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Published online by Cambridge University Press: 01 July 2016
The imago of the peppered moth exists in two sharply distinguished forms, speckled and black. Clarke and Sheppard (1966) have shown that the frequency of the black moth decreases from 99% in Liverpool to 10% in North Wales. If these frequencies are in equilibrium, albeit dynamic equilibrium, then such a gradient is called a cline. Haldane (1948) briefly describes several clines, and then defines a model that may represent the life-cycle of certain species. The assumptions defining his model are as follows.