To clarify the role of the incisors in selective grazing by cattle and horses, experiments were conducted in July 1999, using three Holstein cattle and three thoroughbred horses to investigate bite weight and bite force using hand-constructed swards with 15 loadcells spaced 6×12 cm apart. Leaves of perennial ryegrass (Lolium perenne: PRG) were mixed with leaves of tall fescue (Festuca arundinacea: TF) or reproductive culms of reed canarygrass (Phalaris arundinacea: RCG).

There was little difference between cattle and horses in bite rate, but DM intake rate of horses was almost twice that of cattle in the PRG/TF mixture. In the PRG/RCG mixture, both animals tended to graze only leaves of PRG. Total bite force was 3270–3280 N/min in cattle and 5350–6350 N/min in horses in the PRG/TF mixture. Means of maximum values of one bite force (not total force per bite) were 88±5·5 N in cattle and 141±11·0 N in horses. These values corresponded to 1·6±0·10% of body weight in cattle and 2·2±0·16% in horses, after converting Newton force into kilogram force. Bite area was higher in horses than in cattle. Both cattle and horses did not discriminate between leaves of PRG and TF. In the PRG/RCG mixture, horses prehended both leaves and culms in 33% of total clumps grazed, compared with only 5% in cattle. Many culms escaped from the mouth at the moment of prehension. In mixtures of soft leaves and stiff stems, the lack of upper incisors in cattle may have the advantage over horses. The term ‘comb-out strategy’ is proposed for this selective foraging by cattle. DM intake per bite force was highest in TF leaves (7·5 mg DM/N) grazed by horses and similar (3·7–4·6 mg DM/N) in PRG leaves grazed by both cattle and horses.