Hostname: page-component-586b7cd67f-rcrh6 Total loading time: 0 Render date: 2024-11-25T15:56:55.565Z Has data issue: false hasContentIssue false
  • English
  • Français

A captorhinid-dominated assemblage from the palaeoequatorial Permian of Menorca (Balearic Islands, western Mediterranean)

Published online by Cambridge University Press:  22 July 2021

Rafel MATAMALES-ANDREU Open the ORCID record for Rafel MATAMALES-ANDREU [Opens in a new window] ,
Francesc X. ROIG-MUNAR ,
Oriol OMS ,
Àngel GALOBART  and
Josep FORTUNY Open the ORCID record for Josep FORTUNY [Opens in a new window]
Rafel MATAMALES-ANDREU*
Affiliation:
Institut Català de Paleontologia Miquel Crusafont, Universitat Autònoma de Barcelona, Edifici ICTA-ICP, c/ Columnes s/n, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona, Spain. Email: [email protected] Museu Balear de Ciències Naturals, ctra. Palma-Port de Sóller km 30, 07100 Sóller, Mallorca, Illes Balears, Spain.
Francesc X. ROIG-MUNAR
Affiliation:
Freelance researcher, C/ Carritxaret 18-6, 07749 es Migjorn Gran, Menorca, Illes Balears, Spain.
Oriol OMS
Affiliation:
Departament de Geologia, Universitat Autònoma de Barcelona, avda. de l'Eix Central s/n, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona, Spain.
Àngel GALOBART
Affiliation:
Institut Català de Paleontologia Miquel Crusafont, Universitat Autònoma de Barcelona, Edifici ICTA-ICP, c/ Columnes s/n, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona, Spain. Email: [email protected] Museu de la Conca Dellà, c/ del Museu 4, 25650 Isona i Conca Dellà, Lleida, Spain.
Josep FORTUNY
Affiliation:
Institut Català de Paleontologia Miquel Crusafont, Universitat Autònoma de Barcelona, Edifici ICTA-ICP, c/ Columnes s/n, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona, Spain. Email: [email protected]
*
*Corresponding author.
Get access Rights & Permissions [Opens in a new window]

Abstract

Moradisaurine captorhinid eureptiles were a successful group of high-fibre herbivores that lived in the arid low latitudes of Pangaea during the Permian. Here we describe a palaeoassemblage from the Permian of Menorca (Balearic Islands, western Mediterranean), consisting of ichnites of small captorhinomorph eureptiles, probably moradisaurines (Hyloidichnus), and parareptiles (cf. Erpetopus), and bones of two different taxa of moradisaurines. The smallest of the two is not diagnostic beyond Moradisaurinae incertae sedis. The largest one, on the other hand, shows characters that are not present in any other known species of moradisaurine (densely ornamented maxillar teeth), and it is therefore described as Balearosaurus bombardensis gen. et sp. nov. Other remains found in the same outcrop are identified as cf. Balearosaurus bombardensis gen. et sp. nov., as they could also belong to the newly described taxon. This species is sister to the moradisaurine from the lower Permian of the neighbouring island of Mallorca, and is also closely related to the North American genus Rothianiscus. This makes it possible to suggest the hypothesis that the Variscan mountains, which separated North America from southern Europe during the Permian, were not a very important palaeobiogeographical barrier to the dispersion of moradisaurines. In fact, mapping all moradisaurine occurrences known so far, it is shown that their distribution area encompassed both sides of the Variscan mountains, essentially being restricted to the arid belt of palaeoequatorial Pangaea, where they probably outcompeted other herbivorous clades until they died out in the late Permian.

Keywords

captorhinidaecentral PangaeaHyloidichnusmoradisaurinaepalaeobiogeography
Type
Articles
Information
Earth and Environmental Science Transactions of The Royal Society of Edinburgh , Volume 112 , Issue 2 , June 2021 , pp. 125 - 145
Check for updates
Copyright
Copyright © The Author(s), 2021. Published by Cambridge University Press on behalf of The Royal Society of Edinburgh

Access options

Get access to the full version of this content by using one of the access options below. (Log in options will check for institutional or personal access. Content may require purchase if you do not have access.)

References

9. References

Arche, A. & López-Gómez, J. 1992. Una nueva hipótesis sobre las primeras etapas de la evolución tectonosedimentaria de la cuenca pérmico-triásica del SE de la Cordillera Ibérica. Cuadernos de Geología Ibérica 16, 115–43.Google Scholar
Arribas, J., Gómez-Gras, D., Rosell, J. & Tortosa, A. 1990. Estudio comparativo entre las areniscas paleozoicas y triásicas de la isla de Menorca: evidencias de procesos de reciclado. Revista de la Sociedad Geológica de España 3, 105–116.Google Scholar
Bercovici, A., Diez, J. B., Broutin, J., Bourquin, S., Linol, B., Villanueva-Amadoz, U., López-Gómez, J. & Durand, M. 2009. A palaeoenvironmental analysis of Permian sediments in Minorca (Balearic Islands, Spain) with new palynological and megafloral data. Review of Palaeobotany and Palynology 158, 14–28.CrossRefGoogle Scholar
Berman, D. S., Reisz, R. R. & Eberth, D. A. 1987. Seymouria sanjuanensis (Amphibia, Batrachosauria) from the Lower Permian Cutler Formation of north-central New Mexico and the occurrence of sexual dimorphism in that genus questioned. Canadian Journal of Earth Sciences 24, 1769–84.CrossRefGoogle Scholar
Berman, D. S., Sumida, S. S. & Martens, T. 1998. Diadectes (Diadectomorpha: Diadectidae) from the early Permian of Central Germany, with description of a new species. Annals of Carnegie Museum 67, 53–93.Google Scholar
Berman, D. S. & Sumida, S. S. 1995. New cranial material of the rare diadectid Desmatodon hesperis (Diadectomorpha) from the late Pennsylvanian of central Colorado. Annals of Carnegie Museum 64, 315–36.Google Scholar
Bernardi, M., Petti, F. M., Kustatscher, E., Franz, M., Hartkopf-Fröder, C., Labandeira, C. C., Wappler, T., van Konijnenburg-van Cittert, J. H. A., Peecook, B. R. & Angielczyk, K. D. 2017. Late Permian (Lopingian) terrestrial ecosystems: a global comparison with new data from the low-latitude Bletterbach Biota. Earth-Science Reviews 175, 18–43.CrossRefGoogle Scholar
Borruel-Abadía, V., Barrenechea, J. F., Galán-Abellán, A. B., De la Horra, R., López-Gómez, J., Ronchi, A., Luque, F. J., Alonso-Azcárate, J. & Marzo, M. 2019. Could acidity be the reason behind the Early Triassic biotic crisis on land? Chemical Geology 515, 77–86.CrossRefGoogle Scholar
Bourquin, S., Bercovici, A., López-Gómez, J., Diez, J. B., Broutin, J., Ronchi, A., Durand, M., Arché, A., Linol, B. & Amour, F. 2011. The Permian–Triassic transition and the onset of Mesozoic sedimentation at the northwestern peri-Tethyan domain scale: palaeogeographic maps and geodynamic implications. Palaeogeography, Palaeoclimatology, Palaeoecology 299, 265–80.CrossRefGoogle Scholar
Bourrouilh, R. 1973. Stratigraphie, sédimentologie et téctonique de l’île de Minorque et du Nord-Est de Majorque (Baléares). La terminaison nord-orientale des Cordilleres Bétiques en Méditerranée occidentale. State PhD Dissertation, Université Pierre & Marie Curie, Paris VI, Paris.Google Scholar
Bourrouilh, R. 1983. Estratigrafía, sedimentología y tectónica de la isla de Menorca y del noreste de Mallorca. Memorias del Instituto Geológico y Minero de España 99, 1–672 + 95 pl. + 3 maps.Google Scholar
Brandes, C. & Tiedt, C. 1991. Petrographische Bestandsaufnahme der permotriassischen Sandsteine Menorcas (Balearen, westliches Mittelmeer). Neues Jahrbuch für Geologie und Paläontologie, Monatshefte 1991, 233–42.CrossRefGoogle Scholar
Briere, P. R. 2000. Playa, playa lake, sabkha: proposed definitions for old terms. Journal of Arid Environments 45, 1–7.CrossRefGoogle Scholar
Brocklehurst, N. 2017. Rates of morphological evolution in Captorhinidae: an adaptive radiation of Permian herbivores. PeerJ 5, e3200.CrossRefGoogle ScholarPubMed
Brocklehurst, N. 2020. Olson's Gap or Olson's Extinction? A Bayesian tip-dating approach to resolving stratigraphic uncertainty. Proceedings of the Royal Society B 287, 20200154.CrossRefGoogle ScholarPubMed
Brocklehurst, N., Dunne, E. M., Cashmore, D. D. & Fröbisch, J. 2018. Physical and environmental drivers of Paleozoic tetrapod dispersal across Pangaea. Nature Communications 9, 5216.CrossRefGoogle ScholarPubMed
Broutin, J., Ferrer, J., Gisbert, J. & Nmila, A. 1992. Permière découverte d'une microflore thuringienne dans le faciès saxonien de l’île de Minorque (Baléares, Espagne). Comptes Rendus de l'Académie des Sciences de Paris, s.2 315, 117–22.Google Scholar
Buenaventura, A. 2004. Menorca, Atlas Náutico. Maó: Editorial Menorca S.A.Google Scholar
Calafat, F. 1988. Estratigrafía y sedimentología de la litofacies Buntsandstein de Mallorca. Bachelor's Thesis, Universitat de Barcelona, Barcelona.Google Scholar
Calafat, F., Fornós, J. J., Marzo, M., Ramos, E. & Rodríguez-Perea, A. 1986. Icnología de vertebrados de las facies Buntsandstein de Mallorca. In Cabrera, L. (ed.) XI Congreso Español de Sedimentología. Libro de Resúmenes, 40. Barcelona: Universitat de Barcelona.Google Scholar
Calafat, F., Fornós, J. J., Marzo, M., Ramos-Guerrero, E. & Rodríguez-Perea, A. 1986–1987. Icnología de vertebrados de las facies Buntsandstein de Mallorca. Acta Geològica Hispànica 21–22, 515–20.Google Scholar
Carmona, R. 2004. Estudi d'un seimuriamorfe (Amphibia: Batrachosauria) del Permià superior de Menorca. Bachelor's Thesis, Universitat de Barcelona, Barcelona.Google Scholar
Carroll, R. L., Kuhn, O. & Tatarinov, L. P. 1972. Handbuch der Paläoherpetologie/Encyclopedia of Paleoherpetology. Teil 5B/Part 5B: Batrachosauria (Anthracosauria) Gephyrostegida–Chroniosuchida. Stuttgart and Portland: Gustav Fischer Verlag.Google Scholar
Case, E. C. 1911. A revision of the Cotylosauria of North America. Carnegie Institution of Washington Publication 145, 1–121.Google Scholar
Case, E. C. & Williston, S. W. 1912. A description of the skills of Diadectes lentus and Animasaurus carinatus. American Journal of Science, s. 4 33, 339–48.CrossRefGoogle Scholar
Cisneros, J. C., Angielczyk, K., Kammerer, C. F., Smith, R. M. H., Fröbisch, J., Marsicano, C. A. & Richter, M. 2020a. Captorhinid reptiles from the lower Permian Pedra de Fogo Formation, Piauí, Brazil: the earliest herbivorous tetrapods in Gondwana. PeerJ 8, e8719.CrossRefGoogle Scholar
Cisneros, J. C., Kammerer, C. F., Angielczyk, K. D., Fröbisch, J., Marsicano, C., Smith, R. M. & Richter, M. 2020b. A new reptile from the lower Permian of Brazil (Karutia fortunata gen. et sp. nov.) and the interrelationships of Parareptilia. Journal of Systematic Palaeontology 18, 1939–59.CrossRefGoogle Scholar
Citton, P., Ronchi, A., Maganuco, S., Caratelli, M., Nicosia, U., Sacci, E. & Romano, M. 2019. First tetrapod footprints from the Permian of Sardinia and their palaeontological and stratigraphical significance. Geological Journal 54, 2084–98.CrossRefGoogle Scholar
Contessi, M., Voigt, S. & Bibonne, R. 2018. Permian tetrapod footprints from Tunisia. Ichnos 25, 119–27.CrossRefGoogle Scholar
Cope, E. D. 1896. The reptilian order Cotylosauria. Proceedings of the American Philosophical Society 34, 436–57.Google Scholar
Crespí, D. & Merino, A. 1998. Contribució al coneixement de les mines de coure situades en el Permo-Trias de Menorca. Endins 22, 119–23.Google Scholar
Day, M. O., Ramezani, J., Bowring, S. A., Sadler, P. M., Erwin, D. H., Abdala, F. & Rubidge, B. S. 2015. When and how did the terrestrial mid-Permian mass extinction occur? Evidence from the tetrapod record of the Karoo Basin, South Africa. Proceedings of the Royal Society B 282, 20150834.CrossRefGoogle ScholarPubMed
de Ricqlès, A. & Bolt, J. R. 1983. Jaw growth and tooth replacement in Captorhinus aguti (Reptilia: Captorhinomorpha): a morphological and histological analysis. Journal of Vertebrate Paleontology 3, 7–24.CrossRefGoogle Scholar
de Ricqlès, A. & Taquet, P. 1982. La faune de vertébrés du Permien Supérieur du Niger. I. Le captorhinomorphe Moradisaurus grandis (Reptilia, Cotylosauria). Le crane. Annales de Paléontologie (Vert.-Invert.) 68, 33–106.Google Scholar
Dodick, J. T. & Modesto, S. P. 1995. The cranial anatomy of the captorhinid reptile Labidosaurikos meachami from the lower Permian of Oklahoma. Palaeontology 38, 687–711.Google Scholar
Dutuit, J. M. 1976. Il est probable que les Rhynchocéphales sont représentés dans la faune du Trias marocain. Compte-Rendus de l'Académie des Sciences, Paris 238, 483–86.Google Scholar
Falkingham, P. L. 2012. Acquisition of high resolution three-dimensional models using free, open-source, photogrammetric software. Palaeontologia Electronica 15, 1T:15p.Google Scholar
Franzel, M., Jones, S. J., Meadows, N., Allen, M. B., McCaffrey, K. & Morgan, T. 2021. Basin-scale fluvial correlation and response to the Tethyan marine transgression: an example from the Triassic of central Spain. Basin Research 33, 1–25.CrossRefGoogle Scholar
Gand, G. 1987. Les traces de vertébrés tétrapodes du Permien français: paléontologie, stratigraphie, paléoenvironnements. State PhD Dissertation, Université de Bourgogne, Dijon.Google Scholar
Gand, G. 1993. La palichnofaune de vertébrés tétrapodes du basin permien de Saint-Affrique (Aveyron): comparaisons et conséquences stratigraphiques. Géologie de la France 1, 41–56.Google Scholar
Gand, G., Kerp, H., Parsons, C. & Martínez-García, E. 1997. Palaeoenvironmental and stratigraphic aspects of animal traces and plant remains in Spanish Permian red beds (Peña Sagra, Cantabrian Mountains, Spain). Geobios 30, 295–318.CrossRefGoogle Scholar
Gand, G., Garric, J., Demathieu, G. & Ellenberger, P. 2000. La palichnofaune de vertébrés tétrapodes du Permien superiéur du bassin de Lodève (Languedoc-France). Palaeovertebrata 29, 1–82.Google Scholar
Gand, G., De La Horra, R., Galán-Abellán, B., López-Gómez, J., Barrenechea, J. F., Arche, A. & Benito, M. I. 2010. New ichnites from the Middle Triassic of the Iberian Ranges (Spain): palaeoenvironmental and paleogeographical implications. Historical Biology 22, 40–56.CrossRefGoogle Scholar
Gand, G., Tüysüz, O., Steyer, J. S., Allain, R., Sakınç, M., Sanchez, S., Șengör, A. M. C. & Sen, S. 2011. New Permian tetrapod footprints and macroflora from Turkey (Çakraz Formation, northwestern Anatolia): biostratigraphic and palaeoenvironmental implications. Comptes Rendus Palevol 10, 617–25.CrossRefGoogle Scholar
Gand, G. & Durand, M. 2006. Tetrapod footprint ichno-associations from French Permian basins. Comparisons with other Euramerican ichnofaunas. Geological Society, London, Special Publications 265, 157–77.CrossRefGoogle Scholar
Gilmore, C. W. 1927. Fossil footprints of the Grand Canyon: second contribution. Smithsonian Miscellaneous Collections 80, 1–78 + 21 pls.Google Scholar
Gómez-Gras, D. M. 1987. Estratigrafía física y petrología sedimentaria del Pérmico y Buntsandstein de la isla de Menorca. Bachelor's Thesis, Universitat Autònoma de Barcelona, Barcelona.Google Scholar
Gómez-Gras, D. M. 1992. El Permotrías de las Baleares, de la Cordillera Costero Catalana y de la vertiente mediterránea de la Cordillera Ibérica: Facies y Petrología Sedimentaria. PhD Dissertation, Universitat Autònoma de Barcelona, Barcelona.Google Scholar
Gómez-Gras, D. 1993. El Permotrías de las Baleares y de la vertiente mediterránea de la Cordillera Ibérica y del Maestrat: Facies y Petrología Sedimentaria (Parte II). Boletín Geológico y Minero 104, 467–515.Google Scholar
Gómez-Gras, D. & Alonso-Zarza, A. M. 2003. Reworked calcretes: their significance in the reconstruction of alluvial sequences (Permian and Triassic, Minorca, Balearic Islands, Spain). Sedimentary Geology 158, 299–319.CrossRefGoogle Scholar
Hasiotis, S. T., Platt, B. F., Hembree, D. I. & Everhart, M. J. 2007. The trace-fossil record of vertebrates. In William, M. III (ed.) Trace fossils: concepts, problems, prospects, 196–218. Amsterdam & Oxford: Elsevier Science.CrossRefGoogle Scholar
Haubold, H. 2000. Tetrapodenfährten aus dem Perm – Kenntnisstand und Progress 2000. Hallesches Jahrbuch für Geowissenschaften, B 22, 1–16.Google Scholar
Haubold, H., Hunt, A. P., Lucas, S. G. & Lockley, M. G. 1995. Wolfcampian (Early Permian) vertebrate tracks from Arizona and New Mexico. New Mexico Museum of Natural History and Science Bulletin 6, 135–65.Google Scholar
Heyler, D. & Lessertisseur, J. 1963. Pistes de tétrapodes Permiens dans la région de Lodève (Hérault). Muséum National d'Histoire Naturelle, C 11, 1–100.Google Scholar
Hmich, D., Schneider, J. W., Saber, H., Voigt, S. & El Wartiti, M. 2006. New continental Carboniferous and Permian faunas of Morocco: implications for biostratigraphy, palaeobiogeography and palaeoclimate. Geological Society, London, Special Publications 265, 297–324.CrossRefGoogle Scholar
Hminna, A., Voight, A., Saber, H., Schneider, J. W. & Hmich, D. 2012. On a moderately diverse continental ichnofauna from the Permian Ikakern Formation (Argana Basin, Western High Atlas, Morocco). Journal of African Earth Sciences 68, 15–23.CrossRefGoogle Scholar
Holmes, R. B. 2003. The hind limb of Captorhinus aguti and the step cycle of basal amniotes. Canadian Journal of Earth Sciences 40, 515–26.CrossRefGoogle Scholar
Ivakhnenko, M. F. 2008. Podklass Captorhinomorpha [Subclass Captorhinomorpha]. In Ivakhnenko, M. F. & Kurotchkin, E. N. (eds) Iskopaemye pozvonotchnye Rossii i sopredel'nykh stran: Iskopaemye reptilii i ptitsy, Tchast’ 1 [Fossil vertebrates of Russia and adjacent countries: Fossil reptiles and birds, Part 1], 86–94. Moskva: GEOS.Google Scholar
Jalil, N. E. & Dutuit, J. M. 1996. Permian captorhinid reptiles from the Argana Formation, Morocco. Palaeontology 39, 907–18.Google Scholar
Jung, J. P. & Sumida, S. S. 2017. A juvenile of the multiple-tooth-rowed reptile Labidosaurikos (Eureptilia, Captorhinidae, Moradisaurinae) from the Lower Permian of north-central Texas. PaleoBios 34, 1–5.CrossRefGoogle Scholar
Kissel, R. 2010. Morphology, phylogeny, and evolution of Diadectidae (Cotylosauria: Diadectomorpha). PhD Dissertation, University of Toronto, Toronto.Google Scholar
Kuhn, O. 1961. Fossilium Catalogus. I: Animalia. Pars 99. Reptilia (Supplementum 1(2)). The Hague, Netherlands: Uitgeverij Dr. W. Junk.Google Scholar
Kuhn, O. 1969. Handbuch der Paläoherpetologie/Encyclopedia of Paleoherpetology. Teil 6/Part 6: Cotylosauria. Stuttgart and Portland: Gustav Fischer Verlag.Google Scholar
Laurenti, J. N. 1768. Specimen Medicum, Exhibens Synopsin Reptilium Emendatam cum Experimentis Circa Venena et Antidota Reptilium Austriacorum. Vienna: Typis Joannis Thomae Nobilis de Trattnern.CrossRefGoogle Scholar
LeBlanc, A. R. H., Brar, A. K., May, W. & Reisz, R. R. 2015. Multiple tooth-rowed captorhinids from the early Permian fissure fills of the Bally Mountain Locality of Oklahoma. Vertebrate Anatomy Morphology Palaeontology 1, 35–49.CrossRefGoogle Scholar
LeBlanc, A. R. H. & Reisz, R. R. 2015. Patterns of tooth development and replacement in captorhinid reptiles: a comparative approach for understanding the origin of multiple tooth rows. Journal of Vertebrate Paleontology 35, e919928.CrossRefGoogle Scholar
Leonardi, G. 1987. Glossary and manual of tetrapod footprint palaeoichnology. Brasília: Departamento Nacional de Produção Mineral.Google Scholar
Liebrecht, T., Fortuny, J., Galobart, À, Müller, J. & Sander, P. M. 2017. A large, multiple-tooth-rowed captorhinid reptile (Amniota: Eureptilia) from the Upper Permian of Mallorca (Balearic Islands, Western Mediterranean). Journal of Vertebrate Paleontology 37, e1251936.CrossRefGoogle Scholar
Linol, B., Bercovici, A., Bourquin, S., Diez, J. B., López-Gómez, J., Broutin, J., Durand, M. & Villanueva-Amadoz, U. 2009. Late Permian to Middle Triassic correlations and palaeogeographical reconstructions in south-western European basins: new sedimentological data from Minorca (Balearic Islands, Spain). Sedimentary Geology 220, 77–94.CrossRefGoogle Scholar
Llull, B. & Perelló, L. 2013. La mineria de coure a Menorca. De la prehistòria a l'actualitat. Bolletí de la Societat Arqueològica Lul⋅liana. Revista d'Estudis Històrics 69, 75–93.Google Scholar
Logghe, A., Mujal, E., Marchetti, L., Nel, A., Pouillon, J.-M., Giner, S., Garrouste, R. & Steyer, J.-S. 2021. Hyloidichnus trackways with digit and tail drag traces from the Permian of Gonfaron (Var, France): new insights on the locomotion of captorhinomorph eureptiles. Palaeogeography, Palaeoclimatology, Palaeoecology 573, 110436.CrossRefGoogle Scholar
López-Gómez, J., Alonso-Azcárate, J., Arche, A., Arribas, J., Fernández-Barrenechea, J., Borruel-Abadía, V., Bourquin, S., Cadenas, P., Cuevas, J., De la Horra, R., Díez, J. B., Escudero-Mozo, M. J., Fernández-Viejo, G., Galán-Abellán, B., Galé, C., Gaspar-Escribano, J., Gisbert-Aguilar, J., Gómez-Gras, D., Goy, A., Gretter, N., Heredia-Carballo, N., Lago, M., Lloret, J., Luque, J., Márquez, L., Márquez-Aliaga, A., Martín-Algarra, A., Martín-Chivelet, J., Martín-González, F., Marzo, M., Mercedes-Martín, R., Ortí, F., Pérez-López, A., Pérez-Valera, F., Pérez-Valera, J. A., Plasencia, P., Ramos, E., Rodríguez-Méndez, L., Ronchi, A., Salas, R., Sánchez-Fernández, D., Sánchez-Moya, Y., Sopeña, A., Suárez-Rodríguez, Á., Tubía, J. M., Ubide, T., Valero-Garcés, B., Vargas, H. & Viseras, C. 2019a. Permian-Triassic rifting stage. In Vergés, J. & Kullberg, J. C. (coords) The geology of Iberia: a geodynamic approach. Volume 3: the alpine cycle, 29–112. Cham: Springer.CrossRefGoogle Scholar
López-Gómez, J., Martín-González, F., Heredia, N., De la Horra, R., Barrenechea, J. F., Cadenas, P., Juncal, M., Diez, J. B., Borruel-Abadía, V., Pedreira, D., García-Sansegundo, J., Farias, P., Galé, C., Lago, M., Ubide, T., Fernández-Viejo, G. & Gand, G. 2019b. New lithostratigraphy for the Cantabrian Mountains: a common tectono-stratigraphic evolution for the onset of the Alpine cycle in the W Pyrenean realm, N Spain. Earth-Science Reviews 188, 249–71.CrossRefGoogle Scholar
López-Gómez, J. & Arche, A. 1987. Evolución sedimentológica de la Unidad «Limos y areniscas de Alcotas». Tramo medio de la Facies Buntsandstein del sector SE de la Rama Castellana de la Cordillera Ibérica (Provincias de Cuenca y Valencia). Acta Geològica Hispànica 21–22, 9–18.Google Scholar
López-Gómez, J. & Arche, A. 1992. Las unidades litoestratigráficas del Pérmico y Triásico Inferior y Medio en el sector SE de la Cordillera Ibérica. Estudios Geológicos 48, 123–43.CrossRefGoogle Scholar
Lucas, S. G. 2019. An ichnological perspective on some major events of Paleozoic tetrapod evolution. Bollettino della Società Paleontologica Italiana 58, 223–66.Google Scholar
Lucas, S. G., Krainer, K., Chaney, D. S., DiMichele, W. A., Voigt, S., Berman, D. S. & Henrici, A. C. 2013. The Lower Permian Abo Formation in central New Mexico. New Mexico Museum of Natural History and Science Bulletin 59, 161–80.Google Scholar
Mack, G. H., James, W. C. & Monger, H. C. 1993. Classification of paleosols. Geological Society of America Bulletin 105, 129–36.2.3.CO;2>CrossRefGoogle Scholar
Marchetti, L. 2016. New occurrences of tetrapod ichnotaxa from the Permian Orobic Basin (Northern Italy) and critical discussion of the age of the ichnoassociation. Papers in Palaeontology 2, 363–86.CrossRefGoogle Scholar
Marchetti, L., Avanzini, M. & Conti, M. A. 2013. Hyloidichnus bifurcatus Gilmore, 1927 and Limnopus heterodactylus (King, 1845) from the Early Permian of Southern Alps (N Italy): a new equilibrium in the ichnofauna. Ichnos 20, 202–17.CrossRefGoogle Scholar
Marchetti, L., Santi, G. & Avanzini, M. 2014. The problem of small footprints in paleoichnology: remarks on the early Permian ichnotaxon Erpetopus cassinsi, a local species from southern Alps (northern Italy). Rivista Italiana di Paleontologia e Stratigrafia 120, 129–43.Google Scholar
Marchetti, L., Forte, G., Bernardi, M., Wappler, T., Hartkopf-Fröder, C., Krainer, K. & Kustatscher, E. 2015a. Reconstruction of a Late Cisuralian (Early Permian) floodplain lake environment: palaeontology and sedimentology of the Tregiovo Basin (Trentino–Alto Adige, Northern Italy). Palaeogeography, Palaeoclimatology, Palaeoecology 440, 180–220.CrossRefGoogle Scholar
Marchetti, L., Ronchi, A., Santi, G., Schirolli, P. & Conti, M. A. 2015b. Revision of a classic site for Permian tetrapod ichnology (Collio Formation, Trompia and Caffaro valleys, N. Italy), new evidences for the radiation of captorhinomorph footprints. Palaeogeography, Palaeoclimatology, Palaeoecology 433, 140–55.CrossRefGoogle Scholar
Marchetti, L., Ronchi, A., Santi, G. & Voigt, S. 2015c. The Gerola Valley site (Orobic Basin, Northern Italy): a key for understanding late Early Permian tetrapod ichnofaunas. Palaeogeography, Palaeoclimatology, Palaeoecology 439, 97–116.CrossRefGoogle Scholar
Marchetti, L., Belvedere, M., Voigt, S., Klein, H., Castanera, D., Díaz-Martínez, I., Marty, D., Xing, L., Feola, S., Melchor, R. N. & Farlow, J. O. 2019a. Defining the morphological quality of fossil footprints. Problems and principles of preservation in tetrapod ichnology with examples from the Palaeozoic to the present. Earth-Science Reviews 193, 109–45.CrossRefGoogle Scholar
Marchetti, L., Klein, H., Buchwitz, M., Ronchi, A., Smith, R. M. H., De Klerk, W. J., Sciscio, L. & Groenewald, G. H. 2019b. Permian-Triassic vertebrate footprints from South Africa: ichnotaxonomy, producers and biostratigraphy through two major faunal crises. Gondwana Research 72, 139–68.CrossRefGoogle Scholar
Marchetti, L., Voigt, S., Lucas, S. G., Francischini, H., Dentzien-Dias, P., Sacchi, R., Mangiacotti, M., Scali, S., Gazzola, A., Ronchi, A. & Millhouse, A. 2019c. Tetrapod ichnotaxonomy in Eolian paleoenvironments (Coconino and De Chelly formations, Arizona) and late Cisuralian (Permian) sauropsid radiation. Earth-Science Reviews 190, 148–70.CrossRefGoogle Scholar
Marchetti, L., Voigt, S., Lucas, S. G., Stimson, M. R., King, O. A. & Calder, J. H. 2020a. Footprints of the earliest reptiles: Notalacerta missouriensis – ichnotaxonomy, potential trackmakers, biostratigraphy, palaeobiogeography and palaeoecology. Annales Societatis Geologorum Poloniae 90, 271–90.Google Scholar
Marchetti, L., Voigt, S., Mujal, E., Lucas, S. G., Francischini, H., Fortuny, J. & Santucci, V. L. 2020b. Extending the footprint record of Pareiasauromorpha to the Cisuralian: earlier appearance and wider palaeobiogeography of the group. Papers in Palaeontology 2020, 1–23.Google Scholar
Martí, J., Paniello, X., Pomar, L., Ramos, E. & Rodríguez-Perea, A. 1985. El Triásico de las Baleares. In Mateu Ibars, F. & Marzo, M. (eds) II Coloquio de Estratigrafía y Paleogeografía del Pérmico y Triásico de España, 84–85. La Seu d'Urgell: Institut d'Estudis Ilerdencs and Grupo Español del Mesozoico.Google Scholar
Martin-Closas, C. & Ramos, E. 2005. Palaeogene Charophytes of the Balearic Islands (Spain). Geologica Acta 3, 39–58.Google Scholar
Matamales-Andreu, R., Fortuny, J., Mujal, E. & Galobart, À. 2019. Tetrapod tracks from the Permian of Mallorca (western Mediterranean): preliminary data, biostratigraphic and biogeographic inferences. In The Palaeontological Association, 63rd annual meeting, 15th–21st December 2019, 107. València: The Palaeontological Association.Google Scholar
Melchor, R. N. & Sarjeant, W. A. S. 2004. Small amphibian and reptile footprints from the Permian Carapacha Basin, Argentina. Ichnos 11, 57–78.CrossRefGoogle Scholar
Miall, A. D. 1985. Architectural-element analysis: a new method of facies analysis applied to fluvial deposits. Earth-Science Reviews 22, 261–308.CrossRefGoogle Scholar
Miall, A. D. 2006. The geology of fluvial deposits: sedimentary facies, basin analysis, and petroleum geology. 4th edn. Berlin, Heidelberg, New York: Springer.CrossRefGoogle Scholar
Michel, L. A., Tabor, N. J., Montañez, I. P., Schmitz, M. D. & Davydov, V. I. 2015. Chronostratigraphy and Paleoclimatology of the Lodève Basin, France: evidence for a pan-tropical aridification event across the Carboniferous–Permian boundary. Palaeogeography, Palaeoclimatology, Palaeoecology 430, 118–31.CrossRefGoogle Scholar
Modesto, S. P., Scott, D. M., Berman, D. S., Müller, J. & Reisz, R. R. 2007. The skull and the palaeoecological significance of Labidosaurus hamatus, a captorhinid reptile from the Lower Permian of Texas. Zoological Journal of the Linnean Society 149, 237–62.CrossRefGoogle Scholar
Modesto, S. P., Lamb, A. J. & Reisz, R. R. 2014. The captorhinid reptile Captorhinikos valensis from the lower Permian Vale Formation of Texas, and the evolution of herbivory in eureptiles. Journal of Vertebrate Paleontology 34, 291–302.CrossRefGoogle Scholar
Modesto, S. P., Fear, V. J., Danley, M. M. & Reisz, R. R. 2016. A large moradisaurine tooth plate from the lower Permian of Texas and its biostratigraphic implications. Journal of Vertebrate Paleontology 36, e1221832.CrossRefGoogle Scholar
Modesto, S. P., Scott, D. & Reisz, R. R. 2018. A new small captorhinid reptile from the lower Permian of Oklahoma and resource partitioning among small captorhinids in the Richards Spur fauna. Papers in Palaeontology 4, 293–307.CrossRefGoogle Scholar
Modesto, S. P., Richards, C. D., Ide, O. & Sidor, C. A. 2019. The vertebrate fauna of the upper Permian of Niger — X. The mandible of the captorhinid reptile Moradisaurus grandis. Journal of Vertebrate Paleontology 38, e1531877.CrossRefGoogle Scholar
Moodie, R. L. 1929. Vertebrate footprints from the Red Beds of Texas. American Journal of Science 97, 352–68.CrossRefGoogle Scholar
Moreau, J.-D., Benaouiss, N., Tourani, A., Steyer, J.-S., Laurin, M., Peyer, K., Béthoux, O., Aouda, A. & Jalil, N.-E. 2020. A new ichnofauna from the Permian of the Zat Valley in the Marrakech High Atlas of Morocco. Journal of African Earth Sciences 172, 103973.CrossRefGoogle Scholar
Mujal, E., Fortuny, J., Oms, O., Bolet, A., Galobart, À & Anadón, P. 2016. Palaeoenvironmental reconstruction and early Permian ichnoassemblage from the NE Iberian Basin (Pyrenean Basin). Geological Magazine 153, 578–600.CrossRefGoogle Scholar
Mujal, E., Fortuny, J., Pérez-Cano, J., Dinarès-Turell, J., Ibáñez-Insa, J., Oms, O., Vila, I., Bolet, A. & Anadón, P. 2017. Integrated multi-stratigraphic study of the Coll de Terrers late Permian–Early Triassic continental succession from the Catalan Pyrenees (NE Iberian Peninsula): a geologic reference record for equatorial Pangaea. Global and Planetary Change 159, 46–60.CrossRefGoogle Scholar
Mujal, E., Marchetti, L., Schoch, R. R. & Fortuny, J. 2020. Upper Paleozoic to lower Mesozoic tetrapod ichnology revisited: photogrammetry and relative depth pattern inferences on functional prevalence of autopodia. Frontiers in Earth Science 8, 1–23.CrossRefGoogle Scholar
Müller, J. & Reisz, R. R. 2005. An early captorhinid reptile (Amniota, Eureptilia) from the Upper Carboniferous of Hamilton, Kansas. Journal of Vertebrate Paleontology 25, 561–68.CrossRefGoogle Scholar
Nyakatura, J. A., Allen, V. R., Lauströer, J., Andikfar, A., Danczak, M., Ullrich, H.-J., Hufenbach, W., Martens, T. & Fischer, M. S. 2015. A three-dimensional skeletal reconstruction of the stem amniote Orobates pabsti (Diadectidae): analyses of body mass, centre of mass position, and joint mobility. PLoS ONE 10, e0137284.CrossRefGoogle ScholarPubMed
O'Keefe, F. R., Sidor, C. A., Larsson, H. C., Maga, A. & Ide, O. 2005. The vertebrate fauna of the upper Permian of Niger—III, Morphology and ontogeny of the hindlimb of Moradisaurus grandis (Reptilia, Captorhinidae). Journal of Vertebrate Paleontology 25, 309–19.CrossRefGoogle Scholar
Obrador, A. 1983. El Pérmico de las Baleares. In Martínez-Díaz, C. (ed.). Carbonífero y Pérmico de España, X Congreso Internacional de Estratigrafía y Geología del Carbonífero, 463–470. Madrid: Minsterio de Industria y Energía, Instituto Geológico y Minero de España.Google Scholar
Olroyd, S. L. & Sidor, C. A. 2017. A review of the Guadalupian (middle Permian) global tetrapod fossil record. Earth-Science Reviews 171, 583–97.CrossRefGoogle Scholar
Olson, E. C. 1947. The family Diadectidae and its bearing on the classification of reptiles. Fieldiana: Geology 11, 1–53.Google Scholar
Olson, E. C. 1954. Fauna of the Vale and Choza: 9. Captorhinomorpha. Fieldiana: Geology 10, 211–18.Google Scholar
Olson, E. C. 1962a. Late Permian terrestrial vertebrates, U.S.A. and U.S.S.R. Transactions of the American Philosophical Society 52, 1–224.CrossRefGoogle Scholar
Olson, E. C. 1962b. Permian vertebrates from Oklahoma and Texas. Part II. — The osteology of Captorhinikos chozaensis Olson. Oklahoma Geological Survey Circular 59, 49–68.Google Scholar
Olson, E. C. 1965. New Permian vertebrates from the Chickasha Formation in Oklahoma. Oklahoma Geological Survey Circular 70, 1–70.Google Scholar
Olson, E. C. 1970. New and little known genera and species of vertebrates from the Lower Permian of Oklahoma. Fieldiana: Geology 18, 359–434.Google Scholar
Olson, E. C. 1979. Seymouria grandis n. sp. (Batrachosauria: Amphibia) from the middle Clear Fork (Permian) of Oklahoma and Texas. Journal of Paleontology 53, 720–28.Google Scholar
Olson, E. C. & Barghusen, H. 1962. Permian vertebrates from Oklahoma and Texas. Part I. — Vertebrates from the Flowerpot Formation, Permian of Oklahoma. Oklahoma Geological Survey Circular 59, 4–48.Google Scholar
Olson, E. C. & Beerbower, J. R. 1953. The San Angelo Formation, Permian of Texas, and its vertebrates. The Journal of Geology 61, 389–423.CrossRefGoogle Scholar
Pomar-Gomà, L. 1979. The Triassic of the Balearic Islands. Palma: Geology Department, Palma de Mallorca University.Google Scholar
Postma, G. 1990. Depositional architecture and facies of river and fan deltas: a synthesis. Special publications of the international association of sedimentologists 10, 13–27.Google Scholar
Pretus, J. L. & Obrador, A. 1987. Presencia de restos óseos en el Pérmico de Menorca (nota previa). Bolletí de la Societat d'Història Natural de Balears 31, 149–52.Google Scholar
Ramos, A. & Doubinger, J. 1989. Premières datations palynologiques dans le facies Buntsandstein de l’île de Majorque (Baléares, Espagne). Comptes Rendus de l'Académie des Sciences de Paris, s. 2 309, 1089–94.Google Scholar
Reisz, R. R., Liu, J., Li, J.-L. & Müller, J. 2011. A new captorhinid reptile, Gansurhinus qingtoushanensis, gen. et sp. nov., from the Permian of China. Naturwissenschaften 98, 435–41.CrossRefGoogle ScholarPubMed
Reisz, R. R., Haridy, Y. & Müller, J. 2016. Euconcordia nom. nov., a replacement name for the captorhinid eureptile Concordia Müller and Reisz, 2005 (non Kingsley, 1880), with new data on its dentition. Vertebrate Anatomy Morphology Palaeontology 3, 1–6.CrossRefGoogle Scholar
Reisz, R. R., MacDougall, M. J., LeBlanc, A. R. H., Scott, D. & Nagesan, R. S. 2020. Lateralized feeding behaviour in a Paleozoic reptile. Current Biology 30, 2374–78.CrossRefGoogle Scholar
Reisz, R. R. & Sues, H.-D. 2000. Herbivory in late Paleozoic and Triassic terrestrial vertebrates. In Sues, H.-D. (ed) Evolution of herbivory in terrestrial vertebrates, 9–41. Cambridge, New York, Melbourne, Madrid, Cape Town, Singapore, São Paulo: Cambridge University Press.CrossRefGoogle Scholar
Rodríguez-Perea, A., Ramos-Guerrero, E., Pomar, L., Paniello, X., Obrador, A. & Martí, J. 1987. El Triásico de las Baleares. Cuadernos de Geología Ibérica 11, 295–321.Google Scholar
Romano, M. & Rubidge, B. 2019. Long bone scaling in Captorhinidae: do limb bones scale according to elastic similarity in sprawling basal amniotes? Lethaia 52, 389–402.CrossRefGoogle Scholar
Roscher, M., Stordal, F. & Svensen, H. 2011. The effect of global warming and global cooling on the distribution of the latest Permian climate zones. Palaeogeography, Palaeoclimatology, Palaeoecology 309, 186–200.CrossRefGoogle Scholar
Rosell, J., Arribas, J., Elízaga, E. & Gómez-Gras, D. 1988. Caracterización sedimentológica y petrográfica de la serie roja permo–triásica de la isla de Menorca. Boletín Geológico y Minero 99, 71–82.Google Scholar
Rosell, J., Gómez-Gras, D. & Elízaga, E. 1989a. Mapa Geológico de España. Escala 1:25.000. Isla de Menorca. Cap Menorca y Ciutadella. Segunda serie, primera edición. Madrid: Instituto Tecnológico GeoMinero de España.Google Scholar
Rosell, J., Gómez-Gras, D. & Elízaga, E. 1989b. Mapa Geológico de España. Escala 1:25.000. Isla de Menorca. Cala en Brut i Alaior. Segunda serie, primera edición. Madrid: Instituto Tecnológico GeoMinero de España.Google Scholar
Sàbat, F., Gelabert, B. & Rodríguez-Perea, A. 2018. Minorca, an exotic Balearic island (western Mediterranean). Geologica Acta 16, 411–26.Google Scholar
Santi, G., Marchetti, L., Schirolli, P. & Ronchi, A. 2020. The Cisuralian tetrapod ichnoassociation from Italy: from historical findings to a standard reference status. Journal of Mediterranean Earth Sciences 12, 39–59.Google Scholar
Schneider, J. W., Lucas, S. G., Scholze, F., Voigt, S., Marchetti, L., Klein, H., Opluštil, S., Werneburg, R., Golubevb, V. K., Barrick, J. E., Nemyrovska, Y., Ronchil, A., Day, M. O., Silantiev, V. V., Rößler, R., Saber, H., Linnemann, U., Zharinova, V. & Shen, S.-Z. 2020. Late Paleozoic–early Mesozoic continental biostratigraphy — links to the Standard Global Chronostratigraphic Scale. Palaeoworld 29, 186–238.CrossRefGoogle Scholar
Scotese, C. R. 2014. Atlas of Middle & Late Permian and Triassic Paleogeographic Maps, maps 43–48 from Volume 3 of the PALEOMAP Atlas for ArcGIS (Jurassic and Triassic) and maps 49–52 from Volume 4 of the PALEOMAP Atlas for ArcGIS (Late Paleozoic), Mollweide Projection, PALEOMAP Project: Evanston, Illinois, USA.Google Scholar
Sennikov, A. G. & Golubev, V. K. 2017. Sequence of Permian tetrapod faunas of Eastern Europe and the Permian–Triassic ecological crisis. Paleontological Journal 51, 600–11.CrossRefGoogle Scholar
Sidor, C. A., O'Keefe, F. R., Damiani, R., Steyer, J. S., Smith, R. M. H., Larsson, H. C. E., Sereno, P. C., Ide, O. & Maga, A. 2005. Permian tetrapods from the Sahara show climate-controlled endemism in Pangaea. Nature 434, 886–89.CrossRefGoogle ScholarPubMed
Smith, R. M., Sidor, C. A., Tabor, N. J. & Steyer, J.-S. 2015. Sedimentology and vertebrate taphonomy of the Moradi Formation of northern Niger: a Permian wet desert in the tropics of Pangaea. Palaeogeography, Palaeoclimatology, Palaeoecology 440, 128–41.CrossRefGoogle Scholar
Speksnijder, A. 1985. Anatomy of a strike-slip fault controlled sedimentary basin, Permian of the southern Pyrenees, Spain. Sedimentary Geology 44, 179–223.CrossRefGoogle Scholar
Stappenbeck, R. 1905. Über Stephanospondylus n. g. und Phanerosaurus H. v. Meyer. Zeitschrift der Deutschen Geologischen Gesellschaft 57, 380–437.Google Scholar
Stovall, J. W. 1950. A new cotylosaur from North Central Oklahoma. American Journal of Science 248, 46–54.CrossRefGoogle Scholar
Sullivan, C. & Reisz, R. R. 1999. First record of Seymouria (Vertebrata: Seymouriamorpha) from Early Permian fissure fills at Richards Spur, Oklahoma. Canadian Journal of Earth Sciences 36, 1257–66.CrossRefGoogle Scholar
Sumida, S. S. 1990. Vertebral morphology, alternation of neural spine height, and structure in Permo-Carboniferous tetrapods, and a reappraisal of primitive modes of terrestrial locomotion. University of California Publications in Zoology 122, 1–129 + 3 pls.Google Scholar
Swofford, D. L. 2003. PAUP*. phylogenetic analysis using parsimony (*and other methods). Version 4. Sunderland, Massachusetts: Sinauer Associates.Google Scholar
Tabor, N. J. & Poulsen, C. J. 2008. Palaeoclimate across the Late Pennsylvanian–Early Permian tropical palaeolatitudes: a review of climate indicators, their distribution, and relation to palaeophysiographic climate factors. Palaeogeography, Palaeoclimatology, Palaeoecology 268, 293–310.CrossRefGoogle Scholar
Taquet, P. 1969. Première découverte en Afrique d'un reptile captorhinomorphe (cotylosaurien). Comptes Rendus Hebdomadaires des Séances de l'Academie des Sciences Paris, s. D 268, 779–81.Google Scholar
Vera, J.-A., Alonso-Chaves, F. M., Andreo, B., Arias, C., Azañón, J. M., Balanyá, J. C., Barón, A., Booth-Rea, G., Castro, J. M., Chacón, B., Company, M., Crespo-Blanc, A., Delgado, F., Díaz de Federico, A., Esteras, M., Estévez, A., Fernández, J., Fornós, J. J., Galindo-Zaldívar, J., García-Casco, A., García-Dueñas, V., García-Hernández, M., Garrido, C. J., de Gea, G. A., Gelabert, B., Gervilla, F., González-Lodeiro, F., Jabaloy, A., López-Garrido, A. C., Luján, M., Martín-Algarra, A., Martín-Chivelet, J., Martín-Martín, M., Molina, J. M., Morata, D., Nieto, J. M., Obrador, A., O'Dogherty, L., Orozco, M., Pérez-López, A., Pomar, L., Puga, E., Ramos, E., Rey, J., Rivas, P., Rodríguez-Cañero, R., Ruiz-Cruz, M. D., Ruiz-Ortiz, P. A., Sàbat, F., Sánchez-Gómez, M., Sánchez-Navas, A., Sandoval, J., Sanz de Galdeano, C., Soto, J. I., Torres-Roldán, R. L. & Villas, L. 2004. Capítulo 4. Cordillera Bética y Baleares. In Vera, J.-A. (ed.) Geología de España, 345–464. Madrid: Instituto Tecnológico GeoMinero de España.Google Scholar
Viglietti, P. A., Smith, R. M., Angielczyk, K. D., Kammerer, C. F., Fröbisch, J. & Rubidge, B. S. 2016. The Daptocephalus Assemblage Zone (Lopingian), South Africa: a proposed biostratigraphy based on a new compilation of stratigraphic ranges. Journal of African Earth Sciences 113, 153–64.CrossRefGoogle Scholar
Vjushkov, B. P. & Chudinov, P. K. 1957. Otkrytie Kaptorinid v verkhnnej Permi SSSR [Discovery of a captorhinid in the upper Permian of the USSR]. Doklady Akademii Nauk SSSR 112, 523–36.Google Scholar
Voigt, S. 2005. Die Tetrapodenichnofauna des kontinentalen Oberkarbon und Perm im Thüringer Wald - Ichnotaxonomie, Paläoökologie und Biostratigraphie. Göttingen: Cuvillier Verlag.Google Scholar
Voigt, S., Hminna, A., Saber, H., Schneider, J. W. & Klein, H. 2010. Tetrapod footprints from the uppermost level of the Permian Ikakern Formation (Argana Basin, Western High Atlas, Morocco). Journal of African Earth Sciences 57, 470–78.CrossRefGoogle Scholar
Voigt, S., Lagnaoui, A., Hminna, A., Saber, H. & Schneider, J. W. 2011. Revisional notes on the Permian tetrapod ichnofauna from the Tiddas Basin, central Morocco. Palaeogeography, Palaeoclimatology, Palaeoecology 302, 474–83.CrossRefGoogle Scholar
Voigt, S. & Haubold, H. 2015. Permian tetrapod footprints from the Spanish Pyrenees. Palaeogeography, Palaeoclimatology, Palaeoecology 417, 112–20.CrossRefGoogle Scholar
Voigt, S. & Lucas, S. G. 2015. Permian tetrapod ichnodiversity of the Prehistoric Trackways National Monument (south-central New Mexico, USA). New Mexico Museum of Natural History and Science Bulletin 65, 153–67.Google Scholar
Voigt, S. & Lucas, S. G. 2017. Early Permian tetrapod footprints from central New Mexico. New Mexico Museum of Natural History and Science Bulletin 77, 333–52.Google Scholar
von Meyer, H. 1860. Phanerosaurus Naumanni ans dem Sandstein des Rothliegenden in Deutschland. Paleontographica 7, 248–52 + pl. 27.Google Scholar
White, T. E. 1939. Osteology of Seymouria baylorensis Broili. Bulletin of the Museum of Comparative Zoölogy 85, 323–409 + 3 pls.Google Scholar
Zouicha, A., Voigt, S., Saber, H., Marchetti, L., Hminna, A., El Attari, A., Ronchi, A. & Schneider, J. W. 2021. First record of Permian continental trace fossils in the Jebilet Massif, Morocco. Journal of African Earth Sciences 173, 104015.CrossRefGoogle Scholar
Supplementary material: PDF

Matamales-Andreu et al. supplementary material

Matamales-Andreu et al. supplementary material 1

Download Matamales-Andreu et al. supplementary material(PDF)
PDF 46.2 MB
Supplementary material: File

Matamales-Andreu et al. supplementary material

Matamales-Andreu et al. supplementary material 2

Download Matamales-Andreu et al. supplementary material(File)
File 32.5 KB