A high-resolution carbon isotope profile through the uppermost Neoproterozoic–Lower Cambrian part of the Sukharikha section at the northwestern margin of the Siberian platform shows prominent secular oscillations of δ13C with peak-to-peak range of 6–10 ‰. There are six minima, 1n–6n, and seven maxima 1p–7p, in the Sukharikha Formation and a rising trend of δ13C from the minimum 1n of − 8.6 ‰ to maximum 6p of + 6.4 ‰. The trough 1n probably coincides with the isotopic minimum at the Precambrian–Cambrian boundary worldwide. Highly positive δ13C values of peaks 5p and 6p are typical of the upper portion of the Precambrian–Cambrian transitional beds just beneath the Tommotian Stage in Siberia. A second rising trend of δ13C is observed through the Krasnoporog and lower Shumny formations. It consists of four excursions with four major maxima that can be correlated with Tommotian–Botomian peaks II, IV, V, and VII of the reference profile from the southeastern Siberian platform. According to the chemostratigraphic correlation, the first appearances of the index forms of archaeocyaths are earlier in the Sukharikha section than in the Lena–Aldan region.

The Hengshan–Wutai–Fuping belt is located in the middle segment of the Trans-North China Orogen, a Palaeoproterozoic continental collisional belt along which the Eastern and Western blocks amalgamated to form the North China Craton. The belt consists of the medium- to high-grade Hengshan and Fuping gneiss complexes and the intervening low- to medium-grade Wutai granite–greenstone terrane, and most igneous rocks in the belt are calc-alkaline and have affinities to magmatic arcs. Previous tectonic models assumed that the Hengshan and Fuping gneiss assemblages were an older basement to the Wutai supracrustal rocks, but recent studies indicate that the three complexes constitute a single, long-lived Neoarchaean to Palaeoproterozoic magmatic arc where the Wutai Complex represents an upper crustal domain, whereas the Hengshan and Fuping gneisses represent the lower crustal components forming the root of the arc. The earliest arc-related magmatism in the belt occurred at 2560–2520 Ma, marked by the emplacement of the Wutai granitoids, which was followed by arc volcanism at 2530–2515 Ma, forming the Wutai greenstones. Extension driven by widespread arc volcanism led to the development of a back-arc basin or a marginal sea, which divided the belt into the Hengshan–Wutai island arc (Japan-type) and the Fuping relict arc. At 2520–2480 Ma, subduction beneath the Hengshan–Wutai island arc caused partial melting of the lower crust to form the Hengshan tonalitic–trondhjemitic–granodioritic (TTG) suites, whereas eastward-directed subduction of the marginal sea led to the reactivation of the Fuping relict arc, where the Fuping tonalitic–trondhjemitic–granodioritic suite was emplaced. In the period 2360–2000 Ma, sporadic phases of isolated granitoid magmatism occurred in the Hengshan–Wutai–Fuping region, forming 2360 Ma, c. 2250 Ma and 2000–2100 Ma granitoids in the Hengshan Complex, the c. 2100 Ma Wangjiahui and Dawaliang granites in the Wutai Complex, and the 2100–2000 Ma Nanying granitoids in the Fuping Complex. At c. 1920 Ma, the Hengshan–Wutai island arc underwent an extensional event, possibly due to the subduction of an oceanic ridge, leading to the emplacement of pre-tectonic gabbroic dykes that were subsequently metamorphosed, together with their host rocks, to form medium- to high-pressure granulites. At 1880–1820 Ma, the Hengshan–Wutai–Fuping arc system was juxtaposed, intensely deformed and metamorphosed during a major and regionally extensive orogenic event, the Lüliang Orogeny, which generated the Trans-North China Orogen through collision of the Eastern and Western blocks. The Hengshan–Wutai–Fuping belt was finally stabilized after emplacement of a mafic dyke swarm at 1780–1750 Ma.

Detailed remapping of the Palaeogene Ardnamurchan Centre 3 gabbros, NW Scotland, suggests that this classic sequence of ring-intrusions forms a composite layered lopolith. The area mapped by previous studies as the Great Eucrite gabbro intrusion comprises 70% by area of Centre 3. Field observations suggest that most of the other smaller ring-intrusions of Centre 3 (interior to the Great Eucrite) constitute either distinct petrological facies of the same intrusion, or included country-rock or peridotite blocks. These observations, together with syn-magmatically deformed inward-dipping modal layering, are used here to support the interpretation that significant central sagging occurred in the intrusion at a late stage in its crystallization history.

The lower Williamson Shale at Tryon Park, Rochester, New York State is unique in the co-occurrence of Stimulograptus clintonensis (Hall), previously recorded only from the middle Telychian, and Pterospathodus amorphognathoides amorphognathoides Walliser, previously recorded only from the upper Telychian (Cyrtograptus lapworthi Biozone and higher). These two co-occurring fossils are clearly providing contradictory ages for the lower Williamson Shale. The incorporation of chitinozoan data from both the Williamson and Willowvale shales does not support unequivocally either the graptolite- or conodont-derived biostratigraphy and thus the age of the lower Williamson Shale remains problematical.

Vegetation diversity and pattern changes, and their relation to tectono-sedimentary histories are compared between selected Euramerican Late Palaeozoic coalfields, to understand better the controls on the dynamics of the Pennsylvanian terrestrial ecosystems and to demonstrate the problems with comparing data from various basins. The analysis is based on data from the following basins of different geotectonic and palaeogeographical positions: the cratonic Pennines Basin, the foreland South Wales and Upper Silesia basins, and the fault-related Intra Sudetic and Central and Western Bohemia basins. The analysis indicates that complex factors are responsible for changes in plant diversity and vegetation patterns. These are related to climate, tectonics, preservation potential, sampling biases and the current state of revision of the flora in each basin. Plant diversity patterns in the basins differ because of local controls and/or the character and detail of the available data. Maximum diversity varies among the basins within the Langsettian and Duckmantian substages. Two apparent step-like drops in diversity were detected within coal-bearing strata of most basins: at the Duckmantian/Bolsovian boundary and at the Bolsovian/Asturian boundary. Further and more prominent falls are related to transitions from coal-bearing to non-coal-bearing (mostly red bed) strata or vice versa during Stephanian times. Interpretation of climatic signals recorded in the sedimentary successions indicates that Westphalian and middle Stephanian times were wet intervals, whereas early and late Stephanian times were drier.

Upper Triassic volcanic and sedimentary rocks of the Mamonia Complex in southwestern Cyprus are exposed in erosional windows through the post-Cretaceous cover, where the Mamonia Complex is tectonically imbricated with the Troodos and Akamas ophiolitic suites. Most of these Upper Triassic volcanic rocks have been considered to represent remnants of Triassic oceanic crust and its associated seamounts. New Nd and Pb isotopic data show that the whole Mamonia volcanic suite exhibits features of oceanic island basalts (OIB). Four rock types have been distinguished on the basis of the petrology and chemistry of the rocks. Volcanism began with the eruption of depleted olivine tholeiites (Type 1) and oceanic island tholeiites (Type 2) associated with deep basin siliceous and/or calcareous sediments. The tholeiites were followed by highly phyric alkali basalts (Type 3) interbedded with pelagic Halobia-bearing limestones or white reefal limestones. Strongly LREE-enriched trachytes (Type 4) were emplaced during the final stage of volcanic activity. Nd and Pb isotopic ratios suggest that tholeiites and mildly alkali basalts derived from partial melting of heterogeneous enriched mantle sources. Fractional crystallization alone cannot account for the derivation of trachytes from alkaline basalts. The trachytes could have been derived from the partial melting at depth of mafic material which shares with the alkali basalts similar trace element and isotopic compositions. This is corroborated by the rather similar isotopic compositions of the alkali basalts and trachytes. The correlations observed between incompatible elements (Nb, Th) and εNd and Pb isotopic initial ratios suggest that the Mamonia suite was derived from the mixing of a depleted mantle (DMM) and an enriched component of High μ (μ = 238U/204Pb, HIMU) type. Models using both Nd and Pb isotopic initial ratios suggest that the depleted tholeiites (Type 1) derived from a DMM source contaminated by an Enriched Mantle Type 2 component (EM2), and that the oceanic tholeiites (Type 2), alkali basalts (Type 3) and trachytes (Type 4) were derived from the mixing of the enriched mantle source of the depleted tholeiites with a HIMU component. None of the Mamonia volcanic rocks show evidence of crustal contamination. The Upper Triassic within-plate volcanism likely erupted in a small southerly Neotethyan basin, located north of the Eratosthenes seamount and likely floored by oceanic crust.

The Blattodea (cockroaches sensu lato or roachoids) from the southern coalfields of the UK are reviewed or examined, especially species occurring in more than one coal basin including Mylacris abrupta (Bolton, 1930) comb. nov., and undescribed material from Writhlington (Somerset). It is proposed that a derived mylacrid Mylacris–Sooblatta assemblage is used to extend the latest Westphalian and earliest Stephanian insect zonation from the continent to southern Britain, thus augmenting the European Pennsylvanian scheme.

Ion microprobe (SIMS) dating of zircon from the Miranda do Douro orthogneiss, Central Iberian Zone, Hercynian Iberian belt, defines an Early Ordovician U–Pb age of 496.0 ± 2.6 Ma (95 % conf., MSWD = 1.14) for magmatic zircon crystallization in its granitic protolith. The age contrasts with an earlier, conventional (ID-TIMS) U–Pb zircon age of 618 ± 9(95 % conf.) Ma, now thought to be an artefact of the complex zircon population. Individual SIMS ages for zircon from the rock range from 2700 to 180 Ma and comprise inherited and magmatic zircon, both concordant and common Pb-enriched, and younger, reset ages. The ID-TIMS study seems to have misinterpreted this heterogeneous population as a cogenetic suite consisting of magmatic zircon and its age-reset equivalents produced by recent Pb loss. The 496 ± 3 Ma SIMS age represents the weighted average for 26 magmatic zircon domains located by careful inspection of cathodoluminescence, secondary electron and optical microscopy images of ∼ 700 individual zircon crystals. Inherited zircon is widespread, ranging in age from 2700 to 550 Ma, with age clusters, which are statistically indistinguishable from those known from pre-Hercynian granitic basement material elsewhere in the Central Iberian Zone. Including the present age information, 582 ± 4 Ma (95 % conf., MSWD = 1.02, n = 13) and 619 ± 9 Ma (95 % conf., MSWD = 0.93, n = 7) appear as prevailing inherited zircon age components in basement intrusions in the Central Iberian Zone.

Two suites of granitoids, the Late Jurassic (158 ± 3 Ma) Linglong suite and the Early Cretaceous (130–126 Ma) Guojialing suite, crop out in the northwestern Jiaodong Peninsula, eastern China. The Linglong suite is a monzogranite, comprising alkali feldspar, plagioclase, quartz and Fe-rich biotite. The Guojialing suite includes at least five plutonic bodies of both granodiorite and monzo-granite. The rocks are composed of plagioclase, alkali feldspar, quartz, Mg-rich amphibole and Mg-rich biotite. Both the Linglong and Guojialing suites have adakitic affinity. They are enriched in LREE with high La/Yb ratios and show positive Eu anomalies. The rocks are also enriched in LILE and depleted in HFSE with high Sr/Y ratios. The Linglong granite shows very uniform Sr–Nd isotopic compositions with initial 87Sr/86Sr ratios of 0.7119–0.7126 and εNd (T) values of −21.3 to −21.6, which are similar to those of the local Neoarchaean basement. The Guojialing suite has variable initial 87Sr/86Sr ratios (0.7108–0.7120) and εNd (T) values (−10.8 to −17.2), which are distinct both from those of the Neoarchaean basement and from those of the local enriched lithospheric mantle inferred from the coeval mafic dykes in the studied area. Detailed petrological and geochemical data indicate that the Linglong suite was derived by partial melting of Neoarchaean metamorphic lower-crustal rocks at depth of > 50 km with a eclogite residue, whereas the Guojialing suite was formed by the reaction of delaminated eclogitic crust-derived melt with the upwelling asthenospheric mantle. The petrogenesis of these two contrasting adakitic granitoids suggests intensive lower-crustal delamination during Early Cretaceous times, following a crustal thickening process from the late stage of the Early Jurassic to early stage of the Late Jurassic with crustal thickness of < 32 km to > 50 km, respectively.

Biostratigraphical ranges and palaeogeographical distribution of mid-Givetian to end-Frasnian odontopleurids are investigated. The discovery of Leonaspis rhenohercynica sp. nov. in mid-Givetian strata extends this genus unexpectedly up to the late Middle Devonian. New material of Radiaspis radiata (Goldfuss, 1843) and the first koneprusiine in Britain, Koneprusia? sp., are described from the famous Lummaton shell-bed, Torquay, Devon. New taxa of Koneprusia, K. serrensis, K. aboussalamae, K. brevispina, and K. sp. A and K. sp. B are defined. Ceratocephala (Leonaspis) harborti Richter & Richter, 1926, is revised and reassigned to Gondwanaspis Feist, 2002. Two new species of Gondwanaspis, G. dracula and G. spinosa, plus three others left in open nomenclature, are described from the late Frasnian of Western Australia. A further species of Gondwanaspis, G. prisca, is described from the early Frasnian of Montagne Noire. Species of Gondwanaspis are shown to possess a number of paedomorphic features. A functional analysis suggests that, unlike other odontopleurids, Gondwanaspis actively fed and rested with the same cephalic orientation. The sole odontopleurid survivors of the severe terminal mid-Givetian biocrisis (‘Taghanic Event’) belong to the koneprusiine Koneprusia in the late Givetian and Frasnian, and, of cryptogenic origin, the acidaspidine Gondwanaspis in the Frasnian. Whereas the former became extinct in the late Frasnian at the Lower Kellwasser Event, the latter disappeared, and with it the entire Odontopleuroidea, at the terminal Frasnian Upper Kellwasser global biocrisis.

Western Turkey is an area which has experienced large-scale extension of continental crust. Here we report precise crystallization ages of two intrusions in the central Menderes Massif, the Turgutlu and Salihli granodiorites, using U–Pb dating. Both intrusions occur in the southern footwall of the seismically active Alaşehir graben and were emplaced syntectonically in an extensional top-to-the-NNE shear zone which was active at retrograde greenschist-facies conditions. The U–Pb ages of 16.1 ± 0.2 Ma (monazite, Turgutlu granodiorite) and 15.0 ± 0.3 Ma (allanite, Salihli granodiorite) document that tectonic exhumation of middle-crustal rocks in the central Menderes Massif was already underway at the Early to Middle Miocene transition. Combined with published geochronological, structural and sedimentological data, the new U–Pb ages point to a continued extension since at least 16 Ma. There is no convincing evidence for a late Miocene/Pliocene phase of tectonic shortening.

The Tiñu Formation of Oaxaca State is the only fossiliferous lower Palaeozoic unit between the Laurentian platform in northwest Mexico and Gondwanan successions in Andean South America. The Tiñu traditionally has been referred to the Lower Ordovician (Tremadoc) and regarded as having a provincially mixed fauna with Laurentian, Avalonian, and Gondwanan elements. Bio- and lithostratigraphic re-evaluation demonstrates that the Tiñu is a Gondwanan, passive margin succession. It includes a lower, thin (to 16 m), condensed, uppermost Cambrian Yudachica Member (new). The Yudachica nonconformably overlies middle Proterozoic basement as a result of very high late Late Cambrian eustatic levels. The Yudachica changes from storm-dominated, but slightly dysoxic, shelf facies (fossil hash limestone and shale) in the south to an upper slope facies with debris flows 50 km to the north. Three biostratigraphically distinct depositional sequences comprise the Yudachica. The Yudachica has Gondwanan-aspect trilobites with low-diversity conodonts characteristic of unrestricted marine/temperate facies. The upper Tiñu, or Río Salinas Member (new), is a Lower Ordovician (Tremadoc) depositional sequence that records strong early, but not earliest, Tremadoc eustatic rise marked by graptolite- and olenid-bearing dysoxic mudstones. Higher strata shoal upward into shell-hash limestones and proximal tempestite sandstones with upper lower Tremadocian unrestricted marine/temperate conodonts. New taxa include Orminskia rexroadae Landing gen. et sp. nov. from the Cordylodus andresi Zone; this euconodont is related to hyaline coniform genera best known from Ordovician tropical platform successions. Cornuodus? clarkei Landing sp. nov. resembles the coeval, upper lower Tremadoc tropical species Scalpellodus longipinnatus (Ji & Barnes).

Brian Harland was for many years an editor of this journal. He was also a seminal figure in the origins of the current ‘snowball Earth’ debate, having recognized in 1964 the significance of coupling emerging palaeomagnetic data on palaeolatitude with his interpretations of diamictites. Harland worked extensively in the Arctic and knew well many of the workers involved in the arguments surrounding the origin of diamictites. He thus had a unique perspective on the evidence and the disputes surrounding it. This was his last paper but he was not able to complete it before he died. However, with the help of Professor Ian Fairchild to whom we are indebted, the editors have lightly revised this work which is presented as the personal view of one of the key figures with a very broad stratigraphic appreciation of the problems of ‘snowball Earth’.

Records of Precambrian glaciation onwards from the late nineteenth century led to the concept of one or more major ice ages. This concept was becoming well advanced by the mid 1930s, particularly through the compilation of Kulling in 1934. Even so tillite stratigraphy shows that glaciation was exceptional rather than typical of Earth history. Some Proterozoic tillites, sandwiched between warm marine facies, indicate low, even equatorial palaeolatitudes as determined magnetically, and more recently led to ideas of a snow- and ice-covered ‘snowball Earth’. However, interbedded non-glacial facies as well as thick tillite successions requiring abundant snowfall both militate against the hypothesis of extreme prolonged freezing temperatures referred to here as an ‘iceball Earth’ in which all oceans and seas were sealed in continuous ice cover. On the other hand tropical environments were interrupted by glaciation several times in the Proterozoic, something that did not recur in the Phanerozoic. The term ‘snowball Earth’ is consistent with the established view of extremely widespread Proterozoic glaciation, but the ‘iceball Earth’ version of this is not compatible with the geological record.

Organic-walled palynomorph assemblages from the Kasterlee Formation in northern Belgium provide new insights into the Late Miocene depositional history and palaeoenvironments of the southernmost North Sea Basin. Ranges of key dinoflagellate cysts constrain the unit between 7.5 and 5.32 Ma, that is, a latest Tortonian to Messinian age. The palynomorph assemblage is characterized, amongst others, by Geonettia clineae, an opportunistic species that thrives in mesotrophic, coastal embayments with a low sediment influx. This environmental setting is corroborated by the notable presence of Gramocysta verricula, a species with preference for shallow marine environments. The occurrence of species of the fresh water green alga Pediastrum indicates manifest river discharge in a near-shore environment or embayment. The coastal depositional environment mirrored by the palynomorphs of the Kasterlee Formation succeeds the distinct transgressive and fully marine environments of the underlying Diest Formation in the Campine area. The results from the palynological study, combined with lithological and geophysical data, show that both Upper Miocene formations are two distinct depositional cycles separated by an erosional or regressive phase. The upper boundary of the Kasterlee Formation is correlated with the Me2 sequence boundary at 5.73 Ma. The Kasterlee Formation is herein formally moved from the Lower Pliocene series to the Upper Miocene series. The coastal environment, probably characterized by a shoaling phase, recorded at the border of the southern North Sea Basin, matches the global record of regressive phases in Messinian sedimentary sequences, which are linked to cooling and increasing global ice volume.

Re-description of the left dentary of Chinshakiangosaurus chunghoensis reveals that it possesses an unusual combination of ‘prosauropod’ and ‘sauropod’ character states. Cladistic analysis places Chinshakiangosaurus as one of the most basal sauropods known currently. Mapping of dentary and dental characters onto the most parsimonious topologies yields insights into the sequence of acquisition of a number of feeding-related characters. For example, it seems that basal sauropodomorphs (traditional prosauropod taxa) possessed a fleshy cheek that attached to the mandible along a marked ridge, and that the same structure was present in the most basal sauropods. The early sauropod skull developed a lateral plate that reinforced the bases of the tooth crowns labially, and had wrinkled tooth enamel and a concavity on the mesial portion of the lingual part of each crown, while retaining a fleshy cheek and a relatively weak symphysis. More advanced sauropods (eusauropods) lost the cheek, perhaps in order to increase the gape of the jaws in response to a change in feeding style that involved collection of larger quantities of poor quality foliage.

The Troodos ophiolite in Cyprus and Baer–Bassit ophiolite in Syria together form part of the Tethyan ophiolite belt. They were generated in a supra-subduction zone setting in Late Cretaceous times. As with many of the ophiolite occurrences in this belt, the sequences are closely associated with tectonic ‘coloured mélange’ zones, which contain, among a variety of lithologies, metre- to kilometre-size blocks of metamorphic rocks. Precise 40Ar–39Ar laser step-heating experiments performed on four amphibolites from SW Cyprus and six from NW Syria, yield plateau ages ranging from 75.7±0.3 Ma to 88.9±0.8 Ma in Cyprus and 71.7±0.5 to 88.4±0.4 Ma in Syria. The older limits of these time spans are coeval with the age of the formation of the associated ophiolites. Unlike other metamorphic sole rocks which seem to form in relatively short time spans, these metamorphic rocks found in Cyprus and Syria are interpreted to have formed in Late Cretaceous times by accretion below the overriding Troodos and Baer–Bassit crust for a period of 15–18 Ma. The metamorphic complexes were exhumed by extension and crustal thinning associated with subduction roll-back and the rotation of the overriding plate until the cessation of subduction in Maastrichtian times. In Cyprus, the exhumed metamorphic complex was incorporated into an accretionary prism constructed primarily of the collapsed Mamonia passive margin sequence intercalated with rocks of the Troodos ophiolite during plate collision in the Maastrichtian. Concomitantly, in Syria, the Baer–Bassit ophiolite and subcreted metamorphic complex were emplaced onto the Arabian passive margin and fragmented into blocks and knockers, forming the Baer–Bassit mélange.

A summary is given of the lycophyte species as distributed in fifteen Euramerican coalfields during Asturian (Westphalian D) times. The datasets from the coalfields are analysed using unweighted pair group cluster analyses of Jaccard's Coefficients. Separate analyses were done on the full dataset (the full list of morphospecies) and on a more restricted dataset (only the morphospecies of vegetative organs). Three groupings of coalfields were resolved in the analyses. The two most clearly defined groupings represented the intramontane basin coalfields, and the coalfields of the lowland basins on the Variscan foreland where the lycophyte floras are well documented. The third grouping included those basins with less well-studied lycophyte floras, where the apparently low species diversity has undoubtedly affected the results. More work is needed on these poorly documented floras to produce a more rigorous phytogeographical analysis.

Cretaceous mafic dykes in Fujian province, SE China provide an opportunity to examine the nature of their mantle source and the secular evolution of the Mesozoic lithospheric mantle beneath SE China. The mafic rocks have SiO2 ranging from 47.42 to 55.40 wt %, Al2O3 from 14.0 wt % to 20.4 wt %, CaO from 4.09 to 11.7 wt % and total alkaline (K2O+Na2O) from 2.15 wt % to 6.59 wt %. Two types are recognized based on their REE and primitive mantle-normalized trace element patterns. Type-A is the dominant Mesozoic mafic rock type in SE China and is characterized by enrichment of light rare earth elements (LREE) ((La/Yb)n = 2.85–19.0) and arc-like trace element geochemistry. Type-P has relatively flat REE patterns ((La/Yb)n = 1.68–3.43) and primitive mantle-like trace element patterns except for enrichment of Rb, Ba and Pb. Type-A samples show EMII signatures on the Sr-Nd isotopic diagram, whereas type-P rocks have high initial 143Nd/144Nd ratios (0.5126–0.5128) relative to the type-A rocks (143Nd/144Nd = 0.5124–0.5127). The type-A rocks have 207Pb/204Pb ranging from 15.47 to 15.67 and 206Pb/204Pb from 18.26 to 18.52. All the type-A rocks show a negative correlation between 143Nd/144Nd and 206Pb/204Pb ratios and a positive relationship between 87Sr/86Sr and 206Pb/204Pb ratios, indicating mixing of a depleted mantle source and an EMII component. Geochemical modelling shows that the parental magmas were formed by 5–15 % partial melting of a spinel lherzolite, and contaminated by less than 1 % melt derived from subducted sediment. The type-P magmas were derived from a mantle source unmodified by subduction components. The wide distribution of type-A dykes in SE China suggests that subduction-modified lithospheric mantle was extensive beneath the Cathaysia Block. Geochemical differences between Mesozoic and Cenozoic mafic rocks indicate that lithospheric thinning beneath SE China occurred in two episodes: firstly by heterogeneous modification by subducted components in early Mesozoic times, and later by chemical–mechanical erosion related to convective upwelling of the asthenosphere during Cenozoic times.

The South Wales Coalfield has the most complete Westphalian macrofloral record anywhere on the Variscan Foreland or adjacent basins, with 135 biodiversity-meaningful morphospecies having been recognized. All of the standard macrofloral biozones of the Westphalian Stage have been recognized, although a detailed comparison with the Central Pennines Coalfields has indicated some discrepancies in the relative positions of the biozonal boundaries. Total Species Richness progressively increases through the Langsettian Substage, and then remains relatively stable through most of the Duckmantian and Bolsovian substages. There is a distinct reduction in Total Species Richness towards the top of the Bolsovian Substage, but this partially recovers in the middle Asturian Substage with the appearance of a range of marattialean ferns, and medullosalean and callistophytaleans pteridosperms. There is no evidence of any significant drop in Total Species Richness towards the top of the succession, indicating that conditions at this time were relatively stable. The change from coastal floodplain to alluvial braidplain conditions in middle Bolsovian times correlates with a marked increase in the proportion of medullosalean remains being preserved in the adpression record, reflecting an expansion of the clastic-substrate habitats.

New carbon isotopic data from the Devonian of Ardennes (Belgium) and partly from the Holy Cross Mountains (Poland) highlight an abrupt and high-amplitude negative excursion in the punctata conodont Zone. Published information from Moravia and China suggests that this Middle Frasnian negative excursion, jointly with the preceding large-scale positive shift, should be used as a global chemostratigraphic marker. Causation scenarios for this negative ‘punctata Event’ are correlated neither with major biota turnover nor major sea-level changes, but may be related to: (1) the Alamo Impact Event, that led to (2) the massive dissociation of methane hydrates and (3) the rapid onset of global warming.