The Duntulm Formation of the Bathonian (Middle Jurassic) Great Estuarine Group represents a marine-brackish lagoonal complex which was transgressively established on the drowned delta of the underlying Valtos Sandstone Formation. Duntulm Formation sediments accumulated in adjacent basins, the Sea of the Hebrides-and Inner Hebrides-basins. Litho-, bio-, and palynofacies analysis of these rocks show that the most distinctive lithologies, monotypic shell banks of the oyster Praeexogyra hebridica (Lithofacies 1), accumulated under variable, but distinctly marine conditions. Argillaceous carbonate muds (Lithofacies 2), probably accumulated in the lee of the oyster banks under variably marine-freshwater conditions, while the supralittoral lagoon shores were fringed by algal marshes (Lithofacies 3). The palynology of Lithofacies 3 shows that some marshes accumulated close to the lagoon (dominated by marine dinocysts), while others formed further inland (dominated by terrestrial pollen and spores). In the N of the Sea of the Hebrides Basin, small deltas continued to prograde into the lagoons (Lithofacies 4), and toward the end of Duntulm Formation times, muds and sands colonised by freshwater molluscs (Unio and Neomiodon), the planktonic alga Botryococcus, and dominated by terrestrial pollen and spores (Lithofacies 5), accumulated in this area, representing a basin-wide change from marine to freshwater conditions. The structural high which divided the basins exerted a strong control on lithofacies evolution, effectively preventing any material coarser than silt grade entering the Inner Hebrides Basin. Thickness variation in the Formation is similarly structurally/facies linked, with thick sequences accumulating where sandstones (Lithofacies 4) predominate, and thin sequences occurring close to the structural high.

In the W of Ireland the Ordovician rocks of South Mayo and Clew Bay are now juxtaposed but a comparison of the sedimentary histories of these two sequences shows that they accumulated in basins which were probably separated during most of their history. The large amount of terrigenous detritus present in the Arenig to Llanvirn elements of the South Mayo succession is not manifest in that of Clew Bay until the Llandeilo/Caradoc, by which time sedimentation in South Mayo had ceased. A comparison of the South Mayo Ordovician with that of Girvan in Scotland demonstrates that both sequences had a similar provenance. This source contained an ophiolite, granites and some (probably pre-Dalradian) metamorphic rocks. Sediment dispersal directions for the two sequences are opposite in sense, being primarily northward in South Mayo and southward at Girvan. The two stratigraphies indicate that basement subsidence behaviour in South Mayo was virtually the opposite of that at Girvan where initial shallow water sedimentation was rapidly succeeded by deep water environments at the end of the Llanvirn. The two basins may thus have been marginal to a single Ordovician arc complex. One reason for the opposite sense of basin subsidence may lie in the suggested reversal of subduction polarity during the Ordovician. In this scenario the South Mayo basin may be envisaged as lying to the N of a northward-facing arc during the early Ordovician. A new, northward, subduction direction instigated during the Llanvirn, resulted in a fore-arc basin at Girvan complemented by a closing back-arc basin in South Mayo.

Our knowledge of the primitive and aberrant early tetrapod Crassigyrinus scoticus was based on a partial skull roof and mandibles from the Lower Carboniferous of Gilmerton, Edinburgh, plus a skeleton lacking the hind limb and tail from the Namurian, basal Upper Carboniferous, of Cowdenbeath, Fife. New specimens from Cowdenbeath include the pelvic girdle, presacral and sacral rib and most of the hind limb. The ilium of the girdle had a firm articulation with the vertebral column via the sacral rib. The ischium, separated from the ilium by cartilage, was, like that found with the Cowdenbeath skeleton, ornamented as though a dermal bone. No pubis is preserved. The femur lacks an adductor crest, but has a strongly developed internal trochanter. Tibia and fibula are short stout bones, but axial torsion is present in the fibula rather than the tibia. This, and the structure of the femur, suggests a swimming rather than a walking limb. Probable metatarsals and phalanges are recorded. A skeletal reconstruction and a life restoration of Crassigyrinus are presented in the light of its reconstructed anatomy and physiology.

The ornamentation of the ischium of Crassigyrinus, and that of the colosteid Greererpeton, suggests that the bone may be at least in part dermal. Its homology wth the pelvic fin basal scute of osteolepiform fishes is proposed and the homologies of pectoral and pelvic fins, and thus limbs, discussed.

The monotypic ostracode Entomozoe (pro Entomis) (Superorder Myodocopa) from the Silurian of Scotland, type genus of the Entomozoidae, is revised for the first time since being established in 1861. Based on the type and new collections of Entomozoe tuberosa (Jones, 1861), the genus shows affinity to bolbozoid myodocopes rather than to forms hitherto universally considered as ‘typical’ entomozoid myodocopes. The families Entomozoidae and Bolbozoidae are considered synonymous. In the interests of stability of nomenclature, in particular to preserve the traditionally understood usage of the taxon Entomozoidae, a case must be made to the International Commission on Zoological Nomenclature to set aside any previous designations and to establish a new type-species for Entomozoe.

Entomozoe, from the Llandovery Series, is the earliest known myodocope in the fossil record. It occurs as part of a low diversity, shallow water, nearshore assemblage. In contrast to other, but younger, Silurian bolbozoid myodocopes, it is probably a benthic form. The differences in the anterior shell morphology of Silurian bolbozoid taxa possibly reflect changes both in soft-part morphology and from benthic to pelagic lifestyles within the group.

Both bolbozoid and ‘entomozoid’ myodocopes are assigned to the Order Myodocopida rather than to the Order Halocypridida, though the taxonomic affinity of the ‘entomozoids’ is, at best, uncertain.

Detailed mapping of the Trallval-Ruinsival area of the ultrabasic complex of Rhum has revealed the relationship between the intrusive peridotites, which form a major part of the complex, and the layered ultrabasic rocks. The layered rocks on Trallval are correlated with Units 8 to 15 of the Eastern Layered Series as seen on Hallival and Askival, and part of an additional unit, Unit 16, has also been identified. The layered rocks of Ruinsival and the western part of Trallval form part of the Central Series and have been subdivided into six cyclic units. They are associated with four sets of intrusive breccia zones which converge towards a central region. Four cylindrical plugs of intrusive peridotite occur within the area of outcrop of the Central Series. Criteria are listed for the distinction of intrusive peridotite from conformable peridotites formed in situ. Distinctive textures and structures are described; these include vertical feldspathic streamers, which are interpreted as direct evidence for the upward expulsion of interstitial fluid, and loading structures at the contact between a peridotite layer and underlying allivalite are ascribed to the intrusion of a peridotite sill. A comparison of the Eastern Layered Series and the Central Series suggests that three stages were involved in the formation of the ultrabasic rocks: (i) the accretion of rhythmically layered unconsolidated olivine cumulates; (ii) the upward expulsion of intercumulus liquid; and (iii) the formation of either allivalite (as in the Eastern Layered Series) or ultrabasic breccia zones (as in the Central Series) from the expelled intercumulus liquid, depending on whether the tectonic environment was quiescent or extensional.