Incomplete, but structurally preserved remains of an organism referable to the Nematophytales Lang, are described from the Rhynie Chert as a new genus and species. The fossil is composed of a plexus of non-septate tubular cells of two types—the smaller having uniformly smooth walls, the larger, walls with conspicuous spiral thickenings. Branching of both types is localized in areas referred to as branch-knots. Comparison is made with other members of the Nematophytales which have been shown to possess differentially thickened tubes.