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A new ptyctodont (Placodermi) from the Upper Devonian Martin Formation of Northern Arizona, and an analysis of ptyctodont phylogeny

Published online by Cambridge University Press:  20 May 2016

Heidemarie G. Johnson
Affiliation:
Department of Geology, Box 4099, Northern Arizona University, Flagstaff 86011
David K. Elliott
Affiliation:
Department of Geology, Box 4099, Northern Arizona University, Flagstaff 86011

Abstract

Ptyctodonts from the Upper Devonian Martin Formation on Mt. Elden, northern Arizona, were hitherto known only from tooth plates ascribed to Ptyctodus bradyi. Recent collections include dental elements of P. bradyi as well as body and skull plates and dental elements of a previously unknown ptyctodont, described here as Denisonodus plutonensis new genus and species. Comparison with other ptyctodonts suggests that the new genus has close affinities to other North American forms. As the holotype of Ptyctodus bradyi has been lost a neotype is designated here.

Type
Research Article
Copyright
Copyright © The Paleontological Society 

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References

Beus, S. S. 1973. Devonian Stratigraphy and Paleogeography along the western Mogollon Rim, Arizona. Museum of Northern Arizona Bulletin, 49:39 p.Google Scholar
Beus, S. S. 1978. Late Devonian (Frasnian) invertebrate fossils from the Jerome Member of the Martin Formation, Verde Valley, Arizona. Journal of Paleontology, 52:4954.Google Scholar
Beus, S. S. 1980. Late Devonian (Frasnian) Paleogeography and Paleoenvironments in northern Arizona, p. 5556. In Fouch, T.D. and Magathan, E.R. (eds), Rocky Mountain Section S.E.P.M., Paleozoic Paleogeography of West-Central United States.Google Scholar
Brady, L. F. 1933. New Devonian area in northern Arizona. Pan-Am Geologist, 60:303304.Google Scholar
Brady, L. F. 1934. Elden Mt. Museum of Northern Arizona, Museum Notes, 7:912.Google Scholar
Cooper, G. A., Butt, C., Caster, K. E., Chadwick, G. H., Goldring, W., Kindley, E. M., Kirk, E., Merriam, C. W., Swartz, F. M., Warren, P. S., Warthin, A. S., and Willard, B. 1942. Correlation of the Devonian Sedimentary Formations of North America. Geological Society of America Bulletin, 52:17291794.CrossRefGoogle Scholar
Denison, R. H. 1978. Placodermi. Handbook of Paleoichthyology, Gustav Fischer Verlag, Stuttgart, 128 p.Google Scholar
Denison, R. H. 1983. Further Consideration of Placoderm Evolution. Journal of Vertebrate Paleontology, 3:6983.Google Scholar
Denison, R. H. 1985. A new ptyctodont placoderm, Ptyctodopsis, from the Middle Devonian of Iowa. Journal of Paleontology, 59:511522.Google Scholar
Elliott, D. K., and Ruddell, M. 1988. Eldenosteus, an Upper Devonian placoderm from Mt. Elden, Arizona. 41 st Annual Symposium on Southwestern Geology and Paleontology. Museum of Northern Arizona:4.Google Scholar
Elliott, D. K., Ruddell, M. W., and Johnson, H. G. 1989. Devonian Placoderms from Flagstaff, Arizona. 33rd Annual Meeting of the Arizona-Nevada Academy of Science:46.Google Scholar
Forey, P. L. 1980. Latimeria: a paradoxical fish. Proceedings of the Royal Society of London, Series B, 208:369384.Google Scholar
Forey, P. L., and Gardiner, B. G. 1986. Observations on Ctenurella (Ptyctodontida) and the Classification of placoderm fishes. Zoological Journal of the Linnean Society, 86:4374.Google Scholar
Goloboff, P. A. 1993. NONA. INSUE-Fundacion e Instituto Miguel Lillo, Tucuman, Argentina.Google Scholar
Goujet, D. F. 1984. Placoderm Interrelationships: A new Interpretation with a short review of Placoderm Classifications. Proceedings of the Linnean Society of N.S.W., 107:211243.Google Scholar
Goujet, D. F. and Young, G. C. 1995. Interrelationships of Placoderms revisited, p. 8996. In Lelievre, H., Wenz, S., Blieck, A., and Cloutier, R. (eds). Premiers Vertebrates et vertebrates inferieurs. Geobios Memoir Special, 19.Google Scholar
Gross, W. 1932. Die Arthrodira Wildungens. Geologische und Palaeontologische Abhandlungen, 19:561.Google Scholar
Gross, W. 1933. Die Wirbeltiere des rheinischen Devons. Abhandlungen Preussischer Geologischen Landesanstalt, 154:183.Google Scholar
Gross, W. 1961. Lunaspis broilii and Lunaspis heroldi aus dem Hunsrueckschiefer (Unterdevon, Rheinland). Notizblatt des Hessischen Landesamtes fuer Bodenforschung zu Wiesbaden, 89:1743.Google Scholar
Holm, R. F. 1988. Geologic Map of San Francisco Mountain, Elden Mountain and Dry Lake Hills, Coconino County, Arizona. Miscellaneous Investigations Series, Map 1-1663, US Geological Survey.Google Scholar
Hussakof, L. 1942. Fishes from the Devonian of Arizona. American Museum of Natural History Novitates, 1186:19.Google Scholar
Jaekel, O. 1903. Ueber Rhamphodus nov.gen., einen neuen devonischen Holocephalen von Wildungen, Sitzungsberichte der Gesellschaft naturforschender Freunde Berlin, 8:383393.Google Scholar
Jaekel, O. 1906. Einige Beitraege zur Morphologie der aeltesten Wirbeltiere. Sitzungsberichte der Gesellschaft naturforschender Freunde Berlin, 7:180189.Google Scholar
Jaekel, O. 1919. Die Mundbildung der Placodermen. Sitzungsberichte der Gesellschaft naturforschender Freunde Berlin, 3:73110.Google Scholar
Jaekel, O. 1929. Die Morphogenie der aeltesten Wirbeltiere. Monographien zur Geologie und Palaeontologie, Berlin, 3:1198.Google Scholar
Johnson, H. G., and Elliott, D. K. 1995. A redescription of Eldenosteus arizonensis (Placodermi: Arthrodira) from the Upper Devonian Martin Formation of Northern Arizona. Journal of Vertebrate Paleontology, 15:221234.Google Scholar
Klapper, G., and Johnson, J. G. 1980. Endemism and dispersal of Devonian conodonts. Journal of Paleontology, 54:400455.Google Scholar
Lelievre, H. 1986. Les placodermes et leurs relations phyletiques avec les autres gnathostomes. Oceanis, 12:139149.Google Scholar
Liu, Y.-H. 1973. On the new forms of Polybranchiaspiformes and Petalichthyids from Devonian of south-west China. Vertebrata PalAsiatica, 11:132143.Google Scholar
Liu, Y.-H. 1991. On a new petalichthyid Eurycaraspis incilis gen.et sp.nov., (Placodermi, Pisces) from the Middle Devonian of Zhanyi, Yunnan, p. 139177. In Chang, M.-M., Liu, Y.-H., and Zhang, G.-R. (eds), Early Vertebrates and Related Problems of Evolutionary Biology. Science Press, Beijing, China.Google Scholar
Long, J. A. 1988. Late Devonian fishes from Gogo, Western Australia. National Geographic Research, 4:436450.Google Scholar
M'Coy, F. 1848. On some new fossil fish of the Carboniferous Period. Annual Magazine of Natural History, 2:110.CrossRefGoogle Scholar
Miles, R. S. 1964. On some coccosteomorph arthrodires from the Devonian of Arizona. Arkiv for Zoologi, 22:427460.Google Scholar
Miles, R. S. 1967. Observations on the ptyctodont fish Rhamphodopsis Watson. Zoological Journal of the Linnean Society, 47:99120.Google Scholar
Miles, R. S., and Young, G. C. 1977. Placoderm interrelationships reconsidered in the light of new ptyctodontids from Gogo, Western Australia. Linnean Society Symposium Series, 4:123198.Google Scholar
Orvig, R. 1957. Notes on some Paleozoic lower vertebrates from Spitsbergen and North America. Norsk Geologisk Tidskrift, 37:285353.Google Scholar
Orvig, R. 1962. Y a-t-il une relation directe entre les arthrodires ptyctodontides et les holocephales? Colloques Internationaux du Centre National de la Recherche Scientifique, 104:4961.Google Scholar
Pander, C. H. 1858. Die Ctenodipterinen des devonischen systems. Archiv fuer wissenschaftliche Kunde von Russland:65 p.Google Scholar
Stensio, E. A. 1925. On the head of the Macropetalichthyids with certain remarks on the head of other arthrodires. Field Museum of Natural History, 232:87197.Google Scholar
Stoyanow, A. A. 1936. Correlation of Arizona Paleozoic formations. Geological Society of America Bulletin, 47:459540.CrossRefGoogle Scholar
Watson, D. M. S. 1938. On Rhamphodopsis, a ptyctodont from the Middle Old Red Sandstone of Scotland. Transactions of the Royal Society of Edinburgh, 59:397410.Google Scholar
Westoll, T. S. 1962. Ptyctodontid fishes and the ancestry of Holocephali. Nature, 194:949952.Google Scholar
Woodward, A. S. 1891. Catalogue of the fossil fishes in the British Museum (Natural History). 567 p.Google Scholar
Woodward, A. S. 1941. The head shield of a new macropetalichthyid fish (Notopetalichthyis hillsi, gen.et sp.nov.) from the Middle Devonian of Australia. Annals and Magazine of Natural History, 8:9196.Google Scholar
Young, G. C. 1986. The relationships of placoderm fishes. Zoological Journal of the Linnean Society, 88:157.Google Scholar