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Systematics of Shumardiidae (Trilobita), with new species from the Ordovician of Argentina
Published online by Cambridge University Press: 20 May 2016
Abstract
New shumardiid trilobites from Arenig to Early Caradoc deposits of the Argentine Precordillera include species of a revised Kweichowilla (K. salasae, K. sterrenae, and K. piojensis) and the new genus Changchowilla (C. sanjuanina and C. gracielae, and C.? carrerai). Conophrys fabiani new species occurs in the Early Tremadoc of the western Puna region, northwestern Argentina. Leoforteyia new genus is based on L. hintzei new species from the Ibexian of Utah. Published taxa recognized as new species are Conophrys rushtoni, Conophrys wrighti, Leoforteyia ludvigseni and Shumardia whittingtoni. Phylogenetic analysis suggests the differentiation of Shumardia Billings, 1862; Kweichowilla Chang, 1964; Changchowilla new genus; and Leoforteyia new genus as separate taxa and indicates that Conophrys Callaway, 1877, is a grade group. Silicified material provides a nearly complete growth series for Kweichowilla salasae, including the first protaspides known with confidence for the Shumardiidae. Additionally, ontogenetic material for K. sterrenae, C. sanjuanina, C. riojana (Benedetto and Cañas) and C.? carrerai is described.
Leoforteyia is restricted to the Ibexian-Whiterockian of western Laurentia; species of Shumardia are widespread in the Arenig to Early Ashgill of western Europe, Australia, Laurentia, China and Russia. Changchowilla is recorded in the mid-Late Arenig-Llanvirn of the Precordillera and in the Llanvirn-middle Caradoc of China. Argentine species of Kweichowilla are closely allied to Australian species, suggesting biogeographic affinities of the Precordillera to the eastern margin of Gondwana by the Llanvirn and Caradoc.
Functional morphology, occurrence in fine grained sediments, and association mainly with atheloptic and pelagic trilobites suggest that shumardiids were deposit-feeders that inhabited low light levels in low energy settings, with a limited ability to burrow in soft sediments.
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References
Albanesi, G. L., Ortega, G., and Hunicken, M. A.
1998. High resolution conodont-graptolite biostratigraphy controlling the diachronous boundary between the San Juan and Gualcamayo Formations (Arenig-Llanvirn) in the Argentine Precordillera. In
Bagnoli, G. (ed.), Abstracts Seventh International Conodont Symposium held in Europe, Bologna-Modena.Google Scholar
Astini, R. A.
1995. Geología e interpretación ambiental de la Formación Gualcamayo en su localidad clásica (suroeste de Guandacol y Cordón Perico-Potrerillo), Precordillera septentrional. Revista de la Asociación Geologica Argentina, 49:55–70.Google Scholar
Astini, R. A.
1998. E1 Conglomerado de Las Vacas y el Grupo Trapiche de la Precordillera: Tectónica distensiva en el Ordovícico Tardío. Revista de la Asociación Geológica Argentina, 53:489–503.Google Scholar
Astini, R. A., and Brussa, E. D.
1997. Dos localidades fosilíferas en el conglomerado de Las Vacas (Caradociano) en la Precordillera Argentina. Ameghiniana, 34:114.Google Scholar
Benedetto, J. L., and Herrera, Z. A.
1987. Primer hallazgo de braquiópodos y trilobites en la Formación Trapiche (Ordovícico tardío), Precordillera Argentina. X Congreso Geológico Argentino, San Miguel de Tucumán, 3:73–76.Google Scholar
Benedetto, J. L., Cañas, F. L., and Astini, R. A.
1986. Braquiópodos y trilobites de la zona de transitión entre las Formaciones San Juan y Gualcamayo en el área de Guandacol (La Rioja, Argentina). IV Congreso Argentino de Paleontología y Bioestratigrafía, Mendoza, 1:31–44.Google Scholar
Benedetto, J. L., Sánchez, T. M., Carrera, M. G., Brussa, E. D., and Salas, M. J.
1999. Paleontological constrains on successive positions of Precordillera terrane during the Early Paleozoic, p. 21–42. In
Ramos, V. A., and Keppie, D. (eds.), Laurentia-Gondwana connections before Pangea. Geological Society of America Special Paper, 336.Google Scholar
Burrett, C., Stait, B., and Laurie, J.
1983. Trilobites and microfossils from the Middle Ordovician of Surprise Bay, southern Tasmania, Australia. Memoirs of the Association of Australasian Palaeontologists, 1:177–193.Google Scholar
Callaway, C.
1877. On a new area of Upper Cambrian rocks in South Shropshire, with description of a new fauna. Quarterly Journal of the Geological Society of London, 33:652–672.CrossRefGoogle Scholar
Cañas, F. L.
1999. Facies and sequences of the Late Cambrian-Early Ordovician carbonates of the Argentine Precordillera: a stratigraphic comparison with Laurentian platforms, p. 43–62. In
Keppie, V. A. Ramos and D. (eds.), Laurentia-Gondwana connections before Pangea. Geological Society of America Special Paper, 336.Google Scholar
Chang, W. T., Chen, X., Xu, H.-K., and Chen, J.-Y.
1964.
In Atlas of Paleozoic fossils from North Guizhou, pl. 5–10 (no text). Nanjing. (In Chinese)Google Scholar
Chatterton, B. D. E., and Speyer, S. E.
1997. Trilobite ontogeny, p. 171–247. In
Whittington, H. B. (ed.), Treatise on Invertebrate Paleontology, Pt. O, Trilobita, Revised Edition. Geological Society of America and University of Kansas Press, Lawrence.Google Scholar
Chatterton, B. D. E., Edgecombe, G. D., Vaccari, N. E., and Waisfeld, B. G.
1997. Ontogeny and relationships of the Ordovician odontopleurid trilobite Ceratocara, with new species from Argentina and New York. Journal of Paleontology, 71:108–125.CrossRefGoogle Scholar
Chatterton, B. D. E., Edgecombe, G. D., Vaccari, N. E., and Waisfeld, B. G.
1999. Ontogenies of some Ordovician Telephinidae from Argentina, and larval patterns in the Proetida. Journal of Paleontology, 73:219–239.CrossRefGoogle Scholar
Chatterton, B. D. E., Edgecombe, G. D., Waisfeld, B. G., and Vaccari, N. E.
1998. Ontogeny and systematics of Toernquistiidae (Trilobita, Proetida) from the Ordovician of the Argentine Precordillera. Journal of Paleontology, 72:273–303.CrossRefGoogle Scholar
Clark, T. H.
1924. The paleontology of the Beekmantown series at Levis, Quebec. American Paleontologist Bulletin, 10:1–134.Google Scholar
Dean, W. T.
1973. Ordovician trilobites from the Keele Range, Northwestern Yukon Territory. Geological Survey of Canada Bulletin, 223:1–43.Google Scholar
Ebbestad, J. O. R.
1999. Trilobites of the Tremadoc Bjørkåsholmen Formation in the Oslo Region, Norway. Fossils and Strata, 47:1–118.Google Scholar
Edgecombe, G. D., Chatterton, B. D. E., Vaccari, N. E., and Waisfeld, B. G.
1997. Ontogeny of the proetoid trilobite Stenoblepharum Owens, 1973, and relationships of a new species from the Upper Ordovician of Argentina. Journal of Paleontology, 71:419–433.CrossRefGoogle Scholar
Edgecombe, G. D., Chatterton, B. D. E., Vaccari, N. E., and Waisfeld, B. G.
1999a. Ordovician cheirurid trilobites from the Argentine Precordillera. Journal of Paleontology, 73:1155–1175.CrossRefGoogle Scholar
Edgecombe, G. D., Chatterton, B. D. E., Waisfeld, B. G., and Vaccari, N. E.
1998. Ordovician (Whiterock) calymenid and encrinurid trilobites from the Precordillera of Argentina. Journal of Paleontology, 72:678–697.CrossRefGoogle Scholar
Edgecombe, G. D., Chatterton, B. D. E., Waisfeld, B. G., and Vaccari, N. E.
1999b. Ordovician pliomerid and prosopiscid trilobites from Argentina. Journal of Paleontology, 73:1144–1154.CrossRefGoogle Scholar
Farris, J. S.
1969. A successive approximations approach to character weighting. Systematic Zoology, 18:374–385.CrossRefGoogle Scholar
Fortey, R. A.
1980. The Ordovician trilobites of Spitsbergen, III: Remaining trilobites of the Valhallfonna Formation. Norsk Polarinstitut Skrifter, 171:1–163.Google Scholar
Fortey, R. A., and Droser, M. L.
1999. Trilobites from the base of the type Whiterockian (Middle Ordovician) in Nevada. Journal of Paleontology, 73:182–201.CrossRefGoogle Scholar
Fortey, R. A., and Owens, R. M.
1987. The Arenig Series in South Wales: stratigraphy and paleontology. Bulletin of the British Museum (Natural History) Geology, 41:69–307.Google Scholar
Fortey, R. A., and Owens, R. M.
1991. A trilobite fauna from the highest Shineton Shales in Shropshire, and the correlation of the latest Tremadoc. Geological Magazine, 128:437–464.CrossRefGoogle Scholar
Fortey, R. A., and Owens, R. M.
1999. Feeding habits in trilobites. Palaeontology, 42:429–466.CrossRefGoogle Scholar
Fortey, R. A., and Rushton, A. W. A.
1980.
Acanthopleurella Groom 1902: origin and life habits of a miniature trilobite. Bulletin of the British Museum (Natural History) Geology, 33:79–89.Google Scholar
Goloboff, P. A.
1993. Estimating character weights during tree search. Cladistics, 9:83–91.CrossRefGoogle Scholar
Groom, T. T.
1902. On a new trilobite from the Dictyonema-shales of the Malvern Hills. Geological Magazine, 9:70–73.CrossRefGoogle Scholar
Hammann, W., and Leone, F.
1997. Trilobites of the post-Sardic (Upper Ordovician) sequence of southern Sardinia. Pt. 1. Beringeria, 20:3–217.Google Scholar
Harrington, H. J.
1938. Sobre las faunas del Ordoviciano inferior del norte Argentino. Revista del Museo de La Plata Sectión de Paleontología, 4:109–289.Google Scholar
Harrington, H. J., and Leanza, A.
1957. Ordovician trilobites of Argentina. Department of Geology, University of Kansas, Special Publication, 276 p.Google Scholar
Kielan, Z.
1960. Upper Ordovician trilobites from Poland and some related forms from Bohemia and Scandinavia. Palaeontologica Polonica, 11:1–198.Google Scholar
Kobayashi, T.
1931. Studies on the stratigraphy and palaeontology of the Cambro-Ordovician faunas of Hualienchai and Niuhsintai, south Manchuria. Japanese Journal of Geology and Gography, 8:139–189.Google Scholar
Kobayashi, T.
1937. The Cambro-Ordovician shelly faunas of South America. Journal of the Faculty of Science of the University of Tokyo (section 2), 4:369–522.Google Scholar
Koroleva, M. N.
1964. New Middle Ordovician shumardiid trilobites from Northern Kazakhstan. Paleontologichesky Zhurnal, 1:71–75. (In Russian)Google Scholar
Koroleva, M. N.
1982. Ordovician trilobites of northeastern Kazakhstan. Ministerstvo geologii SSSR. Ministerstvo geologii Kazakhskoi SSSR. Kazakhsky nauchno—issledobatelsky institut (KazIMS), Moscow, p. 3–192. (In Russian)Google Scholar
Lake, P.
1906–1946. A monograph of the British Cambrian Trilobites. Monographs of the Paleontographical Society, 350 p.Google Scholar
Laurie, J. R., and Shergold, J. H.
1996. Early Ordovician trilobite taxonomy and biostratigraphy of the Emanuel Formation, Canning Basin, western Australia, pt. 1. Palaeontographica A, 240:56–103.Google Scholar
Legg, D. P.
1976. Ordovician trilobites and graptolites from the Canning Basin, western Australia. Geologica et Paleontologica, 10, 58 p.Google Scholar
Lu, Y.-H.
1975. Ordovician trilobite faunas of Central and Southwestern China. Palaeontologia Sinica, 152, 463 p.Google Scholar
Lu, Y.-H, and Chang, W.-T.
1974. Ordovician Trilobites, p. 124–136. In
A Handbook of Stratigraphy and Paleontology of South-west China. Science Press, Beijing. (In Chinese)Google Scholar
Lu, Y. H., Chu, C.-L., Chien, Y.-Y., Zhou, Z.-Y., Chen, J.-Y., Liu, G.-W., Yu, W., Chen, X., and Xu, H.-K.
1976. Ordovician biostratigraphy and paleozoogeography of China. Memoirs of the Nanjing Institute of Geology and Palaeontology, Academia Sinica, 7:1–83. (In Chinese)Google Scholar
Ludvigsen, R., and Westrop, S. R.
1986. Classification of the Late Cambrian trilobite Idiomesus Raymond. Canadian Journal of Earth Sciences, 23:300–307.CrossRefGoogle Scholar
Malanca, S.
1996. Morfología y ontogenia de un nuevo Shumardiidae (Trilobita) del Tremadociano de la Sierra de Mojotoro. Memorias del 12° Congreso Geológico de Bolivia, Tarija, 1:391–399.Google Scholar
Marek, L.
1964.
Shumardia Billings, 1862 a Staurocephalus Barrande, 1846 v českém Ordoviku. Časopis Národního Muzea, Praha, 133:153–154.Google Scholar
Moya, M. C., Malanca, S., Hongn, F. D., and Bahlburg, H.
1993. El Tremadoc temprano en la Puna occidental argentina. 12° Congreso Geológico Argentino y 2° Congreso de Exploración de Hidrocarburos, Mendoza, 2:20–30.Google Scholar
Ortega, G., and Brussa, E. D.
1990. La Subzona de Climacograptus bicornis (Caradociano temprano) en la Formatión Las Plantas en su localidad tipo, Precordillera de San Juan, Argentina. Ameghiniana, 27:281–288.Google Scholar
Pek, I., and Vaněk, J.
1989. Index of Bohemian trilobites. Krajské Vlastivědné Muzeum, Olomouc, 68 p.Google Scholar
Peng, S.-C.
1990a. Tremadoc stratigraphy and trilobite faunas of northwestern Hunan. 1: Trilobites from the Nantsinkwan Formation of the Yangtze Platform. Beringeria, 2:3–55.Google Scholar
Peng, S.-C.
1990b. Tremadoc stratigraphy and trilobite faunas of nrthwestern Hunan, 2: Trilobites from the Jiangnan Slope belt. Beringeria, 2:55–171.Google Scholar
Peng, S.-C.
1992. Upper Cambrian biostratigraphy and trilobite faunas of the Cili-Taoyuan area, northwestern Hunan, China. Memoirs of the Association of Australasian Palaeontologists, 13:1–119.Google Scholar
Prendini, L.
2000. Phylogeny and classification of the Superfamily Scorpionoidea Latreille 1802 (Chelicerata, Scorpiones): an exemplar approach. Cladistics, 16:1–78.CrossRefGoogle Scholar
PŘibyl, A., and Vanček, J.
1980a. Neue Erkenntnisse über einige Trilobiten aus dem bömischen Ordovizium. Časopis pro mineralogii a geologii, 25:263–274.Google Scholar
PŘibyl, A., and Vaněk, J.
1980b. Ordovician trilobites of Bolivia. Rozpravy Československé Akademie Věd. Řada Matematických a PŘírodrích Věd, 90:1–90.Google Scholar
Raymond, P. E.
1914. The succession of faunas at Lévis, P.Q.
American Journal of Science, 38:523–530.CrossRefGoogle Scholar
Reed, F. R. C.
1903. The Lower Palaeozoic trilobites of the Girvan district, Ayrshire, Pt. 1. Palaeontographical Society Monograph Publication 270, 57:1–48.Google Scholar
Robison, R. A., and Pantoja-Alor, J.
1968. Tremadocian trilobites from the Nochixtlán region, Oaxaca, Mexico. Journal of Paleontology, 42:767–800.Google Scholar
Ross, R. J.
1967. Some Middle Ordovician brachiopods and trilobites from the Basin Ranges, western United States. US Geological Survey Professional Paper, 523D:D1–D43.Google Scholar
Rushton, A. W. A.
1982. The biostratigraphy and correlation of the Marioneth-Tremadoc Series boundary in North Wales, p. 41–51. In
Dean, M. G., and Basset, W. T. (eds.), The Cambrian-Ordovician boundary: Sections, fossil distributions, and correlations. National Museum of Wales, Geological Series 3.Google Scholar
Sarmiento, G. N., and Rábano, I.
1992. Nuevas precisiones bioestratigráficas sobre la Formación Gualcamayo (Ordovícico inferior) en la Sierra de Villicum, San Juan, Argentina. Zentralblatt für Geologie und Paläontologie, 6:1785–1795.Google Scholar
Sars, M.
1835. Über einige neue oder unvollständig bekännte trilobiten. Isis, 28:333–343.Google Scholar
Sdzuy, K.
1955. Die fauna der Leimitz-Schiefer (Tremadoc). Abhandlungen von der Senckenbergischen Naturforschen-den Gesellschaft, 492:1–74.Google Scholar
Stubblefield, C. J.
1926. Notes on the development of a trilobite, Shumardia pusilla (Sars). Journal of the Linnean Society (Zoology), 36:345–379.Google Scholar
Stubblefield, C. J.
1927. Trilobita, p. 96–146. In
Stubblefield, C. J. and Bullman, O. M. B., The Shineton Shales of the Wrekin district: with notes on their development in other parts of the Shropshire and Herefordshire. Quarterly Journal of the Geological Society of London, 83.Google Scholar
Swofford, D. L.
1999. PAUP∗ 4.0. Phylogenetic analysis using parsimony (and other methods). Sinauer, Sunderland, MA.Google Scholar
Tripp, R. P., Zhou, Z.-Y., and Pan, Z.-Q.
1989. Trilobites from the Upper Ordovician Tangtou Formation, Jiangsu Province, China. Transactions of the Royal Society of Edinburgh, Earth Sciences, 80:25–68.CrossRefGoogle Scholar
Weir, J. A.
1959. Ashgillian trilobites from Co. Clare, Ireland. Palaeontology, 1:369–383.Google Scholar
Weller, S.
1910. Description of new species of Ordovician fossils from China. Proceedings of the United States National Museum, Washington, 32:557–563.CrossRefGoogle Scholar
Whittington, H. B.
1965. Trilobites of the Ordovician Table Head Formation, western Newfoundland. Bulletin of the Museum of Camparative Zoology, 132:277–442.Google Scholar
Wright, A. J., Cooper, R. A., and Simes, J. E.
1994. Cambrian and Ordovician faunas and stratigraphy, Mt Patriarch, New Zealand. New Zealand Journal of Geology and Geophysics, 37:437–476.CrossRefGoogle Scholar
Yin, G.-Z., and Li, S.-J.
1978. Trilobita, p. 385–595. In
Stratigraphical and Palaeontological Team of Guizhou (ed.), Palaeontological Atlas of Southwest China, Guizhou, volume 1, Cambrian-Devonian. Geological Publishing House, Beijing. (In Chinese)Google Scholar
Young, G. E.
1973. An Ordovician (Arenigian) trilobite faunule of great diversity from the Ibex area, western Utah. Brigham Young University Geology Studies, 20:91–115.Google Scholar
Zhang, W.-T.
1979. On the Miomera and Polymera (Trilobita). Scientia Sinica, 10:996–1004. (In Chinese)Google Scholar
Zhang, T.-R.
1981. Trilobita, p. 134–213. In
Geological Serving Team, and Institute of Geological Sciences, Xinjiang Bureau of Geology, and Exploration Department, Xinjiang Bureau of Petroleoum (eds.), Palaeontological Atlas of Northwest China, Xinjiang, Volume 1, Late Proterozoic-Early Palaeozoic. Geological Publishing House, Beijing. (In Chinese)Google Scholar
Zhou, T.-M., Liu, Y.-R., Meng, X.-S., and Sun, Z.-H.
1977. Trilobita, p. 104–266. In
Geological Institute of Hubei, Geological Bureaus of Henan, Hubei, Guandong and Guanxi (eds.), Palaeontological Atlas of South-Central China, volume 1, Palaeozoic. Geological Publishing House, Beijing. (In Chinese)Google Scholar
Zhou, Z.-Y., and Dean, W. T.
1986. Ordovician trilobites from Chedao, Gansu Province, northwest China. Palaeontology, 29:743–786.Google Scholar
Zhou, Z.-Y., and Zhang, J.-L.
1984. Uppermost Cambrian and lowest Ordovician trilobites of north and northeast China, p. 63–163. In
Nanjing Institute of Geology and Palaeontology, Academia Sinica (ed.), Stratigraphy and Palaeontology of Systemic Boundaries in China, Cambrian-Ordovician Boundary, 2.Google Scholar
Zhou, Z.-Y., Webby, B. D., and Yuan, W.-W.
1995. Ordovician trilobites from the Yingan Formation of northwestern Tarim, Xinjiang, northwestern China. Alcheringa, 19:47–72.CrossRefGoogle Scholar