Hostname: page-component-78c5997874-s2hrs Total loading time: 0 Render date: 2024-11-04T21:27:26.125Z Has data issue: false hasContentIssue false

New and revised cyrtospiriferid (Spiriferida) brachiopods from the lower Famennian (Upper Devonian) of Armenia

Published online by Cambridge University Press:  31 March 2022

Vahram Serobyan
Affiliation:
Institute of Geological Sciences of the National Academy of Sciences of the Republic of Armenia-Yerevan, Armenia, 24A, Marshal Baghramyan Avenue, Yerevan 0019, Republic of Armenia , University of Lille, CNRS, UMR 8198- Evo-Eco-Paleo, F-59000 Lille, France , ,
Taniel Danelian*
Affiliation:
University of Lille, CNRS, UMR 8198- Evo-Eco-Paleo, F-59000 Lille, France , ,
Catherine Crônier
Affiliation:
University of Lille, CNRS, UMR 8198- Evo-Eco-Paleo, F-59000 Lille, France , ,
Araik Grigoryan
Affiliation:
Institute of Geological Sciences of the National Academy of Sciences of the Republic of Armenia-Yerevan, Armenia, 24A, Marshal Baghramyan Avenue, Yerevan 0019, Republic of Armenia ,
Bernard Mottequin
Affiliation:
Operational Directorate Earth and History of Life, Royal Belgian Institute of Natural Sciences, rue Vautier 29, B 1000 Brussels, Belgium
*
*Corresponding author.

Abstract

The Upper Devonian sedimentary sequences of Central Armenia, which mainly consist of shallow water, mixed carbonate-siliciclastic deposits, contain abundant and diverse brachiopods that are dominated by spiriferides. Based on newly collected material from the lower Famennian Aramazdospirifer orbelianus brachiopod zone (coeval to the Palmatolepis crepida conodont zone) of Armenia, we here introduce two new cyrtospiriferid genera and fully document their type species, including their intraspecific variability. Pentagonospirifer n. gen. is a monospecific genus assigned to the subfamily Cyrtospiriferinae that is currently known only from the lower Famennian of Armenia; it is likely that its type species, P. abrahamyanae n. sp., evolved from the species Cyrtospirifer verneuili sensu Abrahamyan, known from the Frasnian and lower Famennian of Armenia. The second new genus, Tornatospirifer n. gen., is assigned to the subfamily Cyrtiopsinae and defined on the basis of one of the most biostratigraphically valuable cyrtospiriferid species, namely T. armenicus n. comb., described previously from the lower Famennian Aramazdospirifer orbelianus Zone of Armenia. It is likely that it evolved from a Frasnian ancestral stock of Tiocyrspis Sartenaer, known from northwestern Europe, which probably migrated during the early Frasnian to the north Gondwanan margin. A neotype is selected for Abrahamyan's species because the type material is lost. Epibionts (Cornulites, Hederella) attached to brachiopods are also documented for the first time from the Upper Devonian of Armenia.

UUID: http://zoobank.org/923d232f-cd6a-48bb-8a05-27c689058b65.

Type
Articles
Copyright
Copyright © The Author(s), 2022. Published by Cambridge University Press on behalf of The Paleontological Society

Access options

Get access to the full version of this content by using one of the access options below. (Log in options will check for institutional or personal access. Content may require purchase if you do not have access.)

References

Abrahamyan, M.S., 1957, Brakhiopody verkhnefamenskikh i etrenskikh otlozheniy yugo-zapadnoy Armenii: Yerevan, Izdatelstvo Akademii Nauk Armyanskoy SSR, 142 p. [in Russian]Google Scholar
Abrahamyan, M.S., 1964, Karbon, in Mkrtchian, S.S., Vardaniants, L.A., Gabrielian, A.A, Maghakian, I.G., and Paffenholz, C.N., eds., Geologiya Armyanskoy SSR: Yerevan, Akademiya Nauk Armyanskoy SSR, v. 2, p. 96118. [in Russian]Google Scholar
Abrahamyan, M.S., 1974, Opisanie fauny, Devonskaya sistema, Tip Brachiopoda, Brakhiopody, in Akopian, V.T., ed., Atlas Iskopaemoy Fauny Armyanskoy SSR: Yerevan, Akademiya Nauk Armyanskoy SSR, Institut Geologicheskikh Nauk, p. 4867. [in Russian]Google Scholar
Abrahamyan, M.S., Arakelyan, R.A., and Azizbekov, Sh.A., 1973, Malyy Kavkaz (Yuzhnoe Zakavkaz'ya), in Rzhonsnitskaya, M.A., ed., Kavkazskaya Geosinklinal'naya Oblast i Predkavkaz'e, in Nalivkin, D.V., Rzhonsnitskaya, M.A., Markovski, B.P., eds., Stratigrafiya SSSR, Devonskaya Sistema: Moscow, Nedra, v. 1, p. 210219. [in Russian]Google Scholar
Abrahamyan, M.S., Arakelyan, R.A., and Papoyan, A.S., 1975, Granitsa Devona i Nizhnego Karbona, in Stratigrafiya Karbona i Geologiya Uglenosnykh Formatsii SSSR (Stratigraphy of the Carboniferous and Geology of Coal-Bearing Formations of the USSR): Moscow, Nedra, p. 4955. [in Russian]Google Scholar
Alekseeva, R.E., Afanasjeva, G.A., Grechishnikova, I.A., Oleneva, N.V., and Pakhnevich, A.V., 2018a, Devonian and Carboniferous brachiopods and biostratigraphy of Transcaucasia: Paleontological Journal, v. 52, p. 829967.CrossRefGoogle Scholar
Alekseeva, R.E., Afanasjeva, G.A., Grechishnikova, I.A., Oleneva, N.V., and Pakhnevich, A.V., 2018b, Devonian and Carboniferous brachiopods and biostratigraphy of Transcaucasia (ending): Paleontological Journal, v. 52, p. 9691085.CrossRefGoogle Scholar
Arakelyan, R.A., 1952, Stratigrafiya paleozoyskikh otlozheniy yugo-zapadnoy Armenii i prilegayushchikh chastey Nakhichevanskoy ASSR: Yerevan, Akademiya Nauk Armyanskoy SSR, Institut Geologicheskikh Nauk, 142 p. [in Russian]Google Scholar
Arakelyan, R.A., 1964, Devon, in Mkrtchian, S.S., Vardaniants, L.A., Gabrielian, A.A., Maghakian, I.G., and Paffenholz, C.N., eds., Geologiya Armyanskoy SSR: Yerevan, Akademiya Nauk Armyanskoy SSR, v. 2, p. 4696. [in Russian]Google Scholar
Aristov, V.A., 1994, Konodonty devona–nizhnego karbona Evrazii: soobshhestva, zonalnoye raschleneniye, korrelyatsiya raznofatsialnykh otlozheniy: Trudy Geologicheskogo Instituta Rossiyskoy Akademii Nauk v. 484, p. 1193. [in Russian]Google Scholar
Aristov, V.A., Grechishnikova, I.A., Tschigova, V.A., and Felix, V.P., 1979, Subdivision and correlation of the Famennian and the lower Tournaisian deposits of Transcaucasia (on brachiopods, conodonts and ostracods): Geological Survey of Belgium, Professional Paper, v. 161, p. 8795.Google Scholar
Brice, D., 1971, Étude paléontologique et stratigraphique du Dévonien de l'Afghanistan: Notes et Mémoires sur le Moyen-Orient, v. 11, p. 1364.Google Scholar
Brice, D., 1982, Eodmitria, genre nouveau de brachiopode Cyrtospiriferidae du Frasnien Inférieur et Moyen: Geobios, v. 15, p. 575581.CrossRefGoogle Scholar
Brice, D., Lafuste, J., de Lapparent, A.F., Pillet, J., and Yassini, I., 1973, Étude de deux gisements Paléozoïques (Silurien et Dévonien) de l'Elbourz Oriental (Iran): Annales de la Société Géologique du Nord, v. 93, p. 177218.Google Scholar
Brice, D., Jenny, J., Stampfli, G., and Bigey, F., 1978, Le Dévonien de l'Elbourz Oriental: stratigraphie, paléontologie (Brachiopodes et Bryozoaires), paléogéographie: Rivista Italiana di Paleontologia e Stratigrafia, v. 84, p. 156.Google Scholar
Brice, D., Mistiaen, B., and Rohart, J.-C., 1999, New data on distribution of brachiopods, rugose corals and stromatoporoids in the Upper Devonian of central and eastern Iran paleobiogeographic implications: Annales de la Société Géologique du Nord, v. 2, p. 2132.Google Scholar
Brock, G.A., and Yazdi, M., 2000, Palaeobiogeographic affinities of Late Devonian brachiopods from Iran: Records of the Western Australian Museum, Supplement 58, p. 321334.Google Scholar
Carlson, S.J., 2016, The evolution of Brachiopoda: Annual Review of Earth and Planetary Sciences, v. 44, p. 409438.CrossRefGoogle Scholar
Carter, J.L., and Gourvennec, R., 2006, Spiriferida. Introduction, in Kaesler, R.L., Treatise on Invertebrate Paleontology. Part H, Brachiopoda, 5 (revised): Boulder, Colorado and Lawrence, Kansas, Geological Society of America and University of Kansas Press, p. H1689H1694.Google Scholar
Carter, J.L., Johnson, J.G., Gourvennec, R., and Hou, H.F., 1994, A revised classification of the spiriferid brachiopods: Annals of the Carnegie Museum, v. 63, p. 327374.CrossRefGoogle Scholar
Denayer, J., and Hoşgör, I., 2014, Lower Carboniferous rugose corals from the Arabian Plate: an insight from the Hakkari area (SE Turkey): Journal of Asian Earth Sciences, v. 79, p. 345357.CrossRefGoogle Scholar
de Verneuil, E., 1845, Géologie de la Russie d'Europe et des Montagnes de l'Oural. Paléontologie: Paris, Bertrand, v. 2, p. 1512.Google Scholar
Frederiks, G.N., 1924, Paleontologicheskiye etyudy. 2. O Verkhne-Kamennougol'nykh spiriferidakh Urala: Izvestiya Geologicheskogo Komiteta v. 38, p. 295324. [in Russian]Google Scholar
Gaetani, M., 1965, The geology of the Upper Djadjerud and Lar valleys (North Iran). 2 Palaeontology. Brachiopods and molluscs from Geirud Formation, Member A (Upper Devonian and Tournaisian): Rivista Italiana di Paleontologia e Stratigrafia, v. 71, p. 679771.Google Scholar
Gaetani, M., 1968, Lower Carboniferous brachiopods from Central Elburz: Rivista Italiana di Paleontologia e Stratigrafia, v. 74, p. 665744.Google Scholar
Gatinaud, G., 1949, Contributions à l'étude des Brachiopodes Spiriferidae. I, Exposé d'une nouvelle méthode d'étude de la morphologie externe des Spiriferidae à sinus plissé: Bulletin du Museum National d'Histoire Naturelle, v. 21, p. 153–159, 300–307, 408–413, 487492.Google Scholar
Ginter, M., Hairapetian, V., and Grigorian, A, 2011, Chondrichthyan microfossils from the Famennian and Tournaisian of Armenia: Acta Geologica Polonica, v. 61, p. 153173.Google Scholar
Gourvennec, R., 2006, Upper Devonian brachiopods from Eastern Taurus (Turkey): Geologica Croatica, v. 59, p. 117.CrossRefGoogle Scholar
Gourvennec, R., 2019, Silurian–Devonian Spiriferida and Spiriferinida (brachiopods) from the Tindouf Basin (Algeria): Palaeontographica Abteilung, v. 313, p. 81149.CrossRefGoogle Scholar
Gourvennec, R., and Carter, J.L., 2007, Spiriferida and Spiriferinida, in Seldon, P.A., ed., Treatise on Invertebrate Paleontology. Part H, Brachiopoda 6 (revised): Lawrence, Kansas and Boulder, Colorado, Geological Society of America and University of Kansas Press, H2772H2796.Google Scholar
Grabau, A.W., 1923, Stratigraphy of China. Part 1. Palaeozoic and older : Geological Survey of China, Beijing, 200 p. (201528 [1924]).Google Scholar
Grabau, A.W., 1931, Devonian Brachiopoda of China, I: Devonian Brachiopoda from Yunnan and other districts in South China: China Geological Survey, Palaeontologia Sinica, Ser. B 3, p. 1545.Google Scholar
Grechishnikova, I.A., 1986, Novye Devonskiye spiriferidy Zakavkaz'ya: Zapiski Gornogo Instituta, v. 107, p. 5260. [in Russian]Google Scholar
Grechishnikova, I.A., 1996, New Givetian spiriferids (Brachiopoda) of Transcaucasia: Paleontologicheskiy Zhurnal, v. 4, p. 3437. [in Russian]Google Scholar
Grechishnikova, I.A., and Levitskiy, E.S., 2011, The Famennian–Lower Carboniferous reference section Geran-Kalasi (Nakhichevan Autonomous Region, Azerbaijan): Stratigraphy and Geological Correlation, v. 19, p. 2143.CrossRefGoogle Scholar
Grechishnikova, I.A., Levitskiy, V.S., and Feliks, V.P., 1980, K biostratigrafii srednego Devona Zakavkaz'ya: Byulleten’ Moskovskogo Obshchestva Ispytateley Prirody, Otdel Geologii, v. 55, p. 3950.Google Scholar
Grechishnikova, I.A., Aristov, V.A., Reitlinger, E.A., and Chizhova, V.A., 1982, Biostratigrafiya pogranichnykh otlozhenii devona i karbona Zakavkaz'ya (opornye razrezy): Severo-Vostochnyy Kompleksnyy Nauchno-Issledovatel'skiy Institut Dal'nevostochnogo Nauchnogo Tsentra Akademii Nauk SSSR, p. 138. [in Russian]Google Scholar
Grigoryan, A., Serobyan, V, Randon, C., Mayilyan, R., Avagyan, N., Danelian, T., 2019, A Famennian (Late Devonian) conodont assemblage from brachiopod-rich limestones of the Djravank section (southern Armenia): Proceedings of National Academy of Sciences of the Republic of Armenia, Earth Sciences, v. 72, p. 312.Google Scholar
Hall, J., 1883, Bryozoans of the Upper Heldelberg and Hamilton groups: Transactions of the Albany Institute, v. 10, p. 145197.Google Scholar
Ivanova, E.A., 1972, Osnovnyye zakonomernosti evolyutsii spiriferid (Brachiopoda): Paleontological Journal, v. 3, p. 309320.Google Scholar
Johnson, J.G., 2006, Cyrtospiriferoidea (part), in Kaesler, R.L., ed., Treatise on Invertebrate Paleontology. Part H, Brachiopoda 5 (revised): Boulder, Colorado and Lawrence, Kansas, Geological Society of America and University of Kansas Press, p. H1722H1732.Google Scholar
Johnson, J.G., Klapper, G., and Sandberg, C.A., 1985, Devonian eustatic fluctuations in Euramerica: Bulletin of the Geological Society of America, v. 96, p. 567587.2.0.CO;2>CrossRefGoogle Scholar
Ljashenko, A.I., 1957, Novyi rod Devonskikh brakhiopod Uchtospirifer: Doklady Akademii Nauk SSSR 117, p. 885–888. [in Russian]Google Scholar
Ljashenko, A.I., 1959, Atlas Brakhiopod i Stratigrafiya Devonskikh Otlozhenii Tsentral'nykh Oblastei Russkoi Platformy: Moskva, Gostoptehizdat, 451 p. [in Russian]Google Scholar
Ma, X.-P., and Day, J., 1999, The Late Devonian brachiopod Cyrtiopsis davidsoni Grabau 1923, and related forms from central Hunan of South China: Journal of Paleontology, v. 73, p. 608624.CrossRefGoogle Scholar
Ma, X.-P., and Day, J., 2000, Revision of Tenticospirifer Tien, 1938, and morphologically similar spiriferid brachiopod genera from the Late Devonian (Frasnian) of Eurasia, North America, and Australia: Journal of Paleontology, v. 74, p. 444463.CrossRefGoogle Scholar
Ma, X.-P., and Day, J., 2003, Revision of North American and selected Eurasian Late Devonian (Frasnian) species of Cyrtospirifer and Regelia (Brachiopoda): Journal of Paleontology, v. 77, p. 267292.2.0.CO;2>CrossRefGoogle Scholar
Ma, X.-P., and Day, J., 2007., Morphology and revision of Late Devonian (early Famennian) Cyrtospirifer (Brachiopoda) and related genera from South China and North America: Journal of Paleontology, v. 81, p. 286311.CrossRefGoogle Scholar
Ma, X.-P., Chen, X.Q., Day, J., and Jin., Y.G., 2003., Restudy of the Chinese Upper Devonian cyrtospiriferid (Brachiopoda): Sinospirifer Grabau, 1931: Acta Palaeontologica Sinica, 42, p. 367381.Google Scholar
Mamedov, A.B., and Rzhonsnitskaya, M.A., 1985, Devonian of the south Transcaucasus: zonal subdivision, boundaries of series and stages, correlation: Courier Forschungsinstitut Senckenberg, v. 75, p. 135156.Google Scholar
Martynova, M.V., 1961, Stratigrafiya i brakhiopody Famenskogo yarusa zapadnoy chasti Tsentral'nogo Kazakhstana: Materialy po Geologii Tsentral'nogo Kazakhstana, v. 2, p. 1211. [in Russian]Google Scholar
Mottequin, B., and Brice, D., 2019, Reappraisal of some Upper Devonian (Famennian) spiriferide brachiopods from the Band-e Bayan Domain (Afghanistan): Geobios, v. 52, p. 4765.CrossRefGoogle Scholar
Mottequin, B., Brice, D., and Legrand-Blain, M., 2014, Biostratigraphic significance of brachiopods near the Devonian-Carboniferous boundary: Geological Magazine, v. 151, p. 216228.CrossRefGoogle Scholar
Murchison, R.I., 1840, Description de quelques-unes des coquilles fossiles les plus abondantes dans les couches Dévoniennes du Bas-Boulonnais: Bulletin de la Société Géologique de France, v. 11, p. 250256.Google Scholar
Musabelliu, S., and Zatoń, M., 2018, Patterns of cornulitid encrustation on the Late Devonian brachiopod shells from Russia Proceedings of the Geologists’ Association, v. 129, p. 227234.Google Scholar
Nalivkin, D.V., 1937, Brakhiopody verkhnego i srednego devona i nizhnego karbona Severo-Vostochnogo Kazakhstana: Tsentral'nyi Nauchno-Issledovatel'skii Geologo-Razvedochnyi Institut (TSNIGRI), Trudy, v. 99, p. 1200. [in Russian]Google Scholar
Nalivkin, D.V., 1941, Brakhiopody Glavnogo devonskogo polya, in Borisiak, A.A., and Gekker, R.F., eds., Fauna Glavnogo Devonskogo Polya: Moska, Paleontologicheskiy Institut AN SSSR, v. 1, p. 139221. [in Russian]Google Scholar
Oleneva, N.V., 2016, Devonian brachiopods of the orders Spiriferida and Spiriferinida of the European Russia and Transcaucasia: systematics, shell microstructure: Paleontological Journal, v. 50, p. 12071296.CrossRefGoogle Scholar
Paeckelmann, W., 1942, Beiträge zur Kenntnis Devonischer Spiriferen: Abhandlungen des Reichsamts für Bodenforschung, v. 197, p. 1188.Google Scholar
Pushkin, V.I., 1996, Pripyatispirifer, a new Lower Famennian genus from Belarus: Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre, v. 66, p. 4351.Google Scholar
Reed, F.R.C., 1922, Devonian fossils from Chitral and the Pamirs: Memoirs of the Geological Survey of India: Palaeontologica Indica, n. ser., v. 6, p. 1134.Google Scholar
Richards, P.R., 1974, Ecology of the Cornulitidae: Journal of Paleontology, v. 48, p. 514523.Google Scholar
Rzhonsnitskaya, M.A., 1948, Devonskiye otlozheniya Zakavkaz'ya: Doklady Akademii Nauk SSSR, v. 59, p. 14771480. [in Russian]Google Scholar
Rzhonsnitskaya, M.A., and Mamedov, A.B., 2000, Devonian stage boundaries in the southern Transcaucasus: Courier Forschungsinstitut Senckenberg, v. 225, p. 329333.Google Scholar
Sartenaer, P., 1956a, Signification et importance du genre Cyrtiopsis dans les dépôts Famenniens inferieurs. Première note: Cyrtiopsis murchisoniana (de Verneuil, E., 1845): Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, v. 32, p. 119.Google Scholar
Sartenaer, P., 1956b, Signification et importance du genre Cyrtiopsis dans les dépôts Famenniens inferieurs. Deuxième note: Cyrtiopsis senceliae nov. sp.: Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, v. 32, p. 112.Google Scholar
Sartenaer, P., 1965, Signification et importance du genre Cyrtiopsis dans les dépôts Famenniens inférieurs. Quatrième note. Position systématique et stratigraphique du lectotype de l'espèce Spirifer murchisonianus de Verneuil, E., 1845: Bulletin de la Société Belge de Géologie, de Paléontologie et d'Hydrologie, v. 73, p. 366392.Google Scholar
Sartenaer, P., 1970, Nouveaux genres rhynchonellides (Brachiopodes) du Paléozoïque: Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, v. 46, p. 132.Google Scholar
Sartenaer, P., 1982, The presence and significance of Spirifer bisinus, S. malaisi, S. supradisjunctus and S. seminoi in early Frasnian beds of Western Europe, in Sartenaer, P., ed., Papers on the Frasnian/Givetian Boundary: Brussels, Geological Survey of Belgium, p. 122196.Google Scholar
Sartenaer, P., 1994, Tiocyrspis, a new late Frasnian cyrtiopsid genus from Germany and Belgium, in Königshof, P., and Werner, R. eds., Willi Ziegler-Festschrift II: Courier Forschungsinstitut Senckenberg, v. 169, p. 2943.Google Scholar
Schemm-Gregory, M., 2011, A new species of Cyrtospirifer (Brachiopoda) from the Middle Devonian of the Western Sahara (Northwest Africa): Rivista Italiana di Paleontologia e Stratigrafia, v. 117, p. 313.Google Scholar
Serobyan, V., Grigoryan, A., Mottequin, B., Mayilyan, R., Crônier, C., and Danelian, T., 2019, Biostratigraphy of the Upper Devonian trigonirhynchiid brachiopods (Rhynchonellida) from Armenia: Proceedings of National Academy of Sciences of the Republic of Armenia, Earth Sciences, v. 72, p. 318.Google Scholar
Serobyan, V., Danelian, T., Crônier, C., Grigoryan, A., and Mottequin, B., 2021, Lower Famennian (Upper Devonian) rhynchonellide and athyride brachiopods from the South Armenian Block: Journal of Paleontology, v. 95, p. 527552.CrossRefGoogle Scholar
Serobyan, V., Danelian, T., Crônier, C., Grigoryan, A., and Mottequin, B., 2022, Aramazdospirifer orbelianus (Abich, 1858) n. comb., a new cyrtospiriferid brachiopod genus and a biostratigraphically important species from the lower Famennian (Upper Devonian) of Armenia: Comptes Rendus Palevol, v. 21, p. 145156.Google Scholar
Sidjachenko, A.I., 1961, Verkhnedevonskiy podrod tsirtospiriferid Dmitria: Paleontologicheskii Zhurnal, v. 2, p. 8085.Google Scholar
Sidjachenko, A.I., 1962, Spiriferidy i Stratigrafiya Famenskikh Otlozheniy Tsentral'nogo i Yugo-vostochnogo Karatau: Moskva, Akademiya Nauk SSSR, Sibirskoe Otdelenie, 147 p. [in Russian]Google Scholar
Sokiran, E.V., 2006, Early–Middle Frasnian cyrtospiriferid brachiopods from the East European Platform: Acta Palaeontologica Polonica, v. 51, p. 759772.Google Scholar
Sosson, M., Rolland, Y., Müller, C., Danelian, T., Melkonyan, R., Kekelia, S., Adamia, S., Babazadeh, V., Kangarli, T., Avagyan, A., Galoyan, G., and Mosar, J., 2010, Subductions, obduction and collision in the Lesser Caucasus (Armenia, Azerbaijan, Georgia), new insights, in Sosson, M., Kaymakci, N., Stephenson, E.A., Bergerat, F., and Starostenko, V., eds., Sedimentary Basin Tectonics from the Black Sea and Caucasus to the Arabian Platform: London, Geological Society, Special Publication, v. 340, p. 329352.Google Scholar
Stampfli, G., Von Raumer, J.F., and Borel, G.D., 2002, Paleozoic evolution of pre-Variscan terranes: from Gondwana to the Variscan collision, in Martinez Catalan, J.R., Hatcher, R.D. Jr., Arenas, R., and Diaz Garcia, F., eds, Variscan-Appalachian Dynamics: The Building of the Late Paleozoic Basement: Geological Society of America, Special Paper, v. 364, p. 263280.Google Scholar
Tapanila, L., 2005, Palaeoecology and diversity of endosymbionts in Palaeozoic marine invertebrates: trace fossil evidence: Lethaia, v. 38, p. 8999.CrossRefGoogle Scholar
Taylor, P.D., and Wilson, M.A., 2008, Morphology and affinities of hederelloid “bryozoans”: Virginia Museum of Natural History, Special Publication, v. 15, p. 301309.Google Scholar
Termier, H., and Termier, G., 1949, Essai sur l’évolution des Spiriféridés: Notes et Mémoires, Service Géologique, Division des Mines et de la Géologie (Maroc), v. 74, p. 85112.Google Scholar
Vinn, O., 2010, Adaptive strategies in the evolution of encrusting tentaculitoid tubeworms: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 292, p. 211221.CrossRefGoogle Scholar
Vinn, O., Musabelliu, S., and Zaton, M., 2019, Cornulitids from the Upper Devonian of the Central Devonian Field, Russia: GFF, v. 141, p. 6876.CrossRefGoogle Scholar
von Schlotheim, E.F, 1820, Die Petrefakten-Kunde auf ihrem jetzigen Standpunkte durch die Beshreibung seiner Sammlung versteinerter und fossiler Ueberreste des their-und Planzenreichs der Voiwelt erlaeutert: Gotha, Becker, 437 p.Google Scholar
Waagen, W.H., 1883, Salt Range fossils, I. Productus-Limestone Fossils: Brachiopoda. Palaeontologia Indica, Series, 13–14, p. 391546.Google Scholar
Williams, A., Carlson, S.J., Brunton, C.H.C., Holmer, L.E., and Popov, L. 1996, A supra-ordinal classification of the Brachiopoda: Philosophical Transactions of the Royal Society of London, B, v. 351, p. 11711193.Google Scholar
Wilson, M.A., and Taylor, P.D., 2001, “Pseudobryozoans” and the problem of encruster diversity in the Paleozoic: PaleoBios, v. 21 (supplement to no. 2), p. 134135.Google Scholar
Ziegler, W., and Sandberg, C.A., 1990, The Late Devonian standard conodont zonation: Courier Forschungsinstitut Senckenberg, v. 121, p. 1115.Google Scholar