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The strophomenide brachiopod Ahtiella Öpik in the Ordovician of Gondwana and the early history of the plectambonitoids

Published online by Cambridge University Press:  26 June 2018

Juan L. Benedetto
Juan L. Benedetto*
Affiliation:
Centro de Investigaciones en Ciencias de la Tierra (CICTERRA), Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET) and Universidad Nacional de Córdoba, Vélez Sarsfield 1611, X5016GCA Córdoba, Argentina 〈[email protected]〉
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Abstract

The Precordilleran species Ahtiella argentina Benedetto and Herrera, 1986 is redescribed and illustrated and Monorthis coloradoensis Benedetto, 1998b from northwestern Argentina is reassigned to the genus Ahtiella Öpik, 1932. Ahtiella famatiniana new species from volcaniclastic rocks of the Famatina range (western Argentina) and Ahtiella tunaensis new species from the Precordillera basin (Cuyania terrane) are proposed. Paleogeographic and stratigraphic evidence strongly suggests that Ahtiella originated in the Andean region of Gondwana to further migrate to Avalonia, Baltica, and Cuyania. Contrary to previous assumptions, the fossil record from the Famatina volcaniclastic succession suggests that the plectambonitoid Ahtiella famatiniana n. sp. evolved from the hesperonomiid orthoid Monorthis transversa Benedetto, 2003 that always occurs in the underlying strata. Phylogenetic analysis of Ahtiella species shows that A. famatiniana n. sp. and the Peruvian A. zarelae Villas in Gutiérrez-Marco and Villas, 2007 are not only the earliest species of the genus but also are morphologically intermediate between Monorthis Bates, 1968 and the later and more derived species of Ahtiella from Baltica and Cuyania. If, as empirical evidence presented here shows, Ahtiella originated from Monorthis through a series of minor transformations, then the impressive morphological gap between orthides and strophomenides was bridged through short-time cladogenesis events, suggesting that it might not have a definite discontinuity between the species level evolution and the origin of higher taxa (macroevolution).

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Articles
Information
Journal of Paleontology , Volume 92 , Issue 5 , September 2018 , pp. 768 - 793
Copyright
Copyright © 2018, The Paleontological Society 

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References

Aceñolaza, F.G., Miller, H., and Toselli, A.J., 2002, Proterozoic−early Paleozoic evolution in western South America—A discussion: Tectonophysics, v. 354, p. 121–137. doi: 10.1016/s0040-1951(02)00295-0.Google Scholar
Albanesi, G.L., and Astini, R.A., 2000, New conodont fauna from Suri Formation (Early−Middle Ordovician), Famatina System, western Argentina: Reunión Anual de la Asociación Paleontológica Argentina, Mar del Plata, Abstracts, Ameghiniana, v. 37, no. 4, supplement, p. 68.Google Scholar
Albanesi, G.L., and Astini, R.A., 2002, Faunas de conodontes y Sacabambaspis janvieri (Vertebrata) en el Ordovícico Medio de la Cordillera Oriental Argentina: Implicancias estratigráficas y paleobiogeográficas: Congreso Argentino de Paleontología y Bioestratigrafía, 8th, Corrientes, Argentina, Abstracts, p. 17.Google Scholar
Albanesi, G.L., and Ortega, G., 2002, Advances on conodont-graptolite biostratigraphy of the Ordovician System of Argentina, in Aceñolaza, F.G., ed., Aspects of the Ordovician System in Argentina: INSUGEO Serie Correlación Geológica, v. 16, p. 143–166.Google Scholar
Albanesi, G.L., and Vaccari, N.E., 1994, Conodontos del Arenig en la Formación Suri, Sistema de Famatina: Argentina, Revista Española de Micropaleontología, v. 26, p. 125–146.Google Scholar
Astini, R.A., 1994, Interpretación estratigráfica de la Formación Sepulturas (Ordovícico Inferior) y unidades análogas del Noroeste Argentino: La Aloformación Sepulturas: 5th Reunión Argentina de Sedimentología, San Miguel de Tucumán, Argentina, Abstracts, p. 9–14.Google Scholar
Astini, R.A., 1998, Stratigraphical evidence supporting the rifting, drifting and collision of the Laurentian Precordillera terrane of western Argentina, in Pankhurst, R.J., and Rapela, C.W., eds., The Proto-Andean Margin of Gondwana: Geological Society London Special Publication, v. 142, p. 11–33.Google Scholar
Astini, R.A., 1999, Sedimentary record, volcano-tectonic cyclicity and progressive emergence of an Early Ordovician perigondwanan volcanic arc: The Famatina System, in Kraft, P., and Fatka, O., eds., Quo vadis Ordovician?: Acta Universitatis Carolinae Geologica, v. 43, nos. 1/2 p. 115–118.Google Scholar
Astini, R.A., 2003, The Ordovician proto-Andean basins, in Benedetto, J.L., ed., Ordovician Fossils of Argentina: Córdoba, Argentina, Secretaría de Ciencia y Tecnología, Universidad Nacional de Córdoba, p. 1–74.Google Scholar
Astini, R.A., and Benedetto, J.L., 1996, Paleoenvironmental features and basin evolution of a complex island-arc region in the pre-Andean western Gondwana: The Famatina belt: 3rd International Symposium on Andean Geodynamics, St. Malo, France, Short Papers, p. 755–758. St. Malo.Google Scholar
Astini, R.A., and Dávila, D.F., 2002, El Grupo Cerro Morado (Ordovícico Medio) en el Famatina (28°−29°S), Andes centrales del oeste argentino: Revista Geológica de Chile, v. 29, p. 241–254. doi: 10.4067/S0716-02082002000200006.Google Scholar
Astini, R.A., and Rapalini, A.E., 2003, Discussion. ‘Proterozoic-Early Paleozoic evolution in western South America - a discussion’ in: Tectonophysics, 354: 121–137 (2002): Tectonophysics, v. 366, p. 143–148.Google Scholar
Astini, R.A., Benedetto, J.L., and Vaccari, N.E., 1995, The early Paleozoic evolution of the Argentine Precordillera as a Laurentian rifted, drifted and collided terrane: Geological Society of America Bulletin, v. 107, p. 253–273.Google Scholar
Astini, R.A., Toro, B.A., Waisfeld, B.G., and Benedetto, J.L., 2004a, Revisión estratigráfica de la región de Los Colorados: Un perfil condensado del Paleozoico inferior en el borde occidental del Cordillera Oriental, Provincia de Jujuy: Revista de la Asociación Geológica Argentina, v. 59, p. 243–260.Google Scholar
Astini, R.A., Thomas, W.A., and Yochelson, E.L., 2004b, Salterella in the Argentine Precordillera: An Early Cambrian paleobiogeographic indicator of Laurentian affinity: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 213, p. 125–132. doi: 10.1016/j.palaeo.2004.07.008.Google Scholar
Baldo, E.G., Fanning, C.M., Rapela, C.W., Pankhurst, R.J., Casquet, C., and Galindo, C., 2003, U-Pd SHRIMP dating of rhyolite volcanism in the Famatina belt and K-bentonites in the Precordillera, in Albanesi, G.L., Beresi, M.S., and Peralta, S H., eds., Ordovician from the Andes: Proceedings of the 9th International Symposium on the Ordovician System, INSUGEO Serie Corelación Geológica, v. 17, p. 285–189.Google Scholar
Bancroft, B.B., 1949, Welsh Valentian brachiopods, in Lamont, A., ed., Welsh Valentian Brachiopods and the Strophomena antiquata Group of Fossil Brachiopods: Preprint from The Quarry Managers’ Journal, Mexborough, Yorkshire, UK, Times Printing Company, p. 2–10, pls. 1–2.Google Scholar
Bassett, M.G., 1984, Life strategies in Silurian brachiopods, in Bassett, M.G., and Lawson, J.D., eds., Autoecology of Silurian Ornanisms: The Paleontological Society, Special Paper in Palaeontology, v. 32, p. 237–264.Google Scholar
Bassett, M.G., and Popov, L.E., 2017, Earliest ontogeny of the Silurian orthotetide brachiopod Coolinia and its significance for interpreting strophomenate phylogeny: Lethaia, v. 50, p. 504–510. doi: 10.1111/let.12204.Google Scholar
Bassett, M.G., Popov, L.E., and Holmer, L.E., 2001, Functional morphology of articulatory structures and implications for patterns of musculature in Cambrian rhynchonelliform brachiopods, in Brunton, C.H.C., Cocks, L.R.M., and Long, S.L., eds., Brachiopods Past and Present: London, Taylor & Francis, p. 163–176.Google Scholar
Bassett, M.G., Popov, L.E., and Holmer, L.E., 2002, Brachiopods: Cambrian-Tremadocian precursors to Ordovician radiation events, in Crame, J.A., and Owen, A.W., eds., Palaeobiogeography and Biodiversity Change: The Ordovician and Mesozoic-Cenozoic Radiations: Geological Society London Special Publication, v. 194, p. 13–23.Google Scholar
Bassett, M.G., Popov, L.E., and Egerquist, E., 2008, Early ontogeny of some Ordovician-Silurian strophomenate brachiopods: Significance for interpreting evolutionary relationships within early Rhynchonelliformea: Fossils and Strata, v. 54, p. 13–20.Google Scholar
Bates, D.E.B., 1968, The lower Paleozoic brachiopod and trilobites faunas of Anglesey: Bulletin of the British Museum (Natural History) Geology, v. 16, p. 125–199.Google Scholar
Bates, D.E.B., 1969, Some early Arenig brachiopods and trilobites from Wales: Bulletin of the British Museum (Natural History) Geology, v. 18, p. 1–28.Google Scholar
Benedetto, J.L., 1994, Braquiópodos ordovícicos (Arenigíano) de la Formación Suri en la región de Chaschuil, Sistema de Famatina, Argentina: Ameghiniana, v. 31, p. 221–238.Google Scholar
Benedetto, J.L., 1998a, Early Palaeozoic brachiopods and associated shelly faunas from western Gondwana: Its bearing on the geodynamic history of the pre-Andean margin, in Pankhurst, R.J., and Rapela, C.W., eds., The Proto-Andean Margin of Gondwana: Geological Society London Special Publication, v. 142, p. 57–83.Google Scholar
Benedetto, J.L., 1998b, Early Ordovician (Arenig) brachiopods from the Acoite and Sepulturas formations: Cordillera Oriental, northwestern Argentina: Geologica et Palaeontologica, v. 32, p. 7–27.Google Scholar
Benedetto, J.L., 2001, Silicified Early Ordovician (Arenig) brachiopods from the San Juan Limestone, Argentine Precordillera: Geologica et Palaeontologica, v. 35, p. 1–29.Google Scholar
Benedetto, J.L., 2002, The Ordovician brachiopod faunas of Argentina: Chronology and biostratigraphic succession, in Aceñolaza, F.G, ed., Aspects of the Ordovician System in Argentina: 9th International Symposium on the Ordovician System, San Miguel de Tucumán, Argentina, Proceedings, INSUGEO, Serie Correlación Geológica, v. 16, p. 87–106.Google Scholar
Benedetto, J.L., 2003a, Brachiopods, in Benedetto, J.L., ed., Ordovician Fossils of Argentina: Córdoba, Argentina. Secretaría de Ciencia y Tecnología, Universidad Nacional de Córdoba, p. 187–272.Google Scholar
Benedetto, J.L., 2003b, Early Ordovician (Arenig) brachiopods from volcaniclastic rocks of the Famatina Range, northwest Argentina: Journal of Paleontology, v. 77, p. 212–242. doi: 10.1666/0022-3360(2003)077 0212:EOABFV 2.0.CO;2.Google Scholar
Benedetto, J.L., 2004, The allochthony of the Precordillera ten years later (1993–2003): A new paleobiogeographic test of the microcontinental model: Gondwana Research, v. 7, p. 1027–1039. doi: 10.1016/S1342-937X(05)71082-0.Google Scholar
Benedetto, J.L., 2007, Brachiopod succession in the Lower-Middle Ordovician carbonate platform of the Precordillera terrane, western Argentina: An example of interplay between environmental, biogeographic, and evolutionary processes: Acta Palaeontologica Sinica, v. 46, p. 28–36.Google Scholar
Benedetto, J.L., 2008, Spatial and stratigraphic distribution of the rhynchonelliformean brachiopod Productorthis Kozlowski: Fast migrations or parallel evolution?, in Harper, D.A.T., Long, S.L., and Nielsen, C., eds., Brachiopoda: Fossil and Recent: Fossils and Strata, v. 54, p. 21–29.Google Scholar
Benedetto, J.L., 2009, Chaniella, a new lower Tremadocian (Ordovician) brachiopod from northwestern Argentina and its phylogenetic relationships with basal rhynchonelliforms: Palaeontologische Zeitschrift, v. 83, p. 393–405.Google Scholar
Benedetto, J.L., 2013, Presence of punctae in the ‘plectorthoidean’ brachiopod Famatinorthis turneri (Middle Ordovician) from western Argentina: Implications for early diversification of punctate orthides: Lethaia, v. 42, p. 170–179. doi: 10.1111/j.1502-3931.2012.00330.x.Google Scholar
Benedetto, J.L., and Carrasco, P., 2002, Tremadoc (earliest Ordovician) brachiopods from the Purmamarca region and the Sierra de Mojotoro, Cordillera Oriental of northwestern Argentina: Geobios, v. 35, p. 647–661. doi: 10.1016/S0016-6995(02)00079-7.Google Scholar
Benedetto, J.L., and Foglia, R.D., 2012, Lower and Middle Cambrian rhynchonelliformean brachiopods from the Precordillera terrane of Argentina: Journal of Paleontology, v. 86, p. 273–281. doi: 10.2307/41480192.Google Scholar
Benedetto, J.L., and Herrera, Z.A., 1986, Braquiópodos del Suborden Strophomenidina de la Formación San Juan (Ordovícico temprano), Precordillera de San Juan, Argentina: Actas 4th Congreso Argentino Paleontología y Bioestratigrafía, Mendoza, Argentina, v. 1, p. 113–123.Google Scholar
Benedetto, J.L., and Herrera, Z.A., 1987, Sanjuanella, un nuevo género de la subfamilia Ahtiellinae (Brachiopoda, Plectambonitacea) del Ordovícico de la Precordillera Argentina: Actas 4th Congreso Latinoamericano de Paleontología, Santa Cruz de la Sierra, Bolivia, v. 1, p. 97–109.Google Scholar
Benedetto, J.L., and Muñoz, D.F., 2017, Plectorthoid brachiopods from the Lower Ordovician of NW Argentina: Phylogenetic relationships with Tarfaya Havlíček and the origin of heterorthids: Journal of Systematic Palaeontology, v. 15, p. 43–67. doi: 10.1080/14772019.2016.1144086.Google Scholar
Benedetto, J.L., and Sánchez, T.M., 2003, The Ordovician radiation from a Gondwanan perspective: The early diversification of brachiopods and bivalves on mid- to high- latitude siliciclastic platforms, in Albanesi, G.L., Beresi, M.S., and Peralta, S.H., eds., Ordovician from the Andes, 9th International Symposium on the Ordovician System, San Miguel de Tucumán, Argentina, Proceedings, INSUGEO Serie Corelacion Geologica, v. 17, p. 245–250.Google Scholar
Benedetto, J.L., Dávila, D.F., and Astini, R.A., 2003, La presencia de Ahtiella Öpik y otros braquiópodos en la sucesión volcaniclástica ordovícica de la Sierra de Famatina: Connotaciones bioestratigráficas y paleogeográficas: Ameghiniana, v. 40, p. 80.Google Scholar
Benedetto, J.L., Sorrentino, L., Cech, N., and Sánchez, T.M., 2008, Functional morphology of the Ordovician plectambonitoid brachiopod Inversella (Reinversella) arancibiai Herrera & Benedetto from the San Juan Formation, Argentine Precordillera: Alcheringa, v. 32, p. 53–63. doi: 10.1080/03115510701757373.Google Scholar
Benedetto, J.L., Vaccari, N.E., Waisfeld, B.G., Sánchez, T.M., and Foglia, R.D., 2009, Cambrian and Ordovician paleobiogeography of the South American margin of Gondwana and accreted terranes, in Bassett, M.G., ed., Early Palaeozoic Peri-Gondwanan Terranes: New Insights from Tectonics and Biogeography: The Geological Society London Special Publication, v. 325, p. 199–230.Google Scholar
Benton, M.J., and Pearson, P.N., 2001, Speciation in the fossil record: Trends in Ecology and Evolution, v. 16, p. 405–411. doi: 10.1016/S0169-5347(01)02149-8.Google Scholar
Bergström, S.M., Chen, X., Gutiérrez-Marco, J.C., and Dronov, A., 2009, The new chronostratigraphic classification of the Ordovician System and its relations to major regional series and stages and to δ13C chemostratigraphy: Lethaia, v. 42, p. 97–107. doi: 10.1111/j.1502-3931.2008.00136.x.Google Scholar
Billings, E., 1859, On some new genera and species of Brachiopoda from the Silurian and Devonian rocks of Canada: Canadian Naturalist and Geologist, v. 4, p. 1–131.Google Scholar
Bruton, D.L., and Harper, D.A.T., 1981, Brachiopods and trilobites of the Early Ordovician serpentine Otta Conglomerate, south central Norway: Norks Geologisk Tidsskrift, v. 61, p. 153–181.Google Scholar
Butts, C., 1926, The Paleozoic rocks, in Adams, G.I., Butts, C., Stephenson, L.W., and Cooke, C.W., eds., Geology of Alabama: Alabama Geological Survey Special Report 14, p. 41–230.Google Scholar
Cañas, F.L., 1999, Facies sequences of late Cambrian to Early Ordovician carbonates of the Argentine Precordillera: A physical stratigraphic comparison with Laurentian platforms, in Keppie, D., and Ramos, V.A., eds., Laurentia-Gondwana Connections Before Pangea: Geological Society of America Special Paper 336, p. 43–62.Google Scholar
Carlorosi, J., Heredia, S., and Aceñolaza, G., 2017, Conodontes de la Formación Suri en la región de Chaschuil, Sistema de Famatina, Argentina: Congreso Geológico Argentino, 20th, San Miguel de Tucumán, Argentina, Actas, p. 21–23.Google Scholar
Carlson, S.J., 2016, The evolution of Brachiopoda: Annual Review of Earth and Planetary Sciences, v. 44, p. 409–438. doi: 10.1146/annurev-earth-060115-012348.Google Scholar
Carrera, M.G., 1997, Significado paleoambiental de los poríferos y briozoos de la Formación San Juan (Ordovícico), Precordillera argentina: Ameghiniana, v. 34, p. 179–199.Google Scholar
Carrera, M.G., 2003, Sponges and Bryozoans, in Benedetto, J.L., ed., Ordovician Fossils of Argentina: Córdoba, Argentina, Secretaría de Ciencia y Tecnología, Universidad Nacional de Córdoba, p. 155–185.Google Scholar
Carrera, M.G., and Astini, R.A., 1998, Valoración de las restricciones ambientales durante la transición Arenigiano-Llanvirniano: Ordovícico de la Precordillera Argentina: Revista de la Asociación Geológica Argentina, v. 53, p. 41–56.Google Scholar
Carrera, M.G., and Ernst, A., 2010, Darriwilian bryozoans from the San Juan Formation (Ordovician), Argentine Precordillera: Ameghiniana, v. 47, p. 343–354. doi: 10.5710/AMGH.v47i3.3.Google Scholar
Carrera, M.G., and Rigby, J.K., 1999, Biogeography of the Ordovician sponges: Journal of Paleontology, v. 73, p. 6–37.Google Scholar
Cisterna, C.E., 2001, Volcanismo subácueo en el Eopaleozoico del Sistema de Famatina, noroeste de Argentina: Revista de la Asociación Geológica Argentina, v. 56, p. 16–24.Google Scholar
Cocks, L.R.M., and Rong, J.Y., 1989, Classification and review of the brachiopod superfamily Plectambonitacea: Bulletin of the British Museum of Natural History (Geology), v. 45, p. 77–163.Google Scholar
Cocks, L.R.M., and Rong, J.Y., 2000, Strophomenida, in Kaesler, R.L., ed., Treatise on Invertebrate Paleontology, Part H (Revised), Brachiopoda 2: Boulder, Colorado, and Lawrence, Kansas, Geological Society of America (and University of Kansas Press), p. H216–H349.Google Scholar
Cocks, L.R.M., and Torsvik, T.H., 2002, Earth geography from 500 to 400 million years ago: A faunal and palaeomagnetic review: Journal of the Geological Society London, v. 159, p. 631–644. doi: 10.1144/0016-764901-118.Google Scholar
Congreve, C.R., Krug, A.Z., and Patzkowsky, M.E., 2015, Phylogenetic revision of the Strophomenida, a diverse and ecologically important Palaeozoic brachiopod order: Palaeontology, v. 58, p. 745–758. doi: 10.1111/pala.12177.Google Scholar
Cooper, G.A., 1956, Chazyan and related brachiopods: Smithsonian Miscellaneous Collections, v. 127, p. 1–1245.Google Scholar
Dahlquist, J.A., Pankhurst, R.J., Rapela, C.W., Galindo, C., Casquet, C., Fanning, C.M., Saavedra, J., Baldo, E., and González-Casado, J., 2005, New SHRIMP ages in the Sierra de Famatina, NW Argentina: Implications for the Famatinian Orogen, in Pankhurst, R.J., and Veiga, G.D., eds., Gondwana 12, Geological and Biological Heritage of Gondwana, Mendoza, Argentina, Abstracts, p. 123.Google Scholar
Davidson, T., 1868, On the earliest forms of Brachiopoda hitherto discovered in the British Palaeozoic rocks: Geological Magazine, v. 5, p. 303–316.Google Scholar
Dewing, K., 2004, Shell structure and its bearing on the phylogeny of Late Ordovician-Early Silurian strophomenoid brachiopods from Anticosti Island, Québec: Journal of Paleontology, v. 78, p. 275–286. doi: 10.1666/0022-3360(2004)078 0275:SSAIBO 2.0.C);2.Google Scholar
Eldredge, N., and Gould, S.J., 1972, Punctuated equilibria: An alternative to phyletic gradualism, in Schopf, T.J.M., ed., Models in Paleobiology: San Francisco, California, W.H. Freeman & Company, p. 82–115.Google Scholar
Erwin, D.H., 2007, Disparity: Morphological pattern and developmental context: Palaeontology, v. 50, p. 57–73. doi: 10.1111/j.1475-4983.2006.00614.x.Google Scholar
Espisúa, E., 1968, El Paleozoico inferior del Río de Las Chacritas, Dpto. de Jáchal, provincia de San Juan, con especial referencia al Silúrico: Revista de la Asociación Geológica Argentina, v. 23, p. 297–311.Google Scholar
Fang, Z.J., and Sánchez, T.M., 2012, Origin and early evolution of the Bivalvia, in Seldon, P.A., ed., Treatise [on Invertebrate Paleontology] Online, Part N (Revised), Mollusca 6, Bivalvia 1: Lawrence, Kansas, The Paleontological Institute, 21 p. doi: 10.17161/to.v0i0.4275.Google Scholar
Finney, S.C., 2007, The parautochthonous Gondwanan origin of the Cuyania (greater Precordillera) terrane of Argentina: A reevaluation of evidence used to support an allochthonous Laurentian origin: Geologica Acta, v. 5, p. 127–158.Google Scholar
Goloboff, P.A., and Catalano, S.A., 2016, TNT version 1.5, including a full implementation of phylogenetic morphometrics: Cladistics, v. 32, p. 221–238. doi: 10.1111/cla.12160.Google Scholar
Gómez, F.J., and Astini, R.A., 2015, Sedimentology and sequence stratigraphy from a mixed (carbonate-siliciclastic) rift to passive margin transition: The Early to Middle Cambrian of the Argentine Precordillera: Sedimentary Geology, v. 316, p. 39–61. doi: 10.1016/j.sedgeo.2014.11.003.Google Scholar
Gould, S.J., and Eldredge, N., 1977, Punctuated equilibria: The tempo and mode of evolution reconsidered: Paleobiology, v. 3, p. 115–151.Google Scholar
Gutiérrez-Marco, J.C., and Villas, E., 2007, Brachiopods from the uppermost Lower Ordovician of Peru and their palaeogeographical significance: Acta Palaeontologica Polonica, v. 52, p. 547–562.Google Scholar
Gutiérrez-Marco, J.C., Albanesi, G.L., Sarmiento, G.N., and Carlotto, V., 2008, An Early Ordovician (Floian) conodont fauna from the eastern Cordillera of Peru (Central Andean Basin): Geologica Acta, v. 6, p. 147–160.Google Scholar
Hall, J., and Clarke, J.M., 1892, An introduction to the study of the Brachiopoda: New York Geological Survey, v. 9, p. 1–367.Google Scholar
Harper, D.A.T., MacNiocaill, C., and Williams, S.H., 1996, The palaeogeography of Early Ordovician Iapetus terranes: An integration of faunal and palaeomagnetic constraint: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 121, p. 297–312.Google Scholar
Harper, D.A.T., Cocks, L.R.M., Popov, L.E., Sheehan, P.M., Bassett, M.G., Copper, P., Holmer, L.E., Jin, J., and Rong, J.Y., 2004, Brachiopods, in Webby, B.D., Droser, M., Paris, F., and Percival, I.G., eds., The Great Ordovician Biodiversification Event: New York, Columbia University Press, p. 157–178.Google Scholar
Harper, D.A.T., Owen, A.W., and Bruton, D.L., 2009, Ordovician life around Celtic fringes: Diversifications, extintions and migrations of brachiopod and trilobite faunas at middle latitudes, in Bassett, M.G., ed., Early Palaeozoic Peri-Gondwanan Terranes: New Insights from Tectonics and Biogeography: Geological Society of London Special Publication, v. 325, p. 157–170.Google Scholar
Harper, D.A.T., Rasmussen, C.M.Ø., Liljeroth, M., Blodgett, R.B., Candela, Y., Jin, J., Percival, I.G., Rong, J.Y., Villas, E., and Zhan, R., 2013, Biodiversity, biogeography and phylogeography of Ordovician rhynchonelliform brachiopods: The Geological Society London Memoirs, v. 38, p. 127–144. doi: 10.1144/M38.11.Google Scholar
Harper, D.A.T., Popov, L.E., and Holmer, L.E., 2017, Brachiopods: Origin and early history: Palaeontology, v. 60, p. 609–631. doi: 10.1111/pala.12307.Google Scholar
Harrington, H.J., and Leanza, A.F., 1957, Ordovician trilobites of Argentina: University of Kansas Department of Geology Special Publication, v. 1, 276 p.Google Scholar
Havlíček, V., and Branisa, L., 1980, Ordovician brachiopods of Bolivia: Rozpravy Ceskolovenské Akademie Věd, v. 90, p. 1–53.Google Scholar
Havliček, V., and Josopait, V., 1972, Articulate brachiopods from the Iberian Chais, northeast Spain (middle Cambrian-upper Cambrian-Tremadoc): Neues Jahrbuch für Geologie und Palaöntologie, v. 140, p. 328–353.Google Scholar
Heredia, S.E., Beresi, M., and Peralta, S., 2011, Estratigrafía y bioestratigrafía del Ordovícico Medio del río Las Chacritas, Precordillera Central de San Juan: INSUGEO Serie Correlación Geológica, v. 27, p. 18–26.Google Scholar
Herrera, Z.A., and Benedetto, J.L., 1987, El género Reinversella (Brachiopoda) en el Ordovícico temprano de la Precordillera Argentina: Actas 10th Congreso Geológico Argentino, San Miguel de Tucumán, Argentina, v. 3, p. 77–80.Google Scholar
Herrera, Z.A., and Benedetto., J.L., 1991, Early Ordovician brachiopod faunas from the Precordillera basin, western Argentina: Biostratigraphy and paleobiogeographical affinities, in Mackinnon, D.I., Lee, D.E., and Campbell, J.D, eds., Brachiopods through Time: Rotterdam, The Netherlands, A.A. Balkema, p. 283–301.Google Scholar
Hessland, I., 1949, Investigations of the Lower Ordovician of the Siljan District, Sweden: Notes on Swedish Ahtiella species: Bulletin of Geology, v. 33, p. 511–527.Google Scholar
Jones, O.T., 1928, Plectambonites and some allied genera: Memoirs of the Geological Survey of Great Britain, Palaeontology, v. 1, p. 367–527.Google Scholar
Keller, M., 1999, Argentine Precordillera: Sedimentary and plate tectonic history of a Laurentian crustal fragment in South America: Geological Society of America Special Paper 341, p. 1–131.Google Scholar
Keller, M., 2012, The Argentine Precordillera: A little American carbonate bank, in Derby, J.R., Fritz, R.D., Longacre, S.A., Morgan, W.A., and Sternbach, C.A., eds., The Great American Carbonate Bank: The geology and economic resources of the Cambrian–Ordovician sauk megasequence of Laurentia: AAPG Memoir, v. 98, p. 985–1000.Google Scholar
Kozlowski, R., 1927, Sur certains Orthides ordoviciens des environs de St. Petersburg: Bibliotheca Universitatis Libirae Polonae, v. 17, p. 3–21.Google Scholar
Lamansky, V.V., 1905, Die ältesten silurischen Schichten Russlands: Comité Géologique Mémoires St. Petersburg, n. ser, v. 20, p. 1–203.Google Scholar
Laurie, J.R., 1991, Articulate brachiopods from the Ordovician and lower Silurian of Tasmania, in Jell, P.A., ed., Australian Ordovician Brachiopod Studies: Association of Australasian Paleontologists Memoir, v. 11, p. 1–106.Google Scholar
Lavié, F.J., and Benedetto, J.L., 2016, Middle Ordovician (Darriwilian) linguliform and craniiform brachiopods from the Precordillera (Cuyania terrane) of west-central Argentina: Journal of Paleontology, v. 90, p. 1068–1080. doi: 10.1017/jpa.2016.111.Google Scholar
Lehnert, O.T., Bergström, S.M., and Vaccari, N.E., 1997, Arenig conodonts from the Famatina Range, northwestern Argentina: Faunal affinities and paleogeographic implications: 18th IAS Regional European Meeting of Sedimentology, Heidelberg, Germany, Gaea Heidelbergensis, Abstracts, v. 3, p. 216–217.Google Scholar
MacArthur, R.H., and Wilson, E.O., 1967, The Theory of Island Biogeography: New York, Princeton University Press, 203 p.Google Scholar
Mángano, M.G., and Buatois, L.A., 1996, Shallow marine event sedimentation in a volcanic arc-related setting: The Ordovician Suri Formation, Famatina Range, northwest Argentina: Sedimentary Geology, v. 105, p. 63–90.Google Scholar
Mángano, M.G., and Buatois, L.A., 1997, Slope apron deposition in an arc-related setting: The Vuelta de las Tolas Member (Suri Formation), Famatina Basin, northwest Argentina: Sedimentary Geology, v. 109, p. 155–180.Google Scholar
Mángano, M.G., Astini, R.A., Buatois, L.A., and Dávila, D.F., 2002, The Ordovician System in the Famatina Belt: Depositional and tectonic evolution, in Aceñolaza, F.G., ed., Aspects of the Ordovician System in Argentina: INSUGEO, Serie Correlación Geológica, v. 16, p. 295–312.Google Scholar
Mestre, A., and Heredia, S.E., 2012, Darriwilian species of Histiodella (Conodonta) in the Argentine Precordillera: Alcheringa, v. 36, p. 141–150. doi: 10.1080/03115518.2011.593334.Google Scholar
Mestre, A., and Heredia, S.E., 2013, La Zona de Yangtzeplacognathus crassus (Conodonta), Darriwiliano de la Precordillera Central, San Juan, Argentina: Ameghiniana, v. 50, p. 407–417. doi: 10.5710/AMGH.15.05.2013.558.Google Scholar
Muñoz, D.F., and Benedetto, J.L., 2016, The eoorthid brachiopod Apheoorthina in the Lower Ordovician of NW Argentina and the dispersal pathways along western Gondwana: Acta Palaeontologica Polonica, v. 61, p. 633–644. doi: 10.4202/app.00241.2016.Google Scholar
Neuman, R.B., 1971, An early Middle Ordovician brachiopod assemblage from Maine, New Brunswick, and northern Newfoundland, in Dutro, J.T., Jr., ed., Paleozoic Perspectives: A Paleontological Tribute to G. Arthur Cooper: Smithsonian Contributions to Paleobiology, v. 3, p. 113–124.Google Scholar
Neuman, R.B., 1976, Early Ordovician (late Arenig) brachiopods from Virgin Arm, New World Island, Newfoundland: Geological Survey of Canada Bulletin, v. 261, p. 11–61.Google Scholar
Neuman, R.B., 1984, Geology and paleobiology of islands in Iapetus Ocean: Review and implications: Geological Society of America Bulletin, v. 95, p. 118–1201.Google Scholar
Neuman, R.B., and Bates, D.E.B., 1978, Reassessment of Arenig and Llanvirn age (Early Ordovician) brachiopods from Anglesey, north-west Wales: Palaeontology, v. 21, p. 571–613.Google Scholar
Neuman, R.B., and Harper, D.A.T., 1992, Paleogeographic significance of Arenig-Llanvirn Toquima−Table Head and Celtic brachiopod assemblage, in Webby, B.D., and Laurie, J.R., eds., Global Perspectives on Ordovician Geology: Rotterdam, The Netherlands, Balkema, p. 241–254.Google Scholar
Öpik, A.A., 1930, Brachiopoda Protremata der Estländischen Ordovizischen Kukruse-Stufe: Acta et Commentationes Universitatis Tartuensis, v. A17, p. 1–262.Google Scholar
Öpik, A., 1932, Über die Plectellinen: Acta et Commentationes Universitatis Tartuensis, v. A23, p. 1–85.Google Scholar
Öpik, A., 1933, Über Plectamboniten: Acta et Commentationes Universitatis Tartuensis, v. A24, p. 1–79.Google Scholar
Öpik, A., 1934, Über Klitamboniten: Acta et Commentationes Universitatis Tartuensis, v. A26, no. 5, p. 1–239.Google Scholar
Ortega, G., Albanesi, G.L., and Frigerio, S., 2007, Graptolite-conodont biostratigraphy and biofacies of the Middle Ordovician Cerro Viejo succession, San Juan, Precordillera, Argentina: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 245, p. 245–264. doi: 10.1016/j.palaeo.2006.02.023.Google Scholar
Pander, C.H., 1830, Beiträge zur Geognosie des russichen Reiches: St. Petersburg, Gedrukt bei K. Kray, 165 p.Google Scholar
Pankhurst, R.J., Rapela, C.W., and Fanning, C.M., 2001, Age and origin of coeval TTG, I- and S-type granites in the Famatinian belt of NW Argentina: Transactions of the Royal Society of Edinburgh Earth Sciences, v. 91, p. 151–168. doi: 10.1017/S0263593300007343.Google Scholar
Peralta, S., Heredia, S.E., and Beresi, M., 1999, Upper Arenig–Lower Llanvirn sequence of the Las Chacritas River, Central Precordillera, San Juan Province, Argentina, in Kraft, P., and Fatka, O., eds., Quo vadis Ordovician?: Acta Universitatis Carolinae Geologica, v. 43, 1/2, p. 123–126.Google Scholar
Popov, L.E., Ghobadi Pour, M., Bassett, M.G., and Kebriaee-Zadeh, M., 2009, Billingsellidae and orthidae brachiopods: New insights into earliest Ordovician evolution and biogeography from northern Iran: Palaeontology, v. 52, p. 35–52. doi: 10.1111/j.1475-4983.2008.00833.x.Google Scholar
Ramos, V.A., 2004, Cuyania, an exotic block to Gondwana: Review of a historical success and the present problems: Gondwana Research, v. 7, p. 1009–1026. doi: 10.1016/S1342-937X(05)71081-9.Google Scholar
Sánchez, T.M., 1997, Additional Mollusca (Bivalvia and Rostroconchia) from the Suri Formation, Early Ordovician (Arenig), western Argentina: Journal of Paleontology, v. 71, p. 1046–1054.Google Scholar
Sánchez, T.M., and Babin, C., 1993, Un insolite mollusque bivalve, Catamarcaia n.g. de l’Arenig (Ordovicien inférieur) d’Argentine: Comptes Rendus de l’Academie des Sciences, v. 316, p. 265–271.Google Scholar
Schuchert, C., and Cooper, G.A., 1931, Synopsis of the brachiopod genera of the suborders Orthoidea and Pentameroidea, with notes on the Telotremata: American Journal of Science, ser. 5, v. 22, p. 241–255.Google Scholar
Serra, F., Albanesi, G.L., Ortega, G., and Bergström, S.M., 2015, Biostratigraphy and palaeoecology of Middle-Late Ordovician conodont and graptolite faunas of the Las Chacritas River section, Precordillera of San Juan, Argentina: Geological Magazine, v. 153, p. 813–829. doi: 10.1017/S0016756814000752.Google Scholar
Severgina, L.G., 1967, [New species and genera of the Ordovician brachiopods from the Sayano-Altai Hill district]: Uchenye Zapinski Tomskogo Gosudartvennogo Universiteta, v. 63, p. 120–140. (in Russian).Google Scholar
Sorrentino, L., Benedetto, J.L., and Carrera, M.G., 2009, Diversidad taxonómica y distribución de morfotipos de braquiópodos en la Zona de Ahtiella argentina (Ordovícico Medio), Formación San Juan, Precordillera Argentina: Ameghiniana, v. 46, p. 241–253.Google Scholar
Spjeldnaes, N., 1957, The Middle Ordovician of the Oslo region, Norway 8, Brachiopoda of the suborder Strophomenoidea: Norsk Geologisk Tidsskrift, v. 37, p. 1–214.Google Scholar
Suyarkova, A.A., and Koren, T.N., 2009, Advances in the Ordovician graptolite biostratigraphy of the St. Petersburg area, Russia: Estonian Journal of Earth Sciences, v. 58, p. 3–9. doi: 10.3176/earth.2009.1.01.Google Scholar
Thomas, W.A., and Astini, R.A., 1996, The Argentine Precordillera: A traveller from the Ouachita Embayment of North American Laurentia: Science, v. 273, p. 752–757.Google Scholar
Thomas, W.A., and Astini, R.A., 2003, Ordovician accretion of the Argentine Precordillera terrane to Gondwana: A review: Journal of South American Earth Sciences, v. 16, p. 67–79. doi: 10.1016/S0895-9811(03)00019-1.Google Scholar
Thomas, W.A., Astini, R.A., Mueller, P.A., Gehrels, G.E., and Wooden, J.L., 2004, Transfer of the Argentine Precordillera terrane from Laurentia: Constraints from detrital-zircon geochronology: Geology, v. 32, p. 965–968. doi: 10.1130/G20727.1.Google Scholar
Tolmacheva, T., Egerquist, E., Meidla, T., Vinn, O., and Holmer, L.E., 2003, Faunal composition and dynamics in the Middle Ordovician of the East Baltic: Geological Magazine, v. 140, p. 31–44. doi: 10.1017/S001675680200701X.Google Scholar
Topper, T.P., Harper, D.A.T., and Brock, G.A., 2013, Ancestral billingsellides and the evolution and phylogenetic relationships of early rhynchonelliform brachiopods: Journal of Systematic Palaeontology, v. 11, p. 821–833. doi: 10.1080/14772019.2012.728253.Google Scholar
Toro, B.A., and Brussa, E.D., 1997, Graptolitos de la Formación Suri (Arenig) en el Sistema de Famatina, Argentina: Revista Española de Paleontología, v. 12, p. 175–188.Google Scholar
Ulrich, O., and Cooper, G.A., 1936, New genera and species of Ozarkian and Canadian brachiopods: Journal of Paleontology, v. 10, p. 616–631.Google Scholar
Vaccari, N.E., 1995, Early Ordovician trilobite biogeography of Precordillera and Famatina, western Argentina: Preliminary results, in Cooper, J.D., Droser, M.L., and Finney, S.C., eds., Ordovician Odyssey: Short Papers for the Seventh International Symposium on the Ordovician System, Las Vegas, Nevada: Fullerton, California, Pacific Section Society for Sedimentary Geology, v. 77, p. 193–196.Google Scholar
Vaccari, N.E., and Waisfeld, B.G., 1994, Nuevos trilobites de la Formación Suri (Ordovícico temprano) en la región de Chaschuil, Provincia de Catamarca: Implicancias bioestratigráficas: Ameghiniana, v. 31, p. 73–86.Google Scholar
Waisfeld, B.G., and Vaccari, N.E., 2003, Trilobites, in Benedetto, J.L., ed., Ordovician Fossils of Argentina: Córdoba, Spain, Secretaría de Ciencia y Tecnología, Universidad Nacional de Córdoba, p. 295–409.Google Scholar
Waisfeld, B.G., Vaccari, N.E., Chatterton, B.D.E., and Edgecombe, G.D., 2001, Systematics of Shumardiidae (Trilobita), with new species from the Ordovician of Argentina: Journal of Paleontology, v. 75, p. 827–859. doi: 10.1666/0022-3360(2001)075 0827:SOSTWN 2.0.CO;2.Google Scholar
Waisfeld, B.G., Sánchez, T.M., Benedetto, J.L., and Carrera, M.G., 2003, Early Ordovician (Arenig) faunal assemblages from western Argentina: Biodiversification trends and paleogeographic constraints: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 196, p. 343–373. doi: 10.1016/S0031-0182(03)00464-4.Google Scholar
Webby, B.D., 1992, Ordovician island biotas: New South Wales record and global implications: Journal and Proceedings of the Royal Society of New South Wales, v. 124, p. 51–77.Google Scholar
Wiley, E O., 1979, Annotated Linnaean Hierarchy, with comments on natural taxa and competing systems: Systematic Zoology, v. 28, p. 308–337.Google Scholar
Williams, A., and Curry, G.B., 1985, Lower Ordovician Brachiopoda from the Tourmakeady Limestone, Co. Mayo, Ireland: Bulletin of the British Museum (Natural History) Geology Series, v. 38, p. 183–269.Google Scholar
Williams, A., and Hurst, J.M., 1977, Brachiopod evolution, in Hallam, A., ed., Patterns of Evolution: Amsterdam, Elsevier, p. 79–121.Google Scholar
Williams, A., Carlson, S.J., Brunton, C.H.C., Holmer, L.E., and Popov, L.E., 1996, A supra-ordinal classification of the Brachiopoda: Philosophical Transactions of the Royal Society of London, ser. B, v. 351, p. 1171–1193.Google Scholar
Xu, H.K., and liu, d.y., 1984, Late Early Ordovician brachiopods of southwestern China: Bulletin Nanjing Institute of Geology and Palaeontology, Academia Sinica, v. 8, p. 217–235.Google Scholar
Zalasiewicz, J.A., Taylor, L., Rushton, A.W.A., Loydell, D.K., Rickards, R.B., and Williams, M., 2009, Graptolites in British stratigraphy: Geological Magazine, v. 146, p. 785–850. doi: 10.1017/S0016756809990434.Google Scholar
Zhan, R.B., Jin, J., and Rong, J.Y., 2006, β-diversity fluctuations in Early–Mid Ordovician brachiopod communities of South China: Geological Journal, v. 41, p. 271–288. doi: 10.1002/gj.1040.Google Scholar
Zhan, R.B., Jin, J., Rong, J.Y, and Liang, Y., 2015, The earliest known strophomenoids (Brachiopoda) from early Middle Ordovician rocks of South China: Palaeontology, v. 56, p. 1121–1148. doi: 10.1111/pala.12039.Google Scholar