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Tetralichine trilobites from the Upper Ordovician of Oklahoma and Virginia and phylogenetic systematics of the Tetralichini

Published online by Cambridge University Press:  14 July 2015

Jesse R. Carlucci ,
Stephen R. Westrop  and
Lisa Amati
Jesse R. Carlucci
Affiliation:
1Oklahoma Museum of Natural History and School of Geology and Geophysics, University of Oklahoma, Norman, Oklahoma 73072, USA, ,
Stephen R. Westrop
Affiliation:
1Oklahoma Museum of Natural History and School of Geology and Geophysics, University of Oklahoma, Norman, Oklahoma 73072, USA, ,
Lisa Amati
Affiliation:
2Department of Geology, State University of New York at Potsdam, Potsdam, New York 13676, USA,
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Abstract

New material, including a complete exoskeleton, from the Bromide Formation of Oklahoma, allows a complete evaluation of Probolichas Phleger, 1936 and necessitates a quantitative revision of the lichid tribe Tetralichini Phleger, 1936. A strict consensus tree of nine equally parsimonious trees generated from a branch and bound analysis indicates that the genus Probolichas is monophyletic, and should no longer be restricted as synonymous with, or as a sub-clade of Amphilichas Raymond, 1905. The genus Apatolichas Whittington, 1963 is revised as a synonym of Amphilichas, and the Tetralichini is defined as clade with three genera: Amphilichas, Probolichas, and Lyralichas Weber, 1948. Probolichas kristiae from the Bromide Formation and Amphilichas effnensis from the Effna Formation of Virginia are new species.

Type
Research Article
Information
Journal of Paleontology , Volume 84 , Issue 6 , November 2010 , pp. 1099 - 1120
Copyright
Copyright © The Paleontological Society 

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References

Amati, L. and Westrop, S. 2004. A systematic revision of Thaleops (Trilobita: Illaenidae) with new species from the Middle and Late Ordovician of Oklahoma and New York. Journal of Systematic Palaeontology, 2(3):207–256.CrossRefGoogle Scholar
Amati, L. and Westrop, S. 2006. Sedimentary facies and trilobite biofacies along an Ordovician shelf to basin gradient, Viola Group, South-Central Oklahoma. PALAIOS, 21:516–529.CrossRefGoogle Scholar
Amsden, T. W. and Sweet, W. C. 1983 Upper Bromide Formation and Viola Group (Middle and Upper Ordovician) in Eastern Oklahoma. Oklahoma Geological Survey Bulletin, 132:1–76.Google Scholar
Angelin, N. P. 1854. Palaeontologica Scandinavica Pars II. Academiae Regiae Scientarium Suecanae (Holmiae), 21–92.Google Scholar
Barton, D. C. 1916. A revision of the Cheirurinae, with notes on their evolution. Washington University Studies in Science, 3:101–152.Google Scholar
Beyrich, E. 1845. Ueber einige böhmische Trilobiten, Berlin, 47 p.Google Scholar
Billings, E. 1865. Palaeozoic fossils, containing descriptions and figures of new or little known species of organic remains from the Silurian rocks. Dawson Brothers, Montreal, 426 p.Google Scholar
Bradley, J. H. 1930. Fauna of the Kimmswick Limestone of Missouri and Illinois. Walker Museum, University of Chicago Contributions, 2(219-290).Google Scholar
Bremer, K. 1994. Branch support and tree stability. Cladistics, 10(3):295–304.CrossRefGoogle Scholar
Chatterton, B. D. E. and Ludvigsen, R. 1976. Silicified Middle Ordovician trilobites from the South Nahanni river area, District of Mackenzie, Canada. Palaeontographica, 154:1–106.Google Scholar
Clarke, J. M. 1894. The Lower Silurian trilobites of Minnesota. Minnesota Geology and Natural History Survey, p. 694–759.Google Scholar
Conrad, T. A. 1842. Observations on the Silurian and Devonian systems of the United States, with descriptions of new organic remains. Journal of the Academy of National Sciences Philadelphia, 8:228–235.Google Scholar
Conrad, T. A. 1843. Observations on the lead bearing limestone of Wisconsin, and description of a new genus of trilobites and fifteen new Silurian fossils. Proceedings of the Academy of Natural Sciences of Philadelphia, 1:329–335.Google Scholar
Cooper, B. N. 1944. Geology and mineral resources of the Rukes Garden quadrangle, Virginia. Virginia Geological Survey Bulletin 60.Google Scholar
Cooper, B. N. and Cooper, G. A. 1946. Lower Middle Ordovician stratigraphy of the Shenandoah Valley, Virginia. Bulletin of the Geological Society of America, 57(35-114).CrossRefGoogle Scholar
Cooper, G. A. 1930. Upper Ordovician and Lower Devonian stratigraphy and palaeontology of Percé, Quebec: part 2; new species from the Upper Ordovician of Percé. American Journal of Science, 5(20):385–392.Google Scholar
Cooper, G. A. 1956. Chazyan and related brachiopods. Smithsonian Miscellaneous Collections, 127(Pt. 1, 2):1–245.Google Scholar
Dalman, J. W. 1827. Om Palaeaderna, eller de sa kallade Trilobiterna. Kongliga Svenska vetenskapsadademiens handlingar, 113-152, 226–294.Google Scholar
De Queiroz, K. and Gauthier, J. 1990. Phylogeny as a central principle in taxonomy: phylogenetic definitions of taxon names. Systematic Zoology, 39(4):307–322.CrossRefGoogle Scholar
De Queiroz, K. 2007. Toward an integrated system of clade names. Systematic Biology, 56(6):956–974.CrossRefGoogle ScholarPubMed
Eckert, T. 1951. The fauna of the upper Bromide Formation. Unpublished , University of Oklahoma, Norman, 110 p.Google Scholar
Edgecombe, G. D., Banks, M. R., and Banks, D. M. 2004. Late Ordovician trilobites from Tasmania: Styginidae, Asaphidae and Lichidae. AAPG Memoir, 30:59–77.Google Scholar
Emerson, N., Simo, J. A., Byers, C., and Fournelle, J. 2004. Correlation of (Ordovican, Mohawkian) K-bentonites in the upper Mississippi valley using apatite chemistry: Implications for stratigraphic interpretation of the mixed carbonate-siliciclastic Decorah Formation. Palaeogeography, Palaeoclimatology, Palaeoecology, 210:215–233.CrossRefGoogle Scholar
Esker, G. C. 1964. New species of trilobites from the Bromide Formation (Poolevillve Member) of Oklahoma. Oklahoma Geology Notes, 24:195–209.Google Scholar
Evitt, W. 1951. Some Middle Ordovician trilobites of the families Cheiruridae, Harpidae and Lichidae. Journal of Paleontology, 25(5):587–616.Google Scholar
Farris, J. S. 1969. A Successive Approximations Approach to Character Weighting. Systematic Zoology, 18(4):374–385.CrossRefGoogle Scholar
Fay, R. O. and Graffham, A. A. 1982. Biostratigraphic and paleontological studies, p. 195–209. In Sprinkle, J. (ed.), Echinoderm Faunas from the Bromide formation (Middle Ordovician) of Oklahoma. Vol. 1. University of Kansas Paleontological Contributions, Monograph 1.Google Scholar
Finney, S. C. 1986. Graptolite biofacies and correlation of eustatic, subsidence, and tectonic events in the Middle to Upper Ordovician of North America. PALAIOS, 1:435–461.CrossRefGoogle Scholar
Foerste, A. F. 1919. Notes on Isotelus, Acrolichas, Calymene and Encrinurus. Bulletin of the scientific labs of Denison University, 19:65–81.Google Scholar
Foerste, A. F. 1920. The generic relations of the American Ordovician Lichadidae. American Journal of Science, 4(49):26–50.CrossRefGoogle Scholar
Fortey, R. and Droser, M. 1999. Trilobites from the base of the type Whiterockian (Middle Ordovician) in Nevada. Journal of Paleontology, 73(2):182–201.CrossRefGoogle Scholar
Goloboff, P., Farris, J., and Nixon, K. 2008. TNT, a free program for phylogenetic analysis. Cladistics, 24:774–786.CrossRefGoogle Scholar
Green, J. 1832. A monograph of the trilobites of North America. Joseph Brano, Philadelphia, 93 p.Google Scholar
Gürich, G. 1901. Ueber eine neue Lichas-art aus dem Devon von Neu-Süd Wales und über die Gattung Lichas überhaupt. Neues Jahr-buch für Mineralogie, Geologie und Paläontologie, 14:519–539.Google Scholar
Harris, R. W. 1957. Ostracoda of the Simpson Group. Oklahoma Geological Survey Bulletin, 75(1-333).Google Scholar
Hawle, I. and Corda, A. J. C. 1847. Prodrom einer monographie der böhmischen Trilobiten. Abhandlungen Koeniglichen Böehmischen Gesellschaft der Wissenschaften. J.G Calve, Prague.Google Scholar
Holland, S. and Patzkowsky, M. 1996. Sequence stratigraphy and long-term paleooceanographic changes in the Middle and Upper Ordovician of the eastern United States. In Witzke, B., Ludvigson, G., and Day, J. (eds.), Paleozoic Sequence Stratigraphy: Views from the North American Craton, GSA Special Paper 306. Geological Society of America, Boulder CO.Google Scholar
Karim, T. and Westrop, S. 2002. Taphonomy and paleoecology of Ordovician trilobite clusters, Bromide Formation, South-central Oklahoma. PALAIOS, 17:394–403.2.0.CO;2>CrossRefGoogle Scholar
King, D. T. 1986. Waulsortian-type buildups and resedimented (carbonate-turbidite) facies, early Mississippian Burlington shelf, central Missouri. Journal of Sedimentary Petrology, 56:471–479.Google Scholar
Lazarenko, N. P. 1960. Some new Upper Cambrian trilobites of the northwest part of the Siberian Platform. Trudy Nauchno-Issledovatel'skogo Instituta Geologii Arktiki, 20:38–41.Google Scholar
Longman, M. W. 1982a. Depositional environments, p. 17–29. In Sprinkle, J. (ed.), Echinoderm faunas from the Bromide Formation (Middle Ordovician) of Oklahoma. Vol. 1. University of Kansas Paleontological Contributions, Monograph 1.Google Scholar
Longman, M. W. 1982b. Depositional setting and regional characteristics. In Sprinkle, J. (ed.), Echinoderm faunas from the Bromide Formation (Middle Ordovician) of Oklahoma. The University of Kansas Paleontological Contributions, Monograph 1S.Google Scholar
Ludvigsen, R. 1978. The trilobites Bathyurus and Eomonorachus from the Middle Ordovician of Oklahoma and their biofacies significance. Royal Ontario Museum Life Sciences Contributions, 114:1–18.Google Scholar
Maddison, D. and Maddison, W. 2005. MacClade: Analysis of Phylogeny and Character Evolution, 4.08. Sinauer Associates Inc, Sunderland, Maryland.Google Scholar
Maletz, J., Carlucci, J., and Mitchell, C. 2009. Graptoloid cladistics, taxonomy and phylogeny. Bulletin of Geosciences, 84(1):7–19.CrossRefGoogle Scholar
M'Coy, F. 1849. On the classification of some British fossil Crustacea, with notices of new forms in the University collection at Cambridge. Annual Magazine of Natural History (London), 2(4):161–179.CrossRefGoogle Scholar
Meek, F. B. and Worthen, A. H. 1865. Contributions to the palaeontology of Illinois and other western states. Proceedings of the Academy of Natural Sciences of Philadelphia, 17(5):245–273.Google Scholar
Moore, R. C. 1959. Treatise on invertebrate paleontology, Part O, Arthropoda 1. Geological Society of America and University of Kansas, Boulder, CO and Lawrence, KA, 560 p.Google Scholar
Nixon, K. 2002. WinClada ver. 1.00.08.Google Scholar
Öpik, A. A. 1937. Trilobiten aus Estland. Publications of the Geologic Institutions of the University of Tartu, 52:1–163.Google Scholar
Owen, A. W. 1981. The Ashgill trilobites of the Oslo Region, Norway. Palaeontographica, A, 175:1–88.Google Scholar
Phleger, F. 1936. Lichadian Trilobites. Journal of Paleontology, 10(7):593–615.Google Scholar
Pollitt, J., Fortey, R., and Wills, M. 2005. Systematics of the trilobite families Lichidae Hawle & Corda, 1847 and Lichakephalidae Tripp, 1957: The application of bayesian inference to morphological data. Journal of Systematic Palaeontology, 3(3):225–241.CrossRefGoogle Scholar
Portlock, J. E. 1843. Report on the geology of the Country of Londonderry and of parts of Tyrone and Fermanagh. Longman, Brown, Green, and Longmans, London, 475 p.Google Scholar
Raymond, P. E. 1905. Notes on the names Amphion, Harpina and Platymetopus. American Journal of Science, 4(19):377–378.CrossRefGoogle Scholar
Raymond, P. E. 1910. Notes on Ordovician trilobites IV, new and old species from the Chazy. Annals of the Carnegie Museum, 7:60–80.CrossRefGoogle Scholar
Raymond, P. E. 1925. Some trilobites of the lower Middle Ordovician of eastern North America. Bulletin of the Museum of Comparative Zoology, 67:1–180.Google Scholar
Read, J. F. 1980. Carbonate ramp-to-basin Transitions and foreland basin evolution, Middle Ordovician, Virginia Appalachians. AAPG Bulletin, 64(10):1575–1612.Google Scholar
Read, J. F. 1982. Geometry, facies, and development of Middle Ordovician carbonate buildups, Virginia Appalachians. AAPG Bulletin, 66(2):189–209.Google Scholar
Ross, J. 1972. Fossils from the Ordovician bioherm at Meiklejohn Peak, Nevada. U.S. Government Printing Office, Washington, 685, 1–47 p.Google Scholar
Ruppel, S. and Walker, K. 1977. The ecostratigraphy of the Middle Ordovician of the Southern Appalachians (Kentucky, Tennessee, and Virginia), U.S.A.: A Field Excursion. Studies in Geology, University of Tennessee Department of Geological Sciences, 77–1.Google Scholar
Sabol, J. W. 1958. Geology of the Porterfield Quarry area, Smyth County, Virginia, Unpublished , Virginia Polytechnic Institute, 81 p.Google Scholar
Sdzuy, K. 1955. Die Fauna der Leimintz-Schiefer (Tremadoc). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 492:1–74.Google Scholar
Shaw, F. C. 1968. Early Middle Ordovician Chazy trilobites of New York. New York State Museum and Science Services Memoir 17, 163 p.Google Scholar
Shaw, F. 1974. Simpson Group (Middle Ordovician) Trilobites of Oklahoma. Journal of Paleontology Memoir, 48(6):1–54.CrossRefGoogle Scholar
Shimer, H. W. and Shrock, R. R. 1944. Index Fossils of North America. M.I.T. Press, 837 p.Google Scholar
Slocom, A. W. 1913. New trilobites from the Maquoketa beds of Fayette County, Iowa. Field Museum of Natural History Geological Series, 4:43–83.Google Scholar
Sprinkle, J. 1982. Introduction. In Sprinkle, J. (ed.), Echinoderm faunas from the Bromide Formation (Middle Ordovician) of Oklahoma. University of Kansas Paleontological Contributions, Monograph 1.Google Scholar
Strait, D., Moniz, M., and Strait, P. 1996. Finite Mixture Coding: A new approach to coding continuous characters. Systematic Biology, 45(1):67–78.CrossRefGoogle Scholar
Strong, E.E. and Lipscomb, D. 1999. Character coding and inapplicable characters. Cladistics, 15:363–371.CrossRefGoogle Scholar
Swofford, D. L. 2002. PAUP* Phylogentic Analysis Using Parsimony (*and other methods), Version 4.0b10. Sinauer Associates, Sunderland, Maryland.Google Scholar
Thiele, K. 1993. The holy grail of the perfect character: the cladistic treatment of morphometric data. Cladistics, 9:275–304.CrossRefGoogle ScholarPubMed
Thomas, A. T. and Holloway, D. J. 1988. Classification and phylogeny of the trilobite order Lichida. Philosophical Transactions of the Royal Society of London, 321(1205):179–262.Google Scholar
Törnquist, S. L. 1884. Undersökningar öfver Siljansområdets trilobit-fauna. Sveriges Geologiska Undersokning Afh, 436:1–191.Google Scholar
Tripp, R. P. 1957. The classification and evolution of the superfamily Lichacea (Trilobita). Geological Magazine, 94:104–122.CrossRefGoogle Scholar
Tripp, R. P. 1959. Family Lichidae, p. 579. In Moore, R. C. (ed.), Treatise on Invertebrate Paleontology, Part O, Arthropoda I. Geological Society of America.Google Scholar
Ulrich, E. O. 1892. Two new Lower Silurian species of Lichas (Subgenus Hoplolichas). American Geologist, 10:271–272.Google Scholar
Ulrich, E. O. 1911. Revision of the Paleozoic systems. Geological Society of America Bulletin, 22:281–680.CrossRefGoogle Scholar
Walter, O. T. 1924. Trilobites of Iowa and some related Paleozoic forms. Iowa Geological Survey Annual Reports 1923 and 1924, 31:1–388.Google Scholar
Warburg, E. 1925. The trilobites of the Leptaena Limestone in Dalarne. Bulletin of the Geologic Institute of the University of Upsala, 27:1–446.Google Scholar
Warburg, E. 1939. The Swedish Ordovician and Lower Silurian Lichidae. Kungliga Svenska Vetenskapsakademiens Handlingar, Stockholm, 17(4):1–162.Google Scholar
Webby, B. D. 1974. Upper Ordovician Trilobites from Central New South Wales. Palaeontology, 17(2):203–252.Google Scholar
Weber, V. N. 1932. Trilobites of the Turkestan. Monographs of the Palaeontology of the USSR, 178:3–151.Google Scholar
Weber, V. N. 1948. Trilobites of the Silurian deposits of the USSR. Part 1. The Lower Silurian trilobites. Monografii po Paleontologii SSR, 69:1–111.Google Scholar
Westrop, S. R. and Ludvigsen, R. 1983. Systematics and Paleoecology of Upper Ordovician Trilobites from the Selkirk Member of the Red River Formation, Southern Manitoba. Manitoba Department of Energy and Mines Geological Report GR 82-2, Winnipeg, 51 p.Google Scholar
Whiteaves, J. F. 1897. The fossils of the Galena-Trenton and Black River Formations of Lake Winnipeg and its vicinity. Geological Survey of Canada, 3:192–242.Google Scholar
Whittington, H. B. 1949. Dolichoharpes and the origin of the harpid fringe. American Journal of Science, 247:276–285.CrossRefGoogle Scholar
Whittington, H. B. 1954. Ordovician trilobites from Silliman's Fossil Mount. Geological Society of America Memoir, 62:119–149.Google Scholar
Whittington, H. B. 1963. Middle Ordovician trilobites from Lower Head, western Newfoundland. Bulletin of the Museum of Comparative Zoology, 129:1–118.Google Scholar
Whittington, H. B. and Kelly, S. R. A. 1997. Morphological terms applied to Trilobita. In Kaesler, R. L. (ed.), Treatise on Invertebrate Paleontology, Pt. O, Arthropoda 1. Trilobita (revised). Vol. 1. Introduction, Order Agnostida, Order Redlichiida. Geological Society of America and University of Kansas, Lawrence.Google Scholar
Wilson, J. L. 1975. Carbonate facies in geologic history. Springer-Verlag, New York, 471 p.CrossRefGoogle Scholar