The Cover-plates of the radial grooves are preserved here and there in specimen, notably over the oral centre (Pl. X, Figs. 1, 2, 5, 7, 8), but in B they are preserved over the whole of the grooves and the mouth, though pressed down on to the floor of the grooves and into the peristome (Pl. XI, Fig. 1). Cover-plates are also present in the British Museum specimens, and are almost complete in E 16054.

Each cover-plate corresponds in position with a floor-plate; so that, like the floor-plates, the cover-plates form a double series of alternating plates meeting in a zigzag median suture. The almost straight or slightly rounded outer margin of each cover-plate fits into a bevelled facet just within the rounded edge of the radial groove where the floor-plate begins its downward slope (Pl. X, Figs. 5, 7). This feature is well shown in specimen B, and there the floor-plate also shows, on each, side of the facet, and distinct from the peripodium, a slight depression, possibly for the insertion of a muscle or ligament, or possibly for the reception of the accessory cover-plate shortly to be described.