Raphe-like vestiges have been found regularly in two new taxa which presumably belong to Fragilaria, one of them livingin several lakes of the Kerguelen Islands, the other in the Yellowstone National Park, Wyoming, U.S.A. The latter onepossesses septa-like elements as integrated parts of a girdle band. Other septa-like elements can occur in different speciesof Fragilaria. They represent appendices of the valvocopula, located beneath the transapical costae. Structures like linkingspines on marginal parts of the valve face occur in an Achnanthes species. Most of these characters seem to havebeen unknown in these taxa until recently.