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The taxonomic status of Merychippus insignis Leidy
Published online by Cambridge University Press: 14 July 2015
Extract
The genus Merychippus was erected by Leidy (1857), who named as the type species Merychippus insignis. The type specimen of M. insignis is an immature and incomplete maxilla containing a broken dp2 and a dp3. The type comes from the Bijou Hills of South Dakota, probably from the Barstovian (middle Miocene) Fort Randall Formation of South Bijou Hill in Charles Mix County (Skinner and Taylor, 1967), but possibly from the overlying undifferentiated Ogallala Formation, or possibly from North Bijou Hill in Brule County. Despite this enigmatic type, the species M. insignis has frequently been identified in the fossil record, and the genus Merychippus has grown to include all mesodont horses (Stirton, 1940). Conceptually, the taxon Merychippus is considered a horizontal grade rather than a vertical clade (Simpson, 1945, p. 18). As a horizontal concept, the genus Merychippus derived importance as the ancestral group for the many Mio-Pliocene lineages of hypsodont horses. Today, as horse classification is remodeled along cladistic lines, the taxon Merychippus derives importance from its early naming. Several distinctive horse clades extend upward from a middle Miocene radiation (Stirton, 1940; Quinn, 1955). If M. insignis can be placed within one of these clades, then it is likely that the clade will bear the generic nomen Merychippus because of the antiquity of the name.
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