969) and visual excitation (Wald, 1965). Later on, a similar organization was noticed in hormonal control of metabolism (Bowness, 1964; Stadtman & Chock, 1977, 1978; Chock, Rhee & Stadtman, 1980).