Globular clusters (GCs) and nuclei in dEs are important probes for for studying cosmology and galaxy evolution. GCs and nuclei are usually formed in the high pressures of starbursts, therefore resolved stellar color-magnitude diagrams and integrated colors, magnitudes, or spectra yield ages and metallicities which are snapshots of the conditions during the most violent and important star forming episodes of the galaxies. Most are older, more metal-poor, and more enriched in $\alpha$-elements than the underlying dE field stars. About 25% have formed within the last $\sim$6~Gyr from enriched gas. Many nuclei have the properties of brights GCs but some can be distinguished by large sizes and composite stellar populations. The relatively large numbers of GCs in dEs shows that they are an important source of GCs to galaxies that accrete them.