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Stem parrots (Aves, Halcyornithidae) from the Green River Formation and a combined phylogeny of Pan-Psittaciformes

Published online by Cambridge University Press:  14 July 2015

Daniel T. Ksepka ,
Julia A. Clarke  and
Lance Grande
Daniel T. Ksepka
Affiliation:
1Department of Marine, Earth and Atmospheric Sciences, North Carolina State University, Campus Box 8208, Raleigh, NC 27695, USA and Department of Paleontology, North Carolina Museum of Natural Sciences, Raleigh, NC 27695, USA,
Julia A. Clarke
Affiliation:
2Department of Geological Sciences, Jackson School of Geosciences, The University of Texas at Austin, University Station C1100, Austin, TX 78712, USA,
Lance Grande
Affiliation:
3Division of Collections and Research, Field Museum of Natural History, 1400 S Lake Shore Drive, Chicago, IL 60605, USA,
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Abstract

A new species of stem parrot is reported from the early Eocene Fossil Butte Member of the Green River Formation. Well-preserved specimens including a complete skeleton and referred partial postcranial skeleton reveal new osteological details of the clade Halcyornithidae and expand the known diversity of the Green River avifauna. Cyrilavis colburnorum n. sp. shares key morphologies with the smaller Green River halcyornithid Cyrilavis olsoni. A combined phylogenetic analysis of Pan-Psittaciformes using 101 morphological characters and sequence data from three genes (RAG-1, Z-chromosomal spindlin and cytochrome b) supports the placement of 1) Halcyornithidae, 2) a small clade uniting the Messel fossil taxon Psittacopes lepidus and an unnamed London Clay species, and 3) Quercypsittidae on branches successively closer to the crown clade Psittaciformes. Analysis of the morphological dataset alone yields a similar strict consensus tree with slightly less resolution (one additional branch collapsed). Relationships within the crown clade agree well with recent hypotheses in supporting a basal divergence between the New Zealand Nestoridae and all other extant parrots, and a second major divergence between Cacatuidae and Psittacidae. When fossils are taken into account, many previously proposed synapomorphies of extant parrots are instead optimized as synapomorphies of more inclusive groups including both stem fossil taxa and living taxa. Fossil evidence is consistent with a more recent origin of crown clade Psittaciformes than predicted from divergence dating analyses, and supports environmental shifts and dispersal as key drivers in parrot biogeography.

Type
Research Article
Information
Journal of Paleontology , Volume 85 , Issue 5 , September 2011 , pp. 835 - 852
Copyright
Copyright © The Paleontological Society 

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