Hostname: page-component-586b7cd67f-gb8f7 Total loading time: 0 Render date: 2024-11-28T04:11:47.024Z Has data issue: false hasContentIssue false
  • English
  • Français

REVISION OF THE NEARCTIC SPECIES OF TRICHOCLINOCERA COLLIN (DIPTERA: EMPIDIDAE; CLINOCERINAE)

Published online by Cambridge University Press:  31 May 2012

Bradley J. Sinclair
Bradley J. Sinclair
Affiliation:
Department of Biology, Carleton University, Ottawa, Ontario, Canada K1S 5B6
Get access Rights & Permissions [Opens in a new window]

Abstract

There are 16 species of Trichoclinocera Collin recognized from the Nearctic region, including eight new species. Four of six species-groups of Trichoclinocera are recognized in the Nearctic region, with the following included species: (1) T. comata group — T. cascadensis sp.nov. (type-locality Starvation Creek, Oregon, U.S.A.), T. comata (Melander) comb.nov., and T. rupestris sp.nov. (type-locality Newton Creek, Wyoming, U.S.A.); (2) T. dolicheretma group — T. dolicheretma (Melander); (3) T. lapponica group — T. lapponica (Ringdahl) comb.nov.; (4) T. longipes group — T. agilis sp.nov. (type-locality Fall Creek Falls, Tennessee, U.S.A.), T. ctenistes (Melander), T. dasycoxa sp.nov. (type-locality Ottawa, Ontario, Canada), T. falcata sp.nov. (type-locality Gatineau Park, Quebec, Canada), T. fumosa (Vaillant) comb.nov., T. hamifera (Melander), T. longipes (Walker). T. minor (Melander), T. ozarkensis sp.nov. (type-locality Haw Creek Falls, Arkansas, U.S.A.), and T. pectinifemur sp.nov. (type-locality Phoenixville, Pennsylvania, U.S.A.). The following new synonyms are proposed: T. brunnipennis (Melander) = T. dolicheretma; and Seguyella saltans (Vaillant) = T. hamifera. Lectotypes are designated for T. comata, T. ctenistes, T. dolicheretma, T. brunnipennis, T. hamifera, T. lapponica, and T. stackelbergi Collin. Seguyella Vaillant and Acanthoclinocera Saigusa are newly synonymized with Trichoclinocera and all species formerly placed in these genera are transferred to Trichoclinocera, except Acanthoclinocera glaucescens (Brunetti) which is transferred as Wiedemannia glaucescens (Brunetti) comb.nov.This revision includes a key to Nearctic species, a description of each species, and maps of species distributions. Homologies of male terminalia of Clinocerinae are summarized and the first descriptions of the immature stages of species of Trichoclinocera also are provided. A world list of described species of Trichoclinocera is provided and all species are assigned to one of six species-groups, except T. cummingi sp.nov. (type-locality Robber’s Cave, Oklahoma. U.S.A.), T. fluviatilis (Brunetti) comb.nov., T. fuscipennis Saigusa, and T. maewa Smith, which remain unplaced. The monophyly of Trichoclinocera is justified and phylogenetic relationships to the remaining Clinocerinae and among species-groups are discussed.

Résumé

Le genre Trichoclinocera Collin compte 16 espèces dans la région néarctique et huit d’entre elles sont nouvelles. Quatre des six groupes d’espèces sont représentés dans la région néarctique : (1) le groupe T. comata — T. cascadensis sp.nov. (localité-type, Starvation Creek, Oregon, É.-U.), T. comata (Melander) comb.nov. et T. rupestris sp.nov. (localité-type, Newton Creek, Wyoming, É.-U.), (2) le groupe T. dolicheretma — T. dolicheretma (Melander), (3) le groupe T. lapponica — T. lapponica (Ringdahl) comb.nov., (4) le groupe T. longipes — T. agilis sp.nov. (localité-type, Fall Creek Falls, Tennessee, É.-U.). T. ctenistes (Melander), T. dasycoxa sp.nov. (localité-type, Ottawa, Ontario, Canada), T. falcata sp.nov. (localité-type, Parc de la Gatineau, Québec, Canada), T. fumosa (Vaillant) comb.nov., T. hamifera (Melander), T. longipes (Walker), T. minor (Melander), T. ozarkensis sp.nov. (localité-type, Haw Creek Falls, Arkansas, É.-U.) et T. pectinifemur sp.nov. (localité-type, Phoenixville, Pennsylvanie, É.-U.). Les synonymes suivants sont proposés : T. brunnipennis (Melander) = T. dolicheretma et Seguyella saltans (Vaillant) = T. hamifera. Des lectotypes de T. comata, T. ctenistes, T. dolicheretma, T. brunnipennis, T. hamifera, T. lapponica et T. stackelbergi Collin ont été désignés. Seguyella Vaillant et Acanthoclinocera Saigusa deviennent synonymes de Trichoclinocera et toutes les espèces de ces genres sont transférées au genre Trichoclinocera, sauf Acanthoclinocera glaucescens (Brunetti), qui devient Wiedemannia glaucescens (Brunetti) comb.nov.On trouvera dans cette révision une clé des espèces néarctiques ainsi qu’une description et une carte de répartition de chacune des espèces. Les homologies des organes terminaux des mâles sont exposées brièvement et des descriptions encore inédites des stades immatures des espèces de Trichoclinocera sont présentées. Une liste des espèces de Trichoclinocera à l’échelle mondiale est dressée et toutes les espèces sont assignées à l’un des six groupes, à l’exception de T. cummingi sp.nov. (localité-type, Robber’s Cave, Oklahoma, É.-U.), T. fuscipennis Saigusa et T. maewa Smith, dont la position n’a pas été déterminée. La nature monophylétique du genre Trichoclinocera est démontrée et les relations phylogénétiques entre ce genre et les autres Clinocerinae et entre les groupes d’espèces sont examinées.

[Traduit par la Rédaction]

Type
Articles
Information
The Canadian Entomologist , Volume 126 , Issue 4 , August 1994 , pp. 1007 - 1059
Copyright
Copyright © Entomological Society of Canada 1994

Access options

Get access to the full version of this content by using one of the access options below. (Log in options will check for institutional or personal access. Content may require purchase if you do not have access.)

References

REFERENCES

Allen, R.T. 1990. Insect endemism in the Interior Highlands of North America. Florida Entomologist 73(4): 539–569.CrossRefGoogle Scholar
Bährmann, R. 1960. Vergleichend-morphologische Untersuchungen der männlichen Kopulationsorgane bei Empididen. Beiträge zur Entomologie 10(5/6): 485–540.Google Scholar
Brindle, A. 1969. Taxonomic notes on the larvae of British Diptera. The Entomologist 102: 35–39.Google Scholar
Brindle, A. 1973. Taxonomic notes on the larvae of British Diptera. The Entomologist 106: 249–252.Google Scholar
Brunetti, E. 1913. New Indian Empidae. Records of the Indian Museum 9: 11–45.Google Scholar
Chvála, M. 1983. The Empidoidea (Diptera) of Fennoscandia and Denmark. II. General Part. The families Hybotidae, Atelestidae and Microphoridae. Fauna Entomologica Scandinavica 12: 1–279.Google Scholar
Chvála, M., and Wagner, R.. 1989. Empididae. pp. 228–336 in Soós, A., and Papp, L. (Eds.), Catalogue of Palearctic Diptera. Vol. 6. Therevidae – Empididae. Elsevier Science Publ., Amsterdam. 435 pp.Google Scholar
Collin, J.E. 1941. Some Pipunculidae and Empididae from the Ussuri region on the far eastern border of the U.S.S.R. (Diptera). Proceedings of the Entomological Society of London (B) 10: 218–248.Google Scholar
Collin, J.E. 1961. Empididae. In British Flies. Vol. 6. University Press, Cambridge. 782 pp.Google Scholar
Cumming, J.M., and Cooper, B.E.. 1992. A revision of the Nearctic species of the tachydromiine fly genus Stilpon Loew (Diptera: Empidoidea). The Canadian Entomologist 124: 951–998.CrossRefGoogle Scholar
Cumming, J.M., and Sinclair, B.J.. 1990. Fusion and confusion: Interpretation of male genitalic homologies in the Empidoidea (Diptera). Second International Congress of Dipterology, Bratislava, Abstract Volume, Third Supplement, p. 334.Google Scholar
Delfinado, M.D., and Hardy, D.E. (Eds.). 1973. A Catalog of the Diptera of the Oriental Region. Vol. 1. The University Press of Hawaii, Honolulu. 618 pp.CrossRefGoogle Scholar
Griffiths, G.C.D. 1972. The phylogenetic classification of Diptera Cyclorrhapha, with special reference to the structure of the male postabdomen. Series Entomologica 8: 1–340.Google Scholar
Griffiths, G.C.D. 1980. Preface. pp. v–vii in Griffiths, G.C.D. (Ed.), Flies of the Nearctic Region. Vol. 1. Handbook, Part 1. History of Nearctic Dipterology. E. Schweizerbart'sche Verlagsbuchandlung, Stuttgart. 62 pp.Google Scholar
Kelsey, L.P. 1969. A revision of the Scenopinidae (Diptera) of the World. United States National Museum Bulletin 277: 1–336.Google Scholar
McAlpine, J.F. 1981. Morphology and terminology — Adults [Chapter] 2. pp. 9–63 in McAlpine, J.F. et al. , (Coords.), Manual of Nearctic Diptera. Vol. 1. Agriculture Canada Monograph 27: 1–674.Google Scholar
Melander, A.L. 1902. A monograph of the American Empididae. Part 1. Transactions of the American Entomological Society 28: 195–368.Google Scholar
Melander, A.L. 1928 (1927). Diptera, Fam. Empididae. In Wytsman, P. (Ed.), Genera Insectorum. Fasc. 185. Louis Desmet-Verteneuil, Bruxelles. 434 pp.Google Scholar
Melander, A.L. 1965. Family Empididae (Empidae, Hybotidae). pp. 446–481 in Stone, A. et al. , (Eds.), A Catalog of the Diptera of America North of Mexico. United States Department of Agriculture Handbook 276: 1–1696.Google Scholar
O'Hara, J.E. 1983. (1982). Classification, phylogeny and zoogeography of the North American species of Siphona Meigen (Diptera: Tachinidae). Quaestiones Entomologicae 18: 261–380.Google Scholar
Osten Sacken, C.R. 1878. Catalogue of the described species of Diptera of North America. Smithsonian Miscellaneous Collections 270: 1–276.Google Scholar
Ricker, W.E. 1964. Distribution of Canadian stoneflies. Gewässer und Aswässer 34/35: 50–71.Google Scholar
Ringdahl, O. 1933. Översikt avde i Sverige funna arterna av släktet Atalanta Meig. (Clinocera Meig.) inom fam. Empididae. Entomologisk Tidskrift 54: 277–282.Google Scholar
Rogers, E. 1981. Mating in Wiedemannia (Diptera: Empididae). Journal of Natural History 15: 1013–1018.CrossRefGoogle Scholar
Sabrosky, C.W. 1967. Corrections to A Catalog of the Diptera of America North of Mexico. Bulletin of the Entomological Society of America 13: 115–125.CrossRefGoogle Scholar
Saigusa, T. 1965. Two new species of Clinocerinae from Japan (Diptera, Empididae). Kontyû 33(1): 53–57.Google Scholar
Sinclair, B.J. 1993. Phylogenetic classification of the Clinocerinae, with a Revision of the Nearctic Species of Trichoclinocera Collin and Wiedemannia Zetterstedt (Diptera: Empididae). Ph.D. thesis, Carleton University, Ottawa, Ont.489 pp.Google Scholar
Sinclair, B.J., Cumming, J.M., and Wood, D.M.. 1994. (1993). Homology and phylogenetic implications of male genitalia in Diptera — Lower Brachycera. Entomologica Scandinavica 24(4): 407–423.CrossRefGoogle Scholar
Smith, K.G.V. 1965. Diptera from Nepal. Empididae. Bulletin of the British Museum (Natural History), Entomology 17: 61–112.CrossRefGoogle Scholar
Smith, K.G.V. 1971. A revision of Francis Walker's types of North American Empididae (Diptera). Bulletin of the British Museum (Natural History), Entomology 26(8): 345–370.Google Scholar
Smith, K.G.V. 1975. Family Empididae (Empidae, Hybotidae). pp. 185–211 in Delfinado, M.D., and Hardy, D. E. (Eds.), A Catalog of the Diptera of the Oriental Region. Vol. 2. The University Press of Hawaii, Honolulu. 459 pp.Google Scholar
Soós, A., and Papp, L. (Eds.). 1984. Catalogue of Palearctic Diptera. Vol. 9. Micropezidae–Agromyzidae. Elsevier Science Publ., Amsterdam. 460 pp.Google Scholar
Steyskal, G.C., and Knutson, L.V.. 1981. Empididae [Chapter] 47. pp. 607–624 in McAlpine, J.F. et al. , (Coords.), Manual of Nearctic Diptera. Vol. 1. Agriculture Canada Monograph 27: 1–674.Google Scholar
Teskey, H.J. 1981. Morphology and terminology — Larvae [Chapter] 3. pp. 65–88 in McAlpine, J.F. et al. , (Coords.), Manual of Nearctic Diptera. Vol. 1. Agriculture Canada Monograph 27: 1–674.Google Scholar
Teskey, H.J. 1984. Aquatic Diptera Part One. Larvae of aquatic Diptera. pp. 448–466 in Merritt, R.W., and Cummins, K.W. (Eds.), An Introduction to the Aquatic Insects of North America, 2nd ed. Kendal/Hunt Publ. Co., Dubuque, IA.722 pp.Google Scholar
Ulrich, H. 1971. Zur Skelett- und Muskelanatomie des Thorax der Dolichopodiden und Empididen (Diptera). Veröffentlichungen der Zoologischen Staatssammlung, München 15: 1–44.Google Scholar
Ulrich, H. 1972. Zur Anatomie des Empididen-Hypopygiums (Diptera). Veröffentlichungen der Zoologischen Staatssammlung, München 16: 1–28.Google Scholar
Ulrich, H. 1975. Das Hypopygium von Chelifera precabunda Collin (Diptera, Empididae). Bonner Zoologische Beiträge 26: 264–279.Google Scholar
Vaillant, F. 1952 a (1951). Un Empidide destructeur de simulies. Bulletin de la Société Zoologique de France 76: 371–379. [Though the journal cover bears the date 1951. the issue containing this article was printed in 1952.]Google Scholar
Vaillant, F. 1952 b. Les larves hygropétriques et les nymphes de quelques Empididae Atalantinae (Diptères). Travaux du Laboratoire d'Hydrobiologie et de Pisciculture d'Université de Grenoble 43–44: 39–46.Google Scholar
Vaillant, F. 1960 a. Quelques Empididae Atalantinae des monts des Appalaches [Dipt.]. Bulletin de la Société entomologique de France 65: 117–123.CrossRefGoogle Scholar
Vaillant, F. 1960 b. Quelques Empididae Atalantinae d'Asie russe [Dipt.]. Bulletin de la Société entomologique de France 65: 170–186.CrossRefGoogle Scholar
Vaillant, F. 1965. (1964). Revision des Empididae Hemerodromiinae de France, d'Espagne et d'Afrique du Nord. [Dipt.]. Annales de la Société entomologique de France 133: 143–173. [Date of publication: January 15, 1965.]Google Scholar
Vaillant, F., and Vinçon, G.. 1987. (1986). Quelques Clinocerini (Diptera, Empididae, Hemerodromiinae) nouveaux ou mal connus des Pyrénées. Annals de Limnologie 22: 261–275. [Though the journal cover bears the date 1986. the issue containing this article was printed in 1987.]CrossRefGoogle Scholar
Wagner, R. 1990. Neue Wiedemannia-Arten aus der rhynchops-Gruppe (Diptera, Empididae, Clinocerinae). Entomofauna 11(13): 229–237.Google Scholar
Walker, F. 1849. List of the specimens of Dipterous Insects in the Collection of the British Museum. Part III. pp. 485–687.Google Scholar
Wiley, E.O. 1981. Phylogenetics. The Theory and Practice of Phylogenetic Systematics. John Wiley and Sons, New York, NY. 439 pp.Google Scholar