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IX.—The Placoderm Fish Coccosteus cuspidatus Miller ex Agassiz from the Middle Old Red Sandstone of Scotland. Part I. Descriptive Morphology*

Published online by Cambridge University Press:  06 July 2012

Roger S. Miles
Affiliation:
Department of Geology, University of Newcastle upon Tyne.
T. Stanley Westoll
Affiliation:
Department of Geology, University of Newcastle upon Tyne.

Synopsis

Coccosteus cuspidatus is shown to be the correct name for the arthrodire commonly referred to as Coccosteus decipiens. An almost complete account of the osteology of this species is given and aspects of the restored head, thoracic armour, fins and posterior unarmoured region, the dermal ornamentation, neuromast system and musculature are discussed; the snout, dermal bones of the palate and neural endocranium receive special treatment in view of the part these structures have played in the interpretation of the relationships of the Arthrodira. Changes in the proportions of certain bones during growth are noted and the mode of life is considered. The family Coccosteidae is redefined; the genus Coccosteus and the type species C. cuspidatus are defined and consideration is given to other species included in this genus.

Type
Research Article
Copyright
Copyright © Royal Society of Edinburgh 1968

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References

References to Literature

Adams, L. A., 1919. “A memoir on the phylogeny of the jaw musculature in recent and fossil vertebrates”, Ann. N.Y. Acad. Sci., 28, 120127.Google Scholar
Agassiz, L., 1842. “Report on the fossil fishes of the Devonian system or Old Red Sandstone”, Rep. Br. Ass. Advmt Sci., 1842, 80.Google Scholar
Agassiz, L., 1844. Monographie des Poissons Fossiles du Vieux Grès Rouge ou Système Dévonian (Old Red Sandstone) Des lies Britanniques et de Russie. Neuchâtel.Google Scholar
Allis, E. P., 1889. “The anatomy and development of the lateral line system in Amia calva”, J. Morph., 2, 463568.Google Scholar
Allis, E. P., 1923. “The cranial anatomy of Chlamydoselachus anguineus”, Acta Zool., Stockh., 4, 123221.Google Scholar
Allis, E. P., 1936. “Comparison of the latero-sensory lines, the snout and the cranial roofing bones of the Stegocephali with those in fishes”, J. Anat., 70, 293316.Google ScholarPubMed
Backman, G., 1915. “Die Bauchflosse der Selachier. Abt. 2. Die Bauchflosse der Holocephali”, K. Svenska VetenskAkad. Handl., 53, 163.Google Scholar
Bendix-Almgren, S. E., 1967. “On the fin-structure of the Upper Permian Edestids from East Greenland”, Meddr Dansk Geol. Foren., 17, 147149.Google Scholar
Bryant, W. L., 1934. “The fish fauna of Beartooth Butte, Wyoming. Pts. II and III”, Proc. Am. Phil. Soc., 73, 127162.Google Scholar
Bystrow, A. P., 1957. “The microstructure of dermal bones in Arthrodires”, Acta Zool., Stockh., 38, 239275.CrossRefGoogle Scholar
Case, E. C., 1931. “Arthrodiran remains from the Devonian of Michigan”, Contr. Mus. Geol. Univ. Mich., 3, 163182.Google Scholar
Davidoff, M. V., 1879. “Beitrage zur vergleichenden Anatomie der Gliedmasse der Fische. I”, Morph. Jb., 5, 450520.Google Scholar
Dean, B., 1901. “On two new Arthrodires from the Cleveland Shale of Ohio”, Mem. N.Y. Acad. Sci., 2, 87100.Google Scholar
Dean, B., 1909. “Studies on fossil fishes (sharks, Chimaeroids and Arthrodires)”, Mem. Am. Mus. Nat. Hist., 9, 211287.Google Scholar
Denison, R. H., 1950. “A new Arthrodire from the New York State Devonian”, Am. J. Sci., 248, 565580.Google Scholar
Denison, R. H., 1956. “A review of the habitat of the earliest vertebrates”, Fieldiana, Geol., 11, 359457.Google Scholar
Denison, R. H., 1958. “Early Devonian fishes from Utah. 3. Arthrodira”, Fieldiana, Geol., 11, 459551.Google Scholar
Denison, R. H., 1960. “Fishes of the Devonian Holland Quarry Shale of Ohio”, Fieldiana, Geol., 11, 555613.Google Scholar
Denison, R. H., 1962. “A reconstruction of the shield of the Arthrodire Bryantolepis brachycephalus (Bryant)”, Fieldiana, Geol., 14, 99104.Google Scholar
Duff, P., 1842. Sketch of the Geology of Moray. Elgin.Google Scholar
Dunkle, D. H., 1947. “A new genus and species of Arthrodiran Fish from the Upper Devonian Cleveland Shale”, Scient. Publs Cleveland Mus. Nat. Hist., 8, 103117.Google Scholar
Dunkle, D. H., and Bungart, P. A., 1940. “On one of the least known of the Cleveland Shale Arthrodires”, Scient. Publs Cleveland Mus. Nat. Hist., 8, 2247.Google Scholar
Dunkle, D. H., 1942. “The Infero-gnathal of Titanichthys”, Scient. Publs Cleveland Mus. Nat. Hist., 8, 4959.Google Scholar
Dunkle, D. H., 1943. “Comments on Diplognathus mirabilis Newb.”, Scient. Publs Cleveland Mus. Nat. Hist., 8, 7384.Google Scholar
Dunkle, D. H., 1945. “A new Arthrodiran fish from the Upper Devonian Ohio Shales”, Scient. Publs Cleveland Mus. Nat. Hist., 8, 97102.Google Scholar
Dunkle, D. H., 1946. “The Antero-supragnathal of Gorgonichthys”, Amer. Mus. Nov., 1316, 110.Google Scholar
Eastman, C. R., 1907. “Devonic fishes of the New York Formation”, Mem. N.Y. St. Mus., 10, 1235Google Scholar
Edinger, T., 1956. “The paired pineal organs”, In Progress in Neurobiology. Elsevier, Amsterdam.Google Scholar
Egerton, P. de Grey, 1860. “Palichthyologic Notes no. 12. Remarks on the nomenclature of the Devonian fishes”, Q. Jl Geol. Soc. Lond., 16, 119136.Google Scholar
Gadow, H., and Abbott, E. C., 1895. “On the evolution of the vertebral column of fishes”, Phil. Trans. Roy. Soc., 186, 163221.Google Scholar
Gardiner, B. G., 1963. “Certain Palaeoniscoid fishes and the evolution of the snout in Actinopterygians”, Bull. Br. Mus. Nat. Hist, Geol., 8, 255325.Google Scholar
Garman, S., 1888. “On the lateral canal system of the Selachia and Holocephala”, Bull. Mus. Comp. Zool. Harv., 17, 57119.Google Scholar
Goodrich, E. S., 1930. Studies on the Structure and Development of Vertebrates. Macmillan, London.Google Scholar
Grey, J., 1933. “Studies in animal locomotion”, Proc. Roy. Soc., B, 113, 115124.Google Scholar
Gross, W., 1930. “Die Fische des mittleren Old Red Süd-Livelands”, Geol. Paläont. Abh., 18, 123156.Google Scholar
Gross, W., 1932. “Die Arthrodira Wildungens”, Geol. Paläont. Abh., 19, 161.Google Scholar
Gross, W., 1933 a. “Die Wirbeltiere des rheinischen Devons”, Abh. Preuss. Geol. Landesanst., 154, 183.Google Scholar
Gross, W., 1933 b. “Die Fische des baltischen Devons”, Palaeontographica, 79A, 174.Google Scholar
Gross, W., 1935. “Histologische Studien aus Aussenskelett fossiler Agnathen und Fische”, Palaeontographica, 83A, 160.Google Scholar
Gross, W., 1937. “Die Wirbeltiere des rheinischen Devons II”, Abh. Preuss. Geol. Landesanst., 176, 583.Google Scholar
Gross, W., 1938 a. “Über das Spinale und die angrezenden Knochen der Brachythoraci”, Neues Jb. Miner. Geol. Paläont., 79B, 403418.Google Scholar
Gross, W., 1938 b. “Rhachiosteus pterygiatus n. gen. n. sp. (Euarthrodira, Brachythoraci)”, Decheniana, 97A, 183208.Google Scholar
Gross, W., 1940. “Acanthodier und Placodermen aus Heterostius—Schichten Estland und Lettlands”, Annls Soc Reb. Nat. Invest. Univ. Tartu, 46, 188.Google Scholar
Gross, W., 1950. “Die paläontologische und stratigraphische Bedeutung der Wirbeltierfaunen des Old Reds und der marinen altpaläozoischen Schichten”, Abh. Dt. Akad. Wiss. Berl., Math.-Nat. Kl., 1949, 1130.Google Scholar
Gross, W., 1957. “Mundzähne und Hautzähne des Acanthodier und Arthrodiren”, Palaeontographica, 109A, 140.Google Scholar
Gross, W., 1959. “Arthrodiren aus dem Obersilur der Pragen Mulde”, Palaeontographica, 113A, 135.Google Scholar
Gross, W., 1960. “Tityosteus n. gen. ein Reisenarthrodire aus dem rheinischen Unterdevon”, Paläont. Z., 34, 263274.Google Scholar
Gross, W., 1961. “Lunaspis broilii und Lunaspis heroldi aus dem Hunsrückschiefer (Unterdevon, Rheinland)”, Notizbl. Hess. Landesamt. Bodenforsch. Wiesbaden, 89, 1743.Google Scholar
Gross, W., 1962 a. “Neuuntersuchung der Stensiöellida (Arthrodira, Unterdevon)”, Notizbl. Hess. Landesamt. Bodenforsch. Wiesbaden, 90, 4886.Google Scholar
Gross, W., 1962 b. “Neuuntersuchung der Dolichothoraci aus dem Unterdevon von Overath bei Köln”, Paläont. Z., H. Schmidt-Festband, 4563.CrossRefGoogle Scholar
Gross, W., 1962 c. “Peut-on homologuer les os des arthrodires et des Teleostomes?”, Colloques Int. Cent. Natn. Rech. Scient., 104, 6974.Google Scholar
Heimberg, G., 1949. “Neue Fischfunde aus dem Weissen Jura von Württemberg”, Palaeontographica, 97A, 7598.Google Scholar
Heintz, A., 1929 a. “Die downtonischen und devonischen Vertebraten von Spitzbergen, II Acanthaspida”, Skr. Svalbard Ishavet, 22, 181.Google Scholar
Heintz, A., 1929 b. “Die downtonischen und devonischen Vertebraten von Spitzbergen, III Acanthaspida”, Skr. Svalbard Ishavet, 23, 120.Google Scholar
Heintz, A., 1931 a. “Revision of the structure of Coccosteus decipiens Ag.”, Norsk Geol. Tidsskr., 12, 291313.Google Scholar
Heintz, A., 1931 b. “Untersuchungen tiber den Bau des Arthrodira”, Acta Zool., Stockh., 12, 225239.Google Scholar
Heintz, A., 1932. “The structure of Dinichthys. A contribution to our knowledge of the Arthrodira”, Am. Mus. Nat. Hist., Bashford Dean Mem. Vol., 4, 115224.Google Scholar
Heintz, A., 1933. “Some remarks about the structure of Phlyctaenaspis acadica Whiteaves”, Norsk Geol. Tidsskr., 14, 127144.Google Scholar
Heintz, A., 1934. “Revision of the Estonian Arthrodira. Pt. I. Family Homostiidae, Jaekel”, Arch. Naturk. Eestis, 10, 177290.Google Scholar
Heintz, A., 1938. “Notes on Arthrodira”, Norsk Geol. Tidsskr., 18, 127.Google Scholar
Hills, E. S., 1936. “On certain endocranial structures in Coccosteus”, Geol. Mag., 73, 213226.Google Scholar
Holmgren, N., 1942. “Studies on the head of fishes. Pt. III. The phylogeny of Elasmobranch fishes”, Acta Zool., Stockh., 23, 129261.CrossRefGoogle Scholar
Holmgren, N., and Pehrson, T., 1949. “Some remarks on the ontogenetical development of the sensory lines on the cheek in fishes and amphibia”, Acta Zool., Stockh., 30, 249–234.Google Scholar
Howell, A. B., 1933. “Morphogenesis of the shoulder architecture. Pt. II. Pisces”, Q. Rev. Biol, 8, 434456.Google Scholar
Hussakof, L., 1906. “Studies on the Arthrodira”, Mem. Am. Mus. Nat. Hist., 9, 105154.Google Scholar
Hussakof, L., 1942. “Fishes from the Devonian of Arizona”, Am. Mus. Novit., 1186, 19.Google Scholar
Hussakof, L., and Bryant, W. L., 1918. “Catalog of the fossil fishes in the Museum of the Buffalo Society of Natural Sciences”, Bull. Buffalo Soc. Nat. Sci., 12, 1198.Google Scholar
Jaekel, O., 1902. “Ueber Coccosteus und die Beurteilung der Placodermen”, Sber. Ges. Naturf. Freunde Berl., 1902, 103115.Google Scholar
Jaekel, O., 1906. “Neue Wirbeltierfunde aus dem Devon von Wildungen”, Sber. Ges. Naturf. Freunde Berl., 1906, 7885.Google Scholar
Jaekel, O., 1907. “Ueber Pholidosteus nov. gen. und die Mundbildung und die Körperform der Placodermen”, Sber. Ges. Naturf. Freunde Berl., 1907, 170186.Google Scholar
Jaekel, O., 1919. “Die Mundbildung der Placodermen”, Sber. Ges. Naturf. Freunde Berl., 1919, 73110.Google Scholar
Jaekel, O., 1925. “Das Mundskelett der Wirbeltiere”, Morph. Jb., 55, 402484.Google Scholar
Jarvik, E., 1942. “On the structure of the snout of Crossopterygians and Lower Gnathostomes in general”, Zool. Bidr. Upps., 21, 235675.Google Scholar
Jarvik, E., 1944 a. “On the dermal bones, sensory canals and pit-lines of the skull in Eusthenopteron foordi Whiteaves, with some remarks on E. sävesöderberghi Jarvik”, K. Svenska VetenskAkad. Handl., 21, 348.Google Scholar
Jarvik, E., 1944 b. “On the exoskeletal shoulder-girdle of Teleostomian fishes, with special reference to Eusthenopteron foordi Whiteaves”, K. Svenska VetenskAkad. Handl., 21, 332.Google Scholar
Jarvik, E., 1948. “On the morphology and taxonomy of the Middle Devonian Osteolepid fishes of Scotland”, K. Svenska VetenskAkad. Handl., 25, 1301.Google Scholar
Kemna, A., 1904. “Les récentes déscouvertes de Poissons Fossiles Primitifs (deuxieme notice)”, Bull. Soc. Belge Géol. Paléont. Hydrol, 18, 6578.Google Scholar
Kulczycki, J., 1956. “On the parasphenoid of Brachythoraci”, Acta Palaeont. Pol., 1, 103111.Google Scholar
Kulczycki, J., 1957. “Upper Devonian fishes from the Holy Cross Mountains (Poland)”, Acta Palaeont. Pol., 2, 285380.Google Scholar
Lehman, J. P., 1956. “Les Arthrodires du Dévonien Supérieur du Tafilalet (Sud marocain)”, Notes Mém. Serv. Mines Carte Géol Maroc, 129, 170.Google Scholar
Lehman, J. P., 1959. L'Evolution des Vertébrés Inferiers. Dunod, Paris.Google Scholar
Lehman, J. P., 1962. “A propos de la double articulation de la Cuirasse des Arthrodires”, Colloques Int. Cent. Natn. Rech. Scient., 104, 6368.Google Scholar
Liu, H. T., 1955. “Kiangyousteus, a new Arthrodiran fish from Szechuan China”, Acta Palaeont. Sin., 3, 261274.Google Scholar
McCoy, F., 1848. “On some new Ichthyolites from the Scotch Old Red Sandstone”, Ann. Mag. Nat. Hist., 2, 297312.Google Scholar
McCoy, F., 1855. “Description of the British Palaeozoic fossils in the Geological Museum of the University of Cambridge”. In A Synopsis of the Classification of the British Palaeozoic Rocks (Ed. by Sedgwick, A.). Cambridge.Google Scholar
Miles, R. S., 1962. “Gemuendenaspis n. gen., an Arthrodiran fish from the Lower Devonian Hunsrückschiefer of Germany”, Trans. Roy. Soc. Edinb., 65, 5977.CrossRefGoogle Scholar
Miles, R. S., 1964. “On some Coccosteomorph Arthrodires from the Devonian of Arizona”, Ark. Zool., 16, 427460.Google Scholar
Miles, R. S., 1965. “Ventral thoracic neuromast lines of Placoderm fishes”, Nature, Lond., 206, 524525.Google Scholar
Miles, R. S., 1966 a. “The Placoderm fish Rhachiosteus pterygiatus Gross and its relationships”, Trans. Roy. Soc. Edinb., 66, 377392.Google Scholar
Miles, R. S., 1966 b. “Protitanichthys and some other Coccosteomorph Arthrodires from the Devonian of North America”, K. Svenska VetenskAkad. Handl., 10, 149.Google Scholar
Miles, R. S., 1967 a. “The cervical joint and some aspects of the origin of the Placodermi”, Colloques Int. Cent. Natn. Rech. Scient., 163, 4971.Google Scholar
Miles, R. S., 1967 b. “Observations on the Ptyctodont fish, Rhamphodopsis Watson”, J. Linn. Soc., Zool., 47, 99120.CrossRefGoogle Scholar
Miles, R. S., and Westoll, T. S., 1963. “Two new genera of Coccosteid Arthrodira from the Middle Old Red Sandstone of Scotland and their stratigraphical distribution”, Trans. Roy. Soc. Edinb., 65, 179210.Google Scholar
Miller, H., 1841. The Old Red Sandstone. 1st edn. Edinburgh.Google Scholar
Miller, H., 1848. “On the Asterolepis and other allied genera of fossil fishes from the Old Red Sandstone”, Gdnrs' Chron., 1848, 863.Google Scholar
Miller, H., 1858. Cruise of the Betsy. London.Google Scholar
Murchison, R. I., 1840. “Fishes of the Old Red Sandstone”, Rep. Br. Ass. Advmt Sci., 1840, 99.Google Scholar
Newberry, J. S., 1875. “Descriptions of fossil fishes (the structure and relations of Dinichthys with descriptions of some new fossil forms)”, Rep. Geol. Surv. Ohio., 2, 140.Google Scholar
Nielsen, E., 1942. “Studies on Triassic fishes from East Greenland. I. Glaucolepis and Boreosomus”, Palaeozool. Grønland, 1, 1403.Google Scholar
Obruchev, D. V., 1931. “Über Coccosteus trautscholdi (Eastman)”, Moscow Acad. Sci. URSS, Mus. Geol., 8, 285310.Google Scholar
Obruchev, D. V., 1934. “Über zwei Coccosteus—Funde in der Sowiet-Arktis und den Umfang der Gattung Coccosteus”, Trudy Arkt. Nauchno-Issled Inst., 13, 185190.Google Scholar
Obruchev, D. V., 1964. “Class Placodermi”. In Foundations of Palaeontology. Reference Book for Palaeontology and Geology of the USSR. Agnatha and Fishes. Moscow. (In Russian.)Google Scholar
Obrucheva, O. P., 1954. “On the genus Plourdosteus (Arthrodira) in the Upper Devonian of the USSR”, Dokl. Akad. Nauk USSR, 96, 10551056. (In Russian.)Google Scholar
Obrucheva, O. P., 1959. “Two species of Plourdosteus (Arthrodira) in the Upper Devonian of the USSR”, Paleont. Zh., 3, 7894. (In Russian.)Google Scholar
Obrucheva, O. P., 1962. Armoured Fishes of the Devonian of the USSR (Coccosteidae and Dinichthyidae). Moscow Univ. Press. (In Russian.)Google Scholar
Obrucheva, O. P., 1966. “New facts concerning the Coccosteids (Placodermi) from the Baltic Devonian”, Paleontologiya Stratigr. Pribaltiki, Belorussk., 1, 151189. (In Russian.)Google Scholar
Ørvig, T., 1951. “Histologic studies of Placoderms and fossil Elasmobranchs. I. The endoskeleton, with remarks on the hard tissues of lower vertebrates in general”, Ark. Zool., 2, 321454.Google Scholar
Ørvig, T., 1957 a. “Remarks on the vertebrate fauna of the Lower Upper Devonian of Escuminac Bay, P.Q., Canada, with special reference to the Porolepiform Crossopterygians”, Ark. Zool., 10, 367426.Google Scholar
Ørvig, T., 1957 b. “Notes on some Palaeozoic lower vertebrates from Spitsbergen and North America”, Norsk Geol. Tidsskr., 37, 285353.Google Scholar
Ørvig, T., 1960. “New finds of Acanthodians, Arthrodires, Crossopterygians, Ganoids and Dipnoans in the Upper Middle Devonian Calcareous Flags (Oberer Plattenkalk) of the Bergisch Gladbach-Paffrath Trough. Pt. 1”, Paläont. Z., 34, 295335.Google Scholar
Ørvig, T., 1962. “Y-a-t-il une rélations directe entre les Arthrodires et les Holocéphales”, Colloques Int. Cent. Natn. Rech. Scient., 104, 4960.Google Scholar
Pander, C. H., 1857. Ueber die Placodermen des devonischen Systems. St. Petersburg.Google Scholar
Rayner, D. H., 1963. “The Achanarras Limestone of the Middle Old Red Sandstone, Caithness, Scotland”, Proc. Yorks. Geol. Soc., 34, 117136.Google Scholar
Sedgwick, A., and Murchison, R. I., 1829. “On the structure and relations of the deposits contained between the Primary Rocks and the Oolitic Series in the North of Scotland”, Trans. Geol. Soc. Lond., 3, 125160.Google Scholar
Stensiö, E. A., 1925. “On the head of the Macropetalichthyids with certain remarks on the head of the other Arthrodires”. Publs. Field Mus. Nat. Hist., Geol., 4, 87197.Google Scholar
Stensiö, E. A., 1931. “Upper Devonian vertebrates from East Greenland”, Meddr Grønland, 86, 1207.Google Scholar
Stensiö, E. A., 1934 a. “On the heads of certain Arthrodires. I. Pholidosteus, Leiosteus and Acanthaspids”, K. Svenska VetenskAkad. Handl., 13, 179.Google Scholar
Stensiö, E. A., 1934 b. “On the Placodermi of the Upper Devonian of Greenland. I. Phyllolepis and Arthrodira”, Meddr Grønland, 97, 158.Google Scholar
Stensiö, E. A., 1936. “On the Placodermi of the Upper Devonian of Greenland. Supplement to part I”, Meddr Grønland, 97, 152.Google Scholar
Stensiö, E. A., 1939. “On the Placodermi of the Upper Devonian of Greenland. 2nd supplement to part I”, Meddr Grønland, 97, 133.Google Scholar
Stensiö, E. A., 1942. “On the snout of Arthrodires”, K. Svenska VetenskAkad. Handl., 20, 932.Google Scholar
Stensiö, E. A., 1944. “Contributions to the knowledge of the vertebrate fauna of the Silurian and Devonian of Western Podolia. II. Notes on two Arthrodires from the Downtonian of Podolia”, Ark. Zool., 35A, 183.Google Scholar
Stensiö, E. A., 1945. “On the heads of certain Arthrodires. II. On the cranial and cervical joint of the Dolichothoraci (Acanthaspida)”, K. Svenska VetenskAkad. Handl., 22, 170.Google Scholar
Stensiö, E. A., 1947. “The sensory lines and dermal bones of the cheek in fishes and amphibians”, K. Svenska VetenskAkad. Handl., 2A, 1194.Google Scholar
Stensiö, E. A., 1948. “On the Placodermi of the Upper Devonian of East Greenland. II. Antiarchi: Subfamily Bothriolepinae”, Palaeozool. Grønland, 2, 1622.Google Scholar
Stensiö, E. A., 1959. “On the pectoral fin and shoulder girdle of the Arthrodires”, K. Svenska VetenskAkad. Handl., 8, 1229.Google Scholar
Stensiö, E. A., 1963. “Anatomical studies on the Arthrodiran head. Pt. 1. Preface, geological and geographical distribution, the organisation of the Arthrodires, the anatomy of the head in the Dolichothoraci, Coccosteomorphi and Pachyosteomorphi. Taxonomic appendix”, K. Svenska VetenskAkad. Handl., 9, 1419.Google Scholar
Stensiö, E. A., and Säve-Söderbergh, G., 1938. “Middle Devonian vertebrates from Canning Land and Wegener Peninsula (East Greenland). Pt. 1. Placodermi, Ichthyodorulithes”, Meddr Grønland, 96, 138.Google Scholar
Stetson, H. C., 1930. “Notes on the structure of Dinichthys and Macropetalichthys”, Bull. Mus. Comp. Zool. Harv., 71, 1939.Google Scholar
Tarlo, L. B., 1961. “Psammosteids from the Middle and Upper Devonian of Scotland”, Q. Jl Geol. Soc. Lond., 117, 193213.Google Scholar
Traquair, R. H., 1888. “Notes on the nomenclature of the fishes of the Old Red Sandstone of Great Britain”, Geol. Mag., 73, 501517.Google Scholar
Traquair, R. H., 1890. “On the structure of Coccosteus decipiens”, Ann. Mag. Nat. Hist., 5, 125136.Google Scholar
Traquair, R. H., 1896. “The extinct vertebrate animals of the Moray Firth area”. In A Vertebrate Fauna of the Moray Basin (by Brown, J. A. Harvie and Buckley, T. E.). Douglas, London.Google Scholar
Traquair, R. H., 1897. “Additional notes on the fossil fishes of the Upper Old Red Sandstone of the Moray Firth area”, Proc. Roy. Phys. Soc. Edinb., 13, 376385.Google Scholar
Wallis, F. S., 1928. “The Old Red Sandstone of the Bristol district”, Q. Jl Geol. Soc. Lond., 83, 760789.Google Scholar
Waterston, C. D., 1954. “Catalogue of type and figured specimens of fossil fishes and amphibians in the Royal Scottish Museum, Edinburgh”, Trans. Edinb. Geol. Soc., 16, 190.Google Scholar
Watson, D. M. S., 1931. “On three new species of fish from the Old Red Sandstone of Orkney and Shetland”, Mem. Geol. Surv. Summ. Prog., 1931, 157166.Google Scholar
Watson, D. M. S., 1934. “The interpretation of Arthrodires”, Proc. Zool. Soc. Lond., 1934, 437464.Google Scholar
Watson, D. M. S., 1935. “Fossil fishes of the Orcadian Old Red Sandstone”. In The Geology of the Orkneys (by Wilson, G. V. et al. ). Mem. Geol. Surv. U.K.Google Scholar
Watson, D. M. S., 1937. “The Acanthodian fishes”, Phil. Trans. Roy. Soc., B, 228, 49146.Google Scholar
Watson, D. M. S., 1938. “On Rhamphodopsis, a Ptyctodont from the Middle Old Red Sandstone”, Trans. Roy. Soc. Edinb., 59, 397410.Google Scholar
Watson, D. M. S., 1961. “Some additions to our knowledge of Antiarchi”, Palaeontology, 4, 210220.Google Scholar
Westoll, T. S., 1937. “On the cheek bones in Teleostome fishes”, J. Anat., 71, 362382.Google Scholar
Westoll, T. S., 1944. “The Haplolepidae, a new family of Late Carboniferous bony fishes. A study in taxonomy and evolution”, Bull. Am. Mus. Nat. Hist., 83, 1122.Google Scholar
Westoll, T. S., 1945. “The paired fins of Placoderms”, Trans. Roy. Soc. Edinb., 61, 381398.Google Scholar
Westoll, T. S., 1951. “The vertebrate-bearing strata of Scotland”, 18th Int. Geol. Congr., 2, 520.Google Scholar
Westoll, T. S., 1958. “The lateral fin-fold theory and the pectoral fins of Ostracoderms and early fishes”. In Studies on Fossil Vertebrates. Univ. London: Athlone Press.Google Scholar
Westoll, T. S., 1962. “Ptyctodontid fishes and the ancestry of Holocephali”, Nature, Lond., 194, 949952.Google Scholar
Westoll, T. S., and Miles, R. S., 1963. “On an Arctolepid fish from Gemünden”, Trans. Roy. Soc. Edinb., 65, 139153.Google Scholar
White, E. I., 1951. “The vertebrate faunas of the Old Red Sandstone of the Welsh Borders”, 18th Int. Geol. Congr., 2, 21.Google Scholar
White, E. I., 1952. “Australian arthrodires”, Bull. Br. Mus. Nat. Hist., Geol., 1, 251304.Google Scholar
Williams, E. E., 1959. “Gadow's Arcualia and the development of Tetrapod vertebrae”, Q. Rev. Biol., 34, 132.Google Scholar
Woodward, A. S., 1891. Catalogue of the Fossil Fishes in the British Museum (Natural History), 2. London.Google Scholar