In the course of my already-published studies, the probabilities as to the homology of the last anal vein of the Papilionides primary have varied. From preparations of the pupal wing of the Hesperiades, it has become clear that the fork to second anal at base is the remains of the third anal vein, which is irregular and more extended in the fore wing of the chrysalis and does not attain the outer margin. It is furcate aud connected with the second anal in the pupal stage. On the other hand, the downwardly curved, short, last and free anal vein of the Papilionid primary cannot be homologous with this, as, indeed, I originally contended. This vein reaches the internai margin, and is probably a survival of the fourth anal.