With a diverse and colourful cast of animals, birds, insects and human villagers, Příhody lišky Bystroušky (known in English as The Cunning Little Vixen), is one of Leoš Janáček's most popular, if peculiar, operas. Though nowadays Bystrouška is typically characterised as a charming portrayal of the continuous renewal of life in nature, this idea emerged only gradually from a tangle of competing and occasionally contradictory views in which, however, the complexity of the moral laws by which the opera's inhabitants live was frequently central. By tracing the history of the opera's stagings in the Czech Republic; drawing out themes that developed in the journalistic and critical discourse around those performances; and reading the opera's music and stage action closely, this article argues that the amoral codes of Bystrouška's world not only inhere in the story, its text, and even, perhaps, in the idea of nature's cycle of life itself, but that at certain times in the opera they are also given expression through particular correlations and disjunctions between the opera's music and the physical actions and gestures of the singers on stage.