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XV.—The Placoderm Fish Rhachiosteus pterygiatus Gross and its Relationships*

Published online by Cambridge University Press:  06 July 2012

Roger S. Miles
Affiliation:
Department of Geology, Royal Scottish Museum, Edinburgh.

Synopsis

The holotype and only known specimen of Rhachiosteus pterygiatus Gross is partially redescribed and new restorations are given. Attention is drawn to important points in its osteology and the possible development of a cutaneous sensory system. A definition of the family Rhachiosteidsæ Stensiö is given. This family differs from all other described groups of euarthrodires in the lack of posterior lateral and posterior dorsolateral flank plates. Rhachiosteus is a pachyosteomorph brachythoracid, as defined in the text, and may be fairly closely related in some way to the (coccosteomorph) family Coccosteidsæ. There is no indication that it is closely related to any other known pachyosteomorph, or to other groups of arthrodires, such as the Rhenanida and Ptyctodontida, in which there are no posterior flank plates.

Type
Research Article
Copyright
Copyright © Royal Society of Edinburgh 1966

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References

References to Literature

Boeke, J., 1934. “Freie Nervenendigungen und Endorgane sensibler Nerven”, in Handbuch der vergleichenden Anatomie, 2, Urbam and Schuarzenberg, Berlin-Wien.Google Scholar
Case, E. C., 1931. “Arthrodiran remains from the Devonian of Michigan”, Contr. Mus. Geol. Univ. Mich., 3, 163182.Google Scholar
Denison, R., 1958. “Early Devonian Fishes from Utah. 3. Arthrodira”, Fieldiana, Geol., 11, 461551.Google Scholar
Dunkle, D. H., and Bungart, P. A., 1946. “The Antero-supragnathal of Gorgonichthys”, Amer. Mus. Nov., 1316, 10 pp.Google Scholar
Edinger, T., 1956. “The Paired Pineal Organs”, in Progress in Neurobiology. Elsevier Publ. Co.Google Scholar
Gross, W., 1932. “Die Arthrodira Wildungens”, Geol. Paläont. Abh., 19, 61 pp.Google Scholar
Gross, W., 1933. “Die Wirbeltiere des rheinischen Devons”, Abh. Preuss, Geol. Landesanst., 154, 584.Google Scholar
Gross, W., 1938 a. “Rhachiosteus pterygiatus n. gen., n. sp. (Euarthrodira, Brachythoraci)”, Decheniana, 97A, 183208.Google Scholar
Gross, W., 1938 b. “Über das Spinale und die angrenzenden Knochen der Brachythoraci”, Neues. Jb. Min. Geol. Paläont., 79B, 403418.Google Scholar
Gross, W., 1956. “Über Crossopterygier und Dipnoer aus dem baltischen Oberdevon im Zusammenhang einer vergleichenden Untersuchung des Porenkanalsystems paläozoischer Agnathen und Fische”, K. Svenska VetenskAkad. Handl. 5, 140 pp.Google Scholar
Gross, W., 1960. “Tityosteus n. gen., ein Riesenarthrodire aus dem rheinischen Unterdevon”, Paläont. Z., 34, 263274.CrossRefGoogle Scholar
Gross, W., 1963. “Gemuendina stuertzi Traquair. Neuuntersuchung”, Notizbl. Hess. Landesamt. Bodenforsch., 91, 3673.Google Scholar
Gross, W., 1964. “Über die Randzähne des Mundes, die Ethmoidalregion des Schädels und die Unterkiefersymphyse von Dipterus oervigi n. sp.”, Paläont. Z., 38, 725.CrossRefGoogle Scholar
Heintz, A., 1930. “Eine neue Rekonstruktion von Heterostius ASM”, S.B. Naturf. Ges. Jurjew., 38, 7 pp.Google Scholar
Heintz, A., 1931. “Revision of the Structure of Coccosteus decipiens Ag.”, Norsk Geol. Tidsskr., 12, 291313.Google Scholar
Heintz, A., 1932. “The Structure of Dinichthys. A contribution to our knowledge of the Arthrodira”, Amer. Mus. Nat. Hist., Bashford Dean Mem. Vol., 4, 115224.Google Scholar
Heintz, A., 1933. “Revision of the Estonian Arthrodira. Pt. 1. Family Homostiidsæ Jaekel”, Arch. Naturk. Eestis, 10, 177290.Google Scholar
Jaekel, O., 1929. “Die Morphogenie der ältesten Wirbeltiere”, Monogr. Geol. Palwont., 3, 198 pp.Google Scholar
Jarvik, E., 1948. “On the Morphology and Taxonomy of the Middle Devonian Osteolepid Fishes of Scotland”, K. Svenska VetenskAkad. Handl., 25, 301 pp.Google Scholar
Jux, U., 1960. “Montecaris lehmani, a New Crustacean from the Rhenish Devonian and the Problem of its Systematic Position”, J. Palœont., 34, 11291152.Google Scholar
Jux, U., 1964. “Zur stratigraphischen Gliederung des Devonprofils von Bergisch Gladbach (Rheinisches Schiefergebirge)”, Decheniana, 117, 159174.Google Scholar
Kulczycki, J., 1957. “Upper Devonian Fishes from the Holy Cross Mountains (Poland)”, Acta Palcœont. Polon., 2, 285380.Google Scholar
Lekander, B., 1949. “The Sensory Line System and the Canal Bones in the Head of Some Ostariophysi”, Acta Zool, 30, 131 pp.CrossRefGoogle Scholar
Luther, A., 1913. “Beiträge zur Kenntnis des Kopfskelets der Knorpelganoiden”, Acta Soc. Sci. Fenn., 41, 26 pp.Google Scholar
Mark, E. J., 1953. “Genus Holonema from the Middle Devonian of Estonian S.S.R.”, Loodusuurijate Seltsi Juubelikoguteos, 18531953, Tallinn, 1953, 382396. (In Russian).Google Scholar
Mark, E. J., 1963. ”On the Spinal Plate of the Middle Devonian Arthrodire Homostius”, Geoloogia-Inst. Uurim., 13, 189199. (In Russian with English and Estonian summaries).Google Scholar
Miles, R. S., 1962. “Gemuendenaspis n.gen., an Arthrodiran Fish from the Lower Devonian Hunsrückschiefer of Germany”, Trans. Roy. Soc. Edin., 65, 5977.Google Scholar
Miles, R. S., 1964. “On Some Coccosteomorph Arthrodires from the Devonian of Arizona”, Ark. Zool., 16, 427460.Google Scholar
Miles, R. S., 1965. “Ventral Thoracic Neuromast Lines of Placoderm Fishes”, Nature, Lond., 206, 524525.CrossRefGoogle Scholar
Miles, R. S., 1966. “Protitanichthys and Some Other Coccosteomorph Arthrodires from the Devonian of North America”, K. Svenska VetenskAkad. Handl. [In the press, to be published as 10, no. 3.]Google Scholar
Miles, R. S., and Westoll, T. S., 1963. “Two New Genera of Coccosteid Arthrodira from the Middle Old Red Sandstone of Scotland and their Stratigraphical Distribution”, Trans. Roy. Soc. Edin., 65, 179210.CrossRefGoogle Scholar
Nielsen, E., 1949. “Studies on Triassic Fishes from East Greenland. 2. Australosomus and Birgeria”, Medd. Grønland, 146, 309 pp.Google Scholar
Obruchev, D. V., 1932. “Holonemidœ des russischen Devons”, Trav. Inst. Paléozool. Acad. Sci. U.R.S.S., 2, 97115.Google Scholar
Obruchev, D. V., 1964. “Class Placodermi”, in Foundations of Palœntology. Reference book for palœntology and geology of the USSR. Agnaiha and Fishes. Moscow.Google Scholar
Ørvig, T., 1960. “New Finds of Acanthodians, Arthrodires, Crossopterygians, Ganoids and Dipnoans in the Upper Middle Devonian Calcareous Flags (Oberer Plattenkalk) of the Bergisch Gladbach-Paffrath Trough. Pt. 1”, Paläont. Z., 34, 295335.Google Scholar
Ørvig, T., 1961. “New Finds of Acanthodians, Arthrodires, Crossopterygians, Ganoids and Dipnoans in the Upper Middle Devonian Calcareous Flags (Oberer Plattenkalk) of the Bergisch Gladbach-Paffrath Trough. Pt. 2”, Paläont. Z., 35, 1027.CrossRefGoogle Scholar
Ørvig, T., 1962. “Y a-t-il une rélation directe entre les arthrodires ptyctodontides et les holocéphales?Colloq. Int. Cent. Nat. Rech. Sci., 104.Google Scholar
Simpson, G. G., 1961. Principles of Animal Taxonomy. xii + 247 pp. Oxford University Press.CrossRefGoogle Scholar
Stensiö, E. A., 1934. “On the Heads of Certain Arthrodires. 1. Pholidosteus, Leiosteus and Acanthaspids”, K. Svenska VetenskAkad. Handl., 13, 79 pp.Google Scholar
Stensiö, E. A., 1944. “Contributions to the Knowledge of the Vertebrate Fauna of the Silurian and Devonian of Western Podolia. 2. Notes on Two Arthrodires from the Downtonian of Podolia”, Ark. Zool., 35A, 83 pp.Google Scholar
Stensiö, E. A., 1948. “On the Placodermi of the Upper Devonian of East Greenland. 2. Antiarchi: subfamily Bothriolepinæ with an Attempt at a Revision of the previously described species of that Family”, Palœozool. Grønland., 2, 622 pp.Google Scholar
Stensiö, E. A., 1959. “On the Pectoral Fin and Shoulder Girdle of the Arthrodires”, K. Svenska. VetenskAkad. Handl., 8, 229 pp.Google Scholar
Stensiö, E. A., 1963. “Anatomical Studies on the Arthrodiran Head. Pt. 1. Preface, Geological and Geographical Distribution, the Organisation of the Arthrodires, the Anatomy of the Head in the Dolichothoraci, Coccosteomorphi and Pachyosteomorphi. Taxonomic appendix”, K. Svenska VetenskAkad. Handl., 9, 419 pp.Google Scholar
Watson, D. M. S., 1938. “On Rhamphodopsis, a Ptyctodont from the Middle Old Red Sandstone of Scotland”, Trans. Roy. Soc. Edin., 59, 397410.CrossRefGoogle Scholar
Westoll, T. S., 1945. “The Paired Fins of Placoderms”, Trans. Roy. Soc. Edin., 61, 381398.CrossRefGoogle Scholar
Westoll, T. S., 1958. “The Lateral Fin-Fold Theory and th? Pectoral Fins of Ostracoderms and Early Fishes”, in Studies on Fossil Vertebrates. The University of London: The Athlone Press.Google Scholar
Westoll, T. S., and Miles, R. S., 1963. “On an Arctolepid Fish from Gemünden”, Trans. Roy. Soc. Edin., 65, 139153.CrossRefGoogle Scholar
White, E. I., 1952. “Australian Arthrodires”, Bull. Brit. Mus. (Nat. Hist.) Geol, 1, 249304.Google Scholar
White, E. I., 1962. “A Dipnoan from the Assise de Mazy of Hingeon”, Bull. Inst. Roy. Sci. Nat. Belg., 38, 8 pp.Google Scholar