Hostname: page-component-78c5997874-m6dg7 Total loading time: 0 Render date: 2024-11-20T01:28:16.864Z Has data issue: false hasContentIssue false

Silurian acastacean trilobites of the Americas

Published online by Cambridge University Press:  20 May 2016

Gregory D. Edgecombe*
Affiliation:
Department of Geology, University of Alberta, Edmonton T6G 2E3, Canada

Abstract

Species of the superfamily Acastacea constitute a minor element of North and South American Silurian trilobite faunas. Phacopidina? obsoleta (Ulrich and Delo) (Llandovery, Tennessee) provides the sole Silurian record of the “kloucekiine” grade (Acastacea s.l.); a lectotype is designated for this species. Acaste birminghamensis Norford (mid Llandovery, Alabama) lacks synapomorphies of post-Llandovery Acastidae s.s. (=Acastinae + “Acastavinae” + Asteropyginae), and is the basis for Llandovacaste n. gen. (Acastidae s.l.). A well-preserved sample of Andinacaste cf. A. ledgrandi Eldredge and Braniša from the Catavi Formation (Ludlow–Přídolí), Bolivia, displays apomorphic characters of the genal spines and hypostome shared with Devonian calmoniids. Coaptative structures, including vincular furrow/pits and “bifurcated” thoracic pleural tips, are documented for Andinacaste; similar enrollment morphologies arose in other acastomorph calmoniids. Poorly known Llandovery acastaceans from Paraguay and Venezuela may represent Andinacaste. Acaste zerinae n. sp. from the Pembroke Formation (Přídolí) of Maine is closely comparable to British late Wenlock A. downingiae (Murchison). The Australian Gedinnian acastine placed in Phacopinae indet. longisulcata (Shergold) is designated Acaste lokii n. sp.

Type
Research Article
Copyright
Copyright © The Paleontological Society 

Access options

Get access to the full version of this content by using one of the access options below. (Log in options will check for institutional or personal access. Content may require purchase if you do not have access.)

References

Babcock, L. E., Gray, J., Boucot, A. J., Himes, G. T., and Siegele, P. K. 1990. First Silurian conulariids from Paraguay. Journal of Paleontology, 64:897902.Google Scholar
Balashova, E. A. 1968. Trilobity skal'skogo i borshchovskogo gorizontov Podolii, p. 95122. In Balashov, Z. G. (ed.), Silurijsko-devonskaya Fauna Podolii. Izdatel'stvo Leningradskogo Universiteta, Leningrad.Google Scholar
Baldis, B. A., and Blasco, G. 1976. Jujuyops n. gen., p. 215217. Harringtonacasten. gen., p. 217–220. In Baldis, B. A., Benedetto, J. L., Blasco, G., and Martel, M. E. (eds.), Trilobites Silúrico-Devónicos de la Sierra de Zapla (Noroeste de Argentina). Ameghiniana, 13.Google Scholar
Baldis, B. A., and González, S. B. 1981. Conocimiento actual de la distribución de los trilobites acastidos de Sudamérica. Anais II Congreso Latino-Americano Paleontológia:5969.Google Scholar
Bancroft, B. B. 1949. Upper Ordovician trilobites of zonal value in southeast Shropshire. Proceedings of the Royal Society of London, B, 136:291315.Google Scholar
Bastin, E. S., and Williams, H. S. 1913. Eastport Folio, Maine. U.S. Geological Survey, Folio 192.Google Scholar
Benedetto, J. L., and Martel, M. A. 1976. Australoacaste n. gen., p. 211215. In Baldis, B. A., Benedetto, J. L., Blasco, G., and Martel, M. E. (eds.), Trilobites Silúrico-Devónicos de la Sierra de Zapla (Nordeste de Argentina). Ameghiniana, 13.Google Scholar
Benedetto, J. L., Racheboeuf, P. R., Herrera, Z., Brussa, E. D., and Toro, B. A. 1992. Brachiopodes et biostratigraphie de la Formation de Los Espejos, Siluro-Dévonien de la Précordillère Argentine. Geobios, 25:599637.Google Scholar
Berry, W. B. N., and Boucot, A. J. 1970. Correlation of the North American Silurian Rocks. Geological Society of America, Special Paper 102, 289 p.Google Scholar
Boucot, A. J. 1972. Silurian of Venezuela, p. 5354. In Berry, W. B. N. and Boucot, A. J. (eds.), Correlation of the South American Silurian Rocks. Geological Society of America, Special Paper 133.CrossRefGoogle Scholar
Butts, C. 1940. Geology of the Appalachian Valley in Virginia. Part I. Geologic text and illustrations. Virginia Geological Survey, Bulletin 52, 568 p.Google Scholar
Campbell, K. S. W. 1967. Trilobites of the Henryhouse Formation (Silurian) in Oklahoma. Oklahoma Geological Survey, Bulletin 115, 68 p.Google Scholar
de Carvalho, M. da G. P., and Edgecombe, G. D. 1991. Lower–Early Middle Devonian calmoniid trilobites from Mato Grosso, Brazil, and related species from Paraná. American Museum Novitates, 3022:113.Google Scholar
Clarke, J. M. 1913. Fosseis devonianos do Paraná. Serviço Geologico e Mineralogico de Brazil, Monographia 1:1353.Google Scholar
Clarkson, E. N. K. 1965. Schizochroal eyes and vision in some Silurian acastid trilobites. Palaeontology, 9:129.Google Scholar
Cooper, M. R. 1982. A revision of the Devonian (Emsian–Eifelian) Trilobita from the Bokkeveld Group of South Africa. Annals of the South African Museum, 89, 174 p.Google Scholar
Dean, W. T. 1961. The Ordovician trilobite faunas of south Shropshire, II. Bulletin of the British Museum (Natural History), Geology, 5:311358.Google Scholar
Dean, W. T. 1967. The correlation and trilobite fauna of the Bedinan Formation (Ordovician) in south-eastern Turkey. Bulletin of the British Museum (Natural History), Geology, 15:81123.Google Scholar
Delo, D. M. 1935. A revision of the phacopid trilobites. Journal of Paleontology, 9:402420.Google Scholar
Delo, D. M. 1940. Phacopid trilobites of North America. Geological Society of America, Special Paper 29, 135 p.Google Scholar
Destombes, J. 1972. Les trilobites du sous-ordre des Phacopina de l'Anti-Atlas (Maroc). Notes et Mémoires du Service Géologique du Maroc, 240:1113.Google Scholar
Destombes, J., and Henry, J.-L. 1987. Trilobites Calmoniidae de l'Ordovicien supérieur du Maroc et les origines de la Province Malvino-Cafre. Lethaia, 20:129139.Google Scholar
Edgecombe, G. D. 1991. Morocconites Struve, 1989, a Devonian acastine trilobite (Calmoniidae: Acastinae). American Museum Novitates, 2998:17.Google Scholar
Edgecombe, G. D. 1993. Calmoniid trilobites from the Devonian Fox Bay Formation, Falkland Islands: systematics and biogeography. In Landing, E. (ed.), Studies in Paleontology and Stratigraphy in Honor of Donald W. Fisher. New York State Museum Bulletin, 481:5568.Google Scholar
Eldredge, N. 1971. Patterns of cephalic musculature in the Phacopina (Trilobita) and their phylogenetic significance. Journal of Paleontology, 45:5267.Google Scholar
Eldredge, N. 1979. Cladism and common sense, p. 165198. In Cracraft, J. and Eldredge, N. (eds.), Phylogenetic Analysis and Paleontology. Columbia University Press, New York.Google Scholar
Eldredge, N., and Braniša, L. 1980. Calmoniid trilobites of the Lower Devonian Scaphiocoelia Zone of Bolivia, with remarks on related species. Bulletin of the American Museum of Natural History, 165:181289.Google Scholar
Eldredge, N., and Cracraft, J. 1980. Phylogenetic Patterns and the Evolutionary Process. Method and Theory in Comparative Biology. Columbia University Press, New York, 349 p.Google Scholar
Eldredge, N., and Ormiston, A. R. 1979. Biogeography of Silurian and Devonian trilobites of the Malvinokaffric Realm, p. 147167. In Boucot, A. J. and Gray, J. (eds.), Historical Biogeography, Plate Tectonics and the Changing Environment. Oregon State University Press, Corvallis.Google Scholar
Gandl, J. 1972. Die Acastavinae und Asteropyginae (Trilobita) Keltiberiens (NE-Spanien). Abhandlungen der Senckenbergischen Gesellschaft, 530:1184.Google Scholar
Goldfuss, A. 1843. Systematische Übersicht der Trilobiten und Beschreibung einiger neuen Arten derselben. Neues Jahrbuch für Mineralogie, Geognosie und Petrefaktenkunde, 1843:537567.Google Scholar
Haas, W. 1968. Trilobiten aus dem Silur und Devon von Bithynien (NW-Turkei). Palaeontographica (A), 130:60207.Google Scholar
Haas, W., and Mensink, H. 1969. Asteropyginae aus Afghanistan (Trilobita). Akademie der Wissenschaften und der Literatur, Abhandlungen der Mathematisch-Naturwissenschaftlichen Klasse, 1969:160.Google Scholar
Hall, J. 1860. Description of new species of fossils from the Silurian rocks of Nova Scotia. Canadian Naturalist, 5:144169.Google Scholar
Hammann, W. 1974. Phacopina und Cheirurina (Trilobita) aus dem Ordovizium von Spanien. Senckenbergiana lethaea, 55:1151.Google Scholar
Harrington, H. J. 1972. Silurian of Paraguay, p. 4150. In Berry, W. B. N. and Boucot, A. J. (eds.), Correlation of the South American Silurian Rocks. Geological Society of America, Special Paper 133.CrossRefGoogle Scholar
Hawle, I., and Corda, A. J. C. 1847. Prodrom einer monographie der böhmischen Trilobiten. Abhandlungen der Königlichen Böhemischen Gesellschaft der Wissenschaften, 5:1176.Google Scholar
Henry, J.-L. 1980. Trilobites ordoviciens du Massif Armoricain. Mémoires de la Société Géologique et Minérologique de Bretagne, 22:1250.Google Scholar
Henry, J.-L., and Dolambi, T. 1990. Coaptations chez les trilobites Placoparia (Ordovicien) et Acastoides (Dévonien): un exemple de convergence. Geobios, 23:581586.Google Scholar
Hollard, H. 1963. Les Acastella et quelques autres Dalmanitacea du Maroc présaharien. Leur distribution verticale et ses conséquences pour l'étude de la limite Silurien–Dévonien. Notes et Mémoires du Service Géologique du Maroc, 176:166.Google Scholar
Holloway, D. J., and Neil, J. V. 1982. Trilobites from the Mount Ida Formation (Late Silurian–Early Devonian), Victoria. Proceedings of the Royal Society of Victoria, 94:133154.Google Scholar
Hupé, P. 1953. Classe des Trilobites, p. 44246. In Piveteau, J. (ed.), Traité de Paléontologie, Tome III. Masson et Compagnie, Paris.Google Scholar
Lespérance, P. J. 1991. Vincular furrows in some Early Silurian and Devonian Phacopidae (Trilobita), predominantly from North America. Journal of Paleontology, 65:276294.Google Scholar
Lieberman, B. S., Edgecombe, G. D., and Eldredge, N. 1991. Systematics and biogeography of the “Malvinella group,” Calmoniidae (Trilobita, Devonian). Journal of Paleontology, 65:824843.Google Scholar
Méndez-Alzola, R. 1934. Contribución al conocimiento de la fauna Devónica de Rincón de Alonso. Boletin del Instituto Geológico Perforaciones, 21:2154.Google Scholar
Méndez-Alzola, R. 1938. Fósiles devónicos del Uruguay. Boletin del Instituto Geológico del Uruguay, 24:3115.Google Scholar
Morris, S. F. 1988. A review of British trilobites, including a synoptic review of Salter's monograph. Palaeontographical Society Monograph, 574, 316 p.Google Scholar
Murchison, R. I. 1839. The Silurian system, founded on geological researches in the counties of Salop, Hereford, Radnor, Montgomery, Caermarthen, Brecon, Pembroke, Monmouth, Gloucester, Worcester and Stafford; with descriptions of the coalfields and overlying formations. John Murray, London, 768 p.Google Scholar
Norford, B. S. 1972. Acaste birminghamensis, a new Lower Silurian trilobite from Alabama. Journal of Paleontology, 46:348352.Google Scholar
Orbigny, A. d'. 1842. Voyage dans l'Amerique méridionale, 1826–1833, 3, 4 (Paléontologie). Bertrand, Paris, 188 p.Google Scholar
Pillet, J. 1961. Contribution à l'étude des faunes de trilobites de Zemmour (Mauritanie septentrionale). Université de Dakar, Annales de la Faculté des Sciences, 5:93119.Google Scholar
Pillet, J. 1990. Les faunes trilobitiques de l'Ordovicien Supérieur en Anjou. Mémoire de la Société d'études scientifiques de l'Anjou, 8:123.Google Scholar
Popp, M. T. B. 1989. Paranacaste, um novo gênero de Trilobita de Formação Ponta Grossa, Bacia do Paraná. Anais 11° Congresso Brasileiro de Paleontologia. Sociedade Brasileira de Paleontologia, 1:1935.Google Scholar
Přibyl, A., and Vaněk, J. 1980. Neue Erkenntnisse über einige Trilobiten aus dem böhmischen Ordovizium. Časopis pro mineralogii a geologii, 25:263274.Google Scholar
Price, D. 1974. Trilobites from the Sholeshook Limestone (Ashgill) of South Wales. Palaeontology, 17:841868.Google Scholar
Rabano, I. 1989. Trilobites del Ordovícico medio del sector meridional de la zona Centroibérica español. Publicacions Especiales del Boletin Geológico y Minero, 233 p.Google Scholar
Ramsköld, L. 1985. Silurian phacopid and dalmanitid trilobites from Gotland. Stockholm Contributions in Geology, 40:162.Google Scholar
Reed, F. R. C. 1925. Some new Silurian trilobites. Geological Magazine, 62:6776.Google Scholar
Rehfeld, U., and Mehl, J. 1989. Andinodesma radicostata n. gen. n. sp., a grammysiid taxon from the Lower Devonian Catavi Formation (Bolivia) and its autecological and phylogenetic implications. Paläontologisches Zeitschrift, 63:263279.Google Scholar
Richter, R., and Richter, E. 1952. Phacopacea von der Grenze Emsium/Eifelium (Tril.). Senckenbergiana, 33:79108.Google Scholar
Richter, R., and Richter, E. 1954. Die Trilobiten des Ebbe-Sattels und zu vergleichende Arten (Ordovizium, Gotlandium/Devon). Abhandlungen der Senckbergischen, Naturforschenden Gesellschaft, 488:176.Google Scholar
Salter, J. 1864. A monograph of the British trilobites from the Cambrian, Silurian, and Devonian formations. Palaeontographical Society Monograph, 80 p.Google Scholar
Schrank, E. 1970. Die Trilobiten des Silurs der Bohrung Leba 1 (Ostsecküste der VR Polen). Bericht der deutschen Gesellschaft geologicher Wissenschaft (A), 15:573586.Google Scholar
Sedgwick, A., and M'Coy, F. 1851. A Synopsis of the Classification of the British Palaeozoic Rocks, with a Systematic Description of the British Palaeozoic Fossils found in the Geological Museum of the University of Cambridge. London, Cambridge, 184 p.Google Scholar
Shergold, J. 1966. A revision of Acaste downingiae (Murchison) and related trilobites. Palaeontology, 9:183207.Google Scholar
Shergold, J. 1967. A reappraisal of the North American species of the Siluro-Devonian trilobite genus Scotiella . Postilla, 112:120.Google Scholar
Shergold, J. 1968. On the occurrence of the trilobite genera Acaste and Acastella in Victoria. Proceedings of the Royal Society of Victoria, 81:1930.Google Scholar
Smeenk, Z. 1983. Devonian trilobites of the southern Cantabrian Mountains (northern Spain) with a systematic description of the Asteropyginae. Leidse Geologische Mededelingen, 52:383511.Google Scholar
Šnajdr, M. 1982. Bohemian representatives of the trilobite genera Kloucekia Delo, Phacopidina Bancroft, Sokhreta Hupé and Dalmanitina Reed. Věstnik Ústředniho ústavu geologického, 57:179182.Google Scholar
Šnajdr, M. 1987. New Bohemian Ordovician Dalmanitidae and Calmonidae (Trilobita). Věstnik Ústředniho ústavu geologického, 62:271277.Google Scholar
Struve, W. 1958. Acastavinae n. subfam. Senckenbergiana lethaea, 39:221226.Google Scholar
Struve, W. 1959. Phacopina, p. 461462. Calmoniinae, p. 483–487. In Moore, R. C. (ed.), Treatise on Invertebrate Paleontology. Part O. Arthropoda 1. Geological Society of America, University of Kansas Press, Lawrence.Google Scholar
Struve, W. 1989. Trilobites, p. 559569. In Boucot, A. J., McLure, H. A., Alvarez, F., Ross, J. R. P., Taylor, D. W., Struve, W., Savage, N. M., and Taylor, S. (eds.), New Devonian fossils from Saudi Arabia and their biogeographical affinities. Senckenbergiana lethaea, 69:535–597.Google Scholar
Swartz, F. M. 1925. The Devonian fauna of Bolivia. Johns Hopkins University Studies in Geology, 6:2968.Google Scholar
Thomas, A. T., Owens, R. M., and Rushton, A. W. A. 1984. Trilobites in British stratigraphy. Geological Society, London, Special Report, 16, 78 p.Google Scholar
Timm, J. 1981. Zur trilobitenstratigraphie des Silur/Devon-Grenzbereiches im Ebbe-Antiklinorium (Rheinishes Schiefergebirge). Mitteilungen aus dem Geologisch-Paläontologischen Institut der Universität Hamburg, 50:91108.Google Scholar
Tomczykowa, E. 1962. O trylobicie Acastella prima n. sp. Kwartalnik Geologiczny, 6:260266.Google Scholar
Tomczykowa, E., and Witwicka, E. 1974. Stratigraphic correlation of Podolasian deposits on the basis of ostracodes and trilobites in the Peri-Baltic area of Poland (Upper Silurian). Biuletyn Instytut Geologiczny, 276:5584.Google Scholar
de Tromelin, G. 1877. Étude de la faune du grès silurian de May, Jurques, Camprandré, Mont Robert, etc. (Calvados) avec les observations sur divers fossiles paléozoïques de l'Ouest de la France. Bulletin de la Société Linnéenne de Normandie, 1:582.Google Scholar
Ulrich, E. O., and Bassler, R. S. 1923. American Silurian formations, p. 233270. In Maryland Geological Survey. Silurian. John Hopkins Press, Baltimore.Google Scholar
Ulrich, E. O., and Delo, D. M. 1940. Scotiella obsoleta n. sp., p. 3435. In Delo, D. M. (ed.), Phacopid trilobites of North America. Geological Society of America, Special Paper 29.Google Scholar
Waisfeld, B. G., Toro, B. A., and Brussa, E. D. 1988. Trilobites Silúricos de la Formación Los Espejos, Sector Occidental del Cerro del Fuerte, Precordillera da San Juan, Argentina. Ameghiniana, 25:305320.Google Scholar
Wolfart, R. 1961. Stratigraphie und fauna des älteren Paläozoikums (Silur, Devon) in Paraguay. Geologisches Jahrbuch, 78:29102.Google Scholar
Wolfart, R. 1968. Die Trilobiten aus dem Devon Boliviens und ihre Bedeutung für Stratigraphie und Tiergeographie, p. 5201. In Wolfart, R. and Voges, A. (eds.), Beiträge zur Kenntnis des Devons von Bolivien. Beihefte zum Geologischen Jahrbuch, 74.Google Scholar
Wright, A. J. 1990. Acastid trilobites from the Baton Formation (Early Devonian), New Zealand. New Zealand Journal of Geology and Geophysics, 33:4953.Google Scholar