Hostname: page-component-78c5997874-4rdpn Total loading time: 0 Render date: 2024-11-20T00:08:35.368Z Has data issue: false hasContentIssue false

Late Ordovician and Silurian lichid trilobites from northwestern Canada: Hemiarges, Borealarges, and Richterarges

Published online by Cambridge University Press:  14 July 2015

Margaret J. Campbell
Affiliation:
Department of Earth and Atmospheric Sciences, University of Alberta, Edmonton, T6G 2E3 Canada
Brian D. E. Chatterton
Affiliation:
Department of Earth and Atmospheric Sciences, University of Alberta, Edmonton, T6G 2E3 Canada

Abstract

Four new species of Borealarges, B. fritillus, B. patulus, B. renodis, and B. variabilis, and one new species of Richterarges, R. facetus, are described and one unnamed species, Borealarges sp., discussed. All are from the Wenlock strata of Avalanche Lake sections in the Mackenzie Mountains, Northwest Territories. Borealarges tuckerae Adrain, 1994, the only species reported from both the Arctic and the Mackenzie Mountains, is discussed. Hemiarges avalanchensis n. sp., an Ashgill species from Avalanche Lake section AV 4B just below the Ordovician-Silurian boundary, is described. A phylogenetic analysis based on 14 species of Borealarges, three of Richterarges, and two of Hemiarges, demonstrates that the former two genera are three separate and distinct taxa. Borealarges, a genus that includes some species formerly assigned to Richterarges or Hemiarges, is monophyletic, contains a well-supported internal clade of species, and is not separated into senso stricto and senso lato groupings.

Type
Research Article
Copyright
Copyright © The Paleontological Society 

Access options

Get access to the full version of this content by using one of the access options below. (Log in options will check for institutional or personal access. Content may require purchase if you do not have access.)

References

Adrain, J. M. 1994. The lichid trilobite Borealarges n. gen., with species from the Silurian of Arctic Canada. Journal of Paleontology, 68:10811099.CrossRefGoogle Scholar
Adrain, J. M. 2003. Validity and comparison of the Silurian trilobite genera Borealarges and Dicranogmus with new species from the Canadian Arctic. Canadian Journal of Earth Sciences, 40:749763.CrossRefGoogle Scholar
Campbell, M. J. and Chatterton, B. D. E. 2006. Late Ordovician and Silurian lichid trilobites of northwestern Canada. Eight new species from subfamilies Lichinae, Platylichinae, and Trochurinae. Journal of Paleontology, 80(3):514528.CrossRefGoogle Scholar
Chatterton, B. D. E. and Ludvigsen, R. 1976. Silicified Middle Ordovician trilobites from the South Nahanni River Area, District of Mackenzie, Canada. Palaeontographica A, 154:1106.Google Scholar
Chatterton, B. D. E. and Perry, D. G. 1984. Silurian cheirurid trilobites from the Mackenzie Mountains, northwestern Canada. Paleontographica A, 184:178.Google Scholar
Chatterton, B. D. E., Edgecombe, G. D., and Tuffnell, P. A. 1990. Extinction and migration of trilobites and conodonts of northwestern Canada. Journal of the Geological Society, London, 147:703715.CrossRefGoogle Scholar
Clarke, J. M. 1894. The Lower Silurian trilobites of Minnesota. Final report of the Geological and Natural History Survey of Minnesota, III:695759.Google Scholar
Fortey, R. A. 2000. Trilobite! Eyewitness to Evolution. HarperCollins, London. 269 p.CrossRefGoogle Scholar
Gürich, G. 1901. Über eine neue Lichas-Art aus dem Devon von neu-Süd-Wales und über die Gattung Lichas überhaupt. Neues Jahrbuch für Mineralogie, Geologie und Palaeontologie, 14:519539.Google Scholar
Hall, J. and Clarke, J. M. 1888. Descriptions of the trilobites and other Crustacea of the Oriskany, Upper Helderberg, Hamilton, Portage, Chemung and Catskill groups. New York Geological Survey, Palaeontology, New York, 7, 236 p.Google Scholar
Hawle, I. and Corda, A. J. C. 1847. Prodrom einer Monographie der Bömischen Trilobiten. Prague, 176 p.Google Scholar
Hunda, B. R., Chatterton, B. D. E., and Ludvigsen, R. 2003. Silicified Late Ordovician trilobites from the Mackenzie Mountains, Northwest Territories, Canada. Palaeontographica Canadiana, No. 21, 87 p.Google Scholar
Kaesler, R. L. (ed.). 1997. Treatise on Invertebrate Paleontology, Pt. O, Arthopoda 1. Trilobita, (revised), 1: Introduction, Order Agnostida, Order Redlichiida. Geological Society of America and University of Kansas, Lawrence, 530 p.Google Scholar
Ludvigsen, R. 1979. Fossils of Ontario, Pt. 1, The Trilobites. Royal Ontario Museum, Toronto, 96 p.Google Scholar
Maddison, W. P. and Maddison, D. R. 1992. MacClade: Analysis of Phylogeny and Character Evolution. Version 3.0. Sinauer Associates, Sunderland, Massachusetts, 398 p.Google Scholar
Milne Edwards, H. 1840. Histoire Naturelle des Crustacés, Comprenant l'Anatomie, la Physiologie et la Classification de ces Animaux. Volume 3. Paris, 638 p.Google Scholar
Nowlan, G. S., Mccracken, A. D., and Chatterton, B. D. E. 1988. Conodonts from Ordovician-Silurian Boundary strata, Whittaker Formation, Mackenzie Mountains, Northwest Territories. Geological Survey of Canada Bulletin, 373:199, 22 pl.Google Scholar
Over, D. J. and Chatterton, B. D. E. 1987. Silurian conodonts from the southern Mackenzie Mountains, Northwest Territories, Canada. Geologica et Palaeontologica, 21:149.Google Scholar
Perry, D. G. and Chatterton, B. D. E. 1977. Silurian (Wenlockian) trilobites from Baillie-Hamilton Island, Canadian Arctic Archipelago. Canadian Journal of Earth Sciences, 14:285319.CrossRefGoogle Scholar
Phleger, F. Jr. 1936. Lichadian trilobites. Journal of Paleontology, 10:593615.Google Scholar
Rudkin, D. M., Tripp, R. P., and Ludvigsen, R. 1994. The Ordovician trilobite Hemiarges (Lichidae: Trochurinae) from North America and Greenland, p. 289306. In Landing, E. (ed.), Studies in stratigraphy and paleontology in honor of Donald W. Fisher. New York State Museum Bulletin, No. 481.Google Scholar
Schmidt, F. 1885. Revision der Ostbaltischen Silurischen Trilobiten. Abtheilung 2. Acidaspiden und Lichiden. Mémoires de l'Académie Impériale des Sciences de St-Pétersbourg, 7th series, 33:1127.Google Scholar
Swofford, D. L. 1993. PAUP: Phylogenetic Analysis Using Parsimony, Version 3.1.1. Illinois Natural History Survey, 1989. Champaign.Google Scholar
Thomas, A. T. and Holloway, D. J. 1988. Classification and phylogeny of the trilobite order Lichida. Philosophical Transactions of the Royal Society of London, 321:179262.Google Scholar
Tripp, R. P. 1957. The classification and evolution of the superfamily Lichacea (Trilobita). Geological Magazine, 94:104122.CrossRefGoogle Scholar
Tripp, R. P. 1959. Lichidae, p. 495503. In Moore, R. C. (ed.), Treatise on Invertebrate Paleontology. Volume O (1) (Arthropoda). Geological Society and University of Kansas Press, Lawrence.Google Scholar
Tripp, R. P. and Evitt, W. R. 1981. Silicified Lichidae (Trilobita) from the Middle Ordovician of Virginia. Geological Magazine, 118:665677.CrossRefGoogle Scholar
Wang, K., Chatterton, B. D. E., Attrep, J. M., and Orth, C. J. 1993. Late Ordovician mass extinction in the Selwyn Basin, northwestern Canada: Geochemical, sedimentological and paleontological evidence. Canadian Journal of Earth Sciences, 30:18701880.CrossRefGoogle Scholar
Whittington, H. B. 1961. Silurian Hemiarges (Trilobita) from Cornwallis Island and New York State. Journal of Paleontology, 33:433444.Google Scholar
Whittington, H. B. 2002. Lichidae (Trilobita): Morphology and classification. Journal of Paleontology, 72:306320.2.0.CO;2>CrossRefGoogle Scholar
Whittington, H. B. and Kelly, S. R. A. 1997. Morphological terms applied to Trilobita, p. 313329. In Kaesler, R. L. (ed.), Treatise on Invertebrate Paleontology, Pt. O, Arthopoda 1. Trilobita (revised). Volume 1. Introduction, Order Agnostida, Order Redlichiida. Geological Society of America and University of Kansas, Lawrence.Google Scholar