Two new species of petrified seeds Eosperma edromense, and Anasperma burnense gen. et sp. nov. are described from the Cementstone Group of Berwickshire. Anasperma shows that the anatropous condition existed among Palæozoic ovules; it has a single integument with two lateral apical lobes.
Rachides of Stauropteris berwickensis sp. nov. and associated megasporangia are also described.
An account is given of the theory of the telomic origin of the first ovular integument. The second (outer) integument in Angiosperm ovules may therefore have evolved after the establishment of anatropy, either from the first integument or as an overgrowth of the chalaza. Evidence is cited to support the theory that Angiosperm carpels have evolved from dorsiventral bivalved cupules.