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XIII.—A New Cephalaspid Fish from the Downtonian of Scotland, with Notes on the Structure and Classification of Ostracoderms

Published online by Cambridge University Press:  06 July 2012

T. S. Westoll
Affiliation:
Department of Geology and Mineralogy, University of Aberdeen.

Extract

Fossils have been known to occur in the early “post-Caledonian” sediments of Cowie Harbour, Stonehaven, for more than sixty years. Dr R. Campbell (1911, 1912 a, b, 1913) gave an account of the stratigraphy and recorded a number of fossils, including plant-fragments, arthropods (subsequently discussed by Steamer, 1934, 1935), and fishes (Traquair, 1912; Kiaer and Heintz, 1932 a; Heintz, 1939). The fossinferous “Dictyocaris-band,” exposed on the tidal foreshore, has been visited repeatedly by members of this department, who have collected a considerable quantity of fragmentary remains. During the last few years interesting new forms have been obtained by Dr F. H. Stewart, by the writer, and by senior and research students. Faunal lists and a discussion of the age of the beds occur later in this paper. The Dictyocaris-band outcrops also in the Tewel Burn. A cephalaspid scale from this section (Geol. Surv. Scot., T. 2738c and T. 2739c) is described below.

Type
Research Article
Copyright
Copyright © Royal Society of Edinburgh 1947

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References

References to Literature

Berg, L. S., 1940. “Classification of Fishes, both Recent and Fossil” (Russian, with complete English Translation), Trav. de'l Inst. Zool. Acad. Sci. U.R.S.S., V, livr. 2, Russian, 87345. English, 346–517.Google Scholar
Brotzen, F., 1933 a. “Weigeltaspis nov. gen und die Phylogenie der panzertragenden Heterostraci,” Cbl. Min. Geol. Pal., 1933, B, 648656.Google Scholar
Brotzen, F., 1933 b, 1934. “Die silurischen und devonischen Fischvorkommen in Westpodolien,” I and II, Palaeobiol., V, 423466; VI, 111–130.Google Scholar
Bryant, W. L., 1936. “A study of the Oldest Known Vertebrates, Astraspis and Eriptychius,” Proc. Amer. Phil. Soc., LXXVI, 409427.Google Scholar
Campbell, R., 1911. “Preliminary Note on the Geology of S.E. Kincardineshire,” Geol. Mag., [5], VIII, 6369. (See also Report Brit. Ass., Dundee Meeting, 1912, pp. 461–2.)CrossRefGoogle Scholar
Campbell, R., 1912 a. “Cambrian, Downtonian and Lower Old Red Sandstone Rocks near Stonehaven,” Proc. Geol. Assoc., XXIII, 291294.Google Scholar
Campbell, R., 1912 b. “On the Geology of South-Eastern Kincardineshire,” Proc. Geol. Assoc., XXIII, 295298.CrossRefGoogle Scholar
Campbell, R. 1913. “The Geology of South-Eastern Kincardineshire,” Trans. Roy. Soc. Edin., XLVIII, 923960.CrossRefGoogle Scholar
Forster-Cooper, C., 1940. Parker and Haswell's Text-book of Zoology, 11, 6th ed., revised by Forster-Cooper, C., Macmillan, London.Google Scholar
Gregory, W. K., and Raven, H. C., “Studies on the Origin and Early Evolution of Paired Fins and Limbs (four parts),” Ann. N.Y. Acad. Sci., XLII, 273360.Google Scholar
Heintz, A., 1935. “How the Fishes Learned to Swim,” Smithsonian Report for 1934, 223245.Google Scholar
Heintz, A., 1938. “Über die ältesten bekannten Wirbeltiere,” Naturwissensch., XXVI, heft 4, 4958.CrossRefGoogle Scholar
Heintz, A., 1939. “Cephalaspida from Downtonian of Norway,” Skr. Norské Vitensk.-Akad., Oslo, i, Math.-Nat. Kl., nr. 5, 119 pp., 30 pis.Google Scholar
Hoppe, K. H., 1933. “Phlebolepis elegans Pander aus dem Obersilur von Ösel.” (Bemerkungen zu dem von A. Heintz beschreibenen vollständigen Exemplar von “Coelolepis luhai.”) Cbl. Min. Geol. Pal., Abt. B., 124130 (see also important postscript, same vol., p. 368).Google Scholar
Kiaer, J., 1915. “Upper Devonian Fish Remains from Ellesmere Land, with remarks on Drepanaspis,” Rep. 2d. Norweg. Arct. Exped. Fram., no. XXXIII, 156.Google Scholar
Kiaer, J., 1924. “The Downtonian Fauna of Norway. I. Anaspida. With a geological introduction,” Skr. Vid. selsk. Kristiania, I, no. 6, 1139.Google Scholar
Kiaer, J., 1930. “Ctenaspis, a new genus of Cyathaspidian Fishes,” Skr. Svolb. Ishavet, Oslo, no. XXXIII.Google Scholar
Kiaer, J., 1931. “Hemicyclaspis Murchisoni—faunen i den downtoniske sandsten på Jelöen i Oslo-fjorden,” Norsk. Geol. Tidss., XII, 419431 (Eng. summary, 429–431), 1 pl.Google Scholar
Kiaer, J., and Heintz, A., 1932 a. “The Downtonian and Devonian Vertebrates of Spitzbergen. IV. Sub-order Cyathaspida (A Preliminary Report),” edited by Heintz, A., Skr. Svalb. Ishavet, Oslo, nr. LII, 26 pp.Google Scholar
Kiaer, J., and Heintz, A., 1932 b. “New Coelolepids from the Upper Silurian on Oesel (Esthonia),” Arch. Naturk. Estlands, (1), X, 169174.Google Scholar
Kiaer, J., and Heintz, A., 1935. “The Downtonian and Devonian Vertebrates of Spitzbergen. V. Sub-order Cyathaspida, Part I,” Skr. Svalb. Ishavet, Oslo, nr. XL.Google Scholar
Lungershausen, L., and Nikiforova, O., 1942. “On the Stratigraphical Relation of Silurian Beds of Podolia to Analagous Beds of Some Other Districts of Western Europe,” C.R. (Doklady) Acad. Sci., U.R.S.S., XXXIV, 6266.Google Scholar
Obruchev, Dm., 1944. “An attempted restoration of Psammolepis paradoxa,” C.R. (Doklady) Acad. Sci., U.R.S.S., XLII, 143145.Google Scholar
Peach, B. N., and Horne, J., 1899. “The Silurian Rocks of Britain. I. Scotland,” Mem. Geol. Surv. U.K.Google Scholar
Robertson, G. M., 1935 a. “The Ostracoderm Genus Dartmuthia Patten,” Amer. Journ. Sci., XXIX, 323335.CrossRefGoogle Scholar
Robertson, G. M., 1935 b. “Oeselaspis, a New Genus of Ostracoderm,” Amer. Journ. Sci., XXIX, 453461.Google Scholar
Robertson, G. M., 1938 a. “The Tremataspidae,” Amer. Journ. Sci., XXXV, 172206, 273–296.CrossRefGoogle Scholar
Robertson, G. M., 1938 b. “New Genera of Ostracoderms from the Upper Silurian of Oesel,” Journ. Paleont., XII, 486493.Google Scholar
Robertson, G. M., 1939 a. “The Status of Cephalaspis schrenckii Pander from the Upper Silurian of Oesel,” Journ. Geol., XLVII, 649657.CrossRefGoogle Scholar
Robertson, G. M., 1939 b. “An Upper Silurian Vertebrate Horizon, with Description of a New Species, Cephalaspis oeselensis,” Trans. Kansas Acad. Sci., XLII, 357363.Google Scholar
Robertson, G. M., 1940 a. “Witaaspis patteni, a New Ostracoderm from the Upper Silurian of Oesel,” Trans. Kansas Acad. Sci., XLIII, 297298.CrossRefGoogle Scholar
Robertson, G. M., 1940 b. “The Sensory Canal System in some Early Vertebrates,” Trans. Kansas Acad. Sci., XLIII, 467471.Google Scholar
Romer, A. S., 1933. Vertebrate Paleontology, Univ. Chicago Press.Google Scholar
Smith, Homer W., 1939. “Studies in the Physiology of the Kidney,” Porter Lectures, ser. 9, Univ. Kansas.Google Scholar
Stensiö, E. A., 1927. “The Downtonian and Devonian Vertebrates of Spitzbergen. Part 1, Family Cephalaspidae,” Skr. Svalb. Ishavet, nr. XII, Oslo.Google Scholar
Stensiö, E. A., 1932. Cephalaspids of Great Britain, British Museum (Natural History), London.CrossRefGoogle Scholar
Stetson, H. C., 1928 a. “A New American Thelodus,” Amer. Journ. Sci., CCXVI, 221231.CrossRefGoogle Scholar
Stetson, H. C., 1928 b. “A Restoration of the Anaspid Birkenia elegans Traquair,” Journ. Geol., XXXVI, 458470.Google Scholar
Stetson, H. C., 1931. “Studies on the Morphology of the Heterostraci,” Journ. Geol., XXXIX, 141154.CrossRefGoogle Scholar
Størmer, L., 1934. “Merestomata from the Downtonian Sandstone of Ringerike, Norway,” Skr. Vidensk.-Akad., Oslo, Math.-Nat. II, n, no. 10, 1125.Google Scholar
Størmer, L., 1935. “Dictyocaris Salter, a large Crustacean from the Upper Silurian and Downtonian,” Norsk. geol. tidss., XV, 267298.Google Scholar
Størmer, L., 1936. “Mixopterus dolichoschelus (Laurie M. S.), a Downtonian Eurypterid from Scotland,” Summ. Progr. Geol. Surv. Lond., year 1934, pt. II, 4147.Google Scholar
Traquair, R. H., 1899 a. “On Thelodus pagei Powrie sp. from the Old Red Sandstone of Forfarshire,” Trans. Roy. Soc. Edin., XXXIX, 595602.Google Scholar
Traquair, R. H., 1899 b. “Report on Fossil Fishes collected by the Geological Survey of Scotland in the Silurian Rocks of the South of Scotland,” Trans. Roy. Soc. Edin., XXXIX, 827864.Google Scholar
Traquair, R. H., 1905. “Supplementary Report on Fossil Fishes collected by the Geological Survey of Scotland in the Upper Silurian Rocks of Scotland,” Trans. Roy. Soc. Edin., XL, 879888.Google Scholar
Traquair, R. H., 1912. “Notes on the Fish-remains collected by Campbell, Messrs R., Gordon, W. T., and Peach, B. N. in Palaeozoic Strata at Cowie, Stonehaven,” Report Brit. Assoc, Dundee Meeting, 1912, p. 463. (Also in Geol. Mag., [5], IX, 511.)Google Scholar
Westoll, T. S., 1936. “On the Structures of the Dermal Ethmoid Shield of Osteolepis,” Geol. Mag., LXXIII, 157171.CrossRefGoogle Scholar
Westoll, T. S., 1942. “The Earliest Panzergruppen,” Aberdeen Univ. Rev., XXIX, 114122.Google Scholar
White, E. I., 1935. “The Ostracoderm Pteraspis Kner, and the Relationships of the Agnathous Vertebrates,” Phil. Trans., ser. B, CCXXV, 381457.Google Scholar
Wills, L. J., 1935. “Rare and new Ostracoderm Fishes from the Downtonian of Shropshire,” Trans. Roy. Soc. Edin., LVIII, 427447.Google Scholar
Woodward, A. S. Sir, 1933. “Report on the Fish-remains from the Little Missenden Boring,” Summ. Progr. Geol. Surv. Lond., year 1932, pt. II, 132134, 140.Google Scholar