In the preceding chapters, we showed why the idea that living organisms are really driven by their genes is a profound misunderstanding of how living systems work. On the contrary, they are open systems at all levels of organisation. How things work at the molecular level is constrained and regulated at the cellular level. The interaction of cells is regulated at the tissue level, and tissues at the organ level, and organs at the system level. The system is regulated by the behaviour of the organisms, and organisms by social and ecological interactions. The psychosocial level is unique. If there is a privileged level of causation, then it lies at the psychosocial level and not at the level of genes. This is the level at which wilful agency is initiated and organisms can be genuinely selfish or altruistic. In truth you cannot be selfish if you do not have the choice to be altruistic, which is why selfishness cannot be applied at a genetic level, neither metaphorically nor literally.