A microbioherm-bearing hardground within the middle Silurian (Wenlock) Massie Formation near Napoleon, southeastern Indiana, United States is encrusted by the attachment structures of numerous pelmatozoan echinoderms. Among the most common of these holdfasts are multi-plated discoidal structures representing the thecal attachments of diploporite ‘cystoids’. This large population of holdfasts permits the first detailed taphonomic and palaeoecologic study of hardground diploporite attachments, allowing for increased morphological understanding of these rarely studied structures and facilitating identification of holdfasts in deposits where they might have been overlooked or misidentified. The biostratinomic sequence commences with detachment of thecae, followed by weathering of isolated discoidal holdfasts to bring out radiating canal structures and plate sutures, eventually leading to removal of the interior floor to expose the underlying substrate. Continued exposure can result in separation of component holdfast plates, though cementation to the substrate prevents scattering of plates. Diagenetic precipitation of pyrite occurred after burial; the large size of crystals suggests late diagenesis, perhaps seeded by early diagenetic pyrite crystallites produced by decay of ligamentary tissue. Extrinsic taphonomic factors include overgrowth of holdfasts by laminar stenolaemate bryozoans and other echinoderm attachment structures. Diploporite holdfasts are not bored and are absent on microbioherms. Taphonomic data indicate the time-averaged nature of this hardground and its diploporite assemblage and permit prediction of similar occurrences at major flooding surfaces.

The upper Wenlock Series (Homerian Stage) of the northern Midland Platform, England, contains numerous volcanic bentonite clay layers. At Wren's Nest Hill, Dudley, 15 bentonites have been investigated and comparisons with the type-Wenlock have been made by means of two key sections along Wenlock Edge, Shropshire. In total 22 bentonites have been investigated and their clay and sand-grade mineralogies determined. Rare earth element (REE) and yttrium concentrations of apatite grains contained within ten of the bentonites have been established allowing geochemical fingerprinting as an indication of provenance of source magmas and identification of geochemical marker beds. Based on the analysis of REE and yttrium concentrations it seems likely that the majority of these bentonites originated from a granodiorite magmatic source. Comparisons with published Llandovery and lower Wenlock age bentonites indicate generally more enrichment in light REEs relative to heavy REEs. In addition, close geochemical similarities between bentonites along Wenlock Edge and at Wren's Nest Hill strongly argue for their presence as precise stratigraphic equivalents within the upper Much Wenlock Limestone Formation. These correlations are further supported by geophysical data from borehole wire-line logs across the West Midlands. Finally, a chemically distinct mid-Homerian episode of volcanism is identified and represents a potentially important marker interval between the study area and other similarly aged bentonites reported from the Island of Gotland, Sweden.

Sanidine composition and bulk geochemistry of volcanic ash beds from the East Baltic indicate the subalkaline nature of the volcanism near the margins of the Baltica plate during the Silurian. Several bentonites in the Wenlock include a previously unknown sanidine with 48 to 58 mol % of the Na+Ca component. In contrast to the earlier Telychian volcanism, sodium-rich sanidine occurs in ash beds which originate from relatively moderately evolved dacitic magma. The studied material from two drill cores integrated with previous research enables production of a more complete list of 49 volcanic eruption layers for the lower to middle Wenlock in the East Baltic. This updated list of bentonites characterized by their sanidine compositions forms a good basis for future integrated bio- and chemostratigraphic correlations in northern Europe.

Integrated graptolite, conodont and chitinozoan biostratigraphical data are presented from the Llandovery and Wenlock of the Kolka-54 core, Latvia. Correlations between graptolite and chitinozoan biozones are consistent with those published from other East Baltic sections and the Welsh Basin. While most correlations between graptolite and conodont biozones agree with those presented in previous studies, there are important exceptions. Significantly, we report here the discovery of Distomodus staurognathoides Biozone conodonts in the lowest Aeronian Demirastrites triangulatus graptolite Biozone. The base of the D. staurognathoides Biozone was previously considered to lie much higher in the Aeronian. Also it is shown that Walliserodus survived the late Wenlock Mulde Event, during which it was considered previously to have become extinct.

δ13Corg and TOC data are presented from the upper spiralis Biozone (Telychian, Llandovery, Silurian) through to the upper Sheinwoodian (Wenlock, Silurian) of the Banwy River section, Wales. In laminated hemipelagites from the Telychian, δ13Corg values rise through the upper lapworthi Biozone to a maximum in the lower insectus Biozone after which they decline slightly. The most conspicuous feature of the δ13Corg curve is the prolonged positive excursion in the Sheinwoodian, commencing in the upper murchisoni Biozone and ending in strata yielding Monograptus flexilis. This Sheinwoodian positive δ13C excursion in the Banwy River section correlates precisely with that recognized in the East Baltic. The interval with the highest δ13Corg values also records the highest TOC values, suggesting that for the Sheinwoodian at least burial of carbon may have contributed to the positive δ13C excursion. Bioturbated strata yield very low TOC values; whether the δ13Corg values from these beds reflect a primary signal or the result of biostratinomic or diagenetic modification is uncertain.

Forty-six samples, taken from the Upper Ordovician and lower Silurian strata of the Girvan district, Midland Valley of Scotland, yield moderately well-preserved and diverse assemblages of chitinozoans. The area was chosen to study the composition of the chitinozoan assemblages, rarely described before in Scotland, which lay at the south-eastern margin of the Laurentia palaeocontinent. These assemblages are compared with those of the same age on other palaeocontinents, representing a critical time when both the Iapetus and Tornquist oceans were closing, thus forming the Caledonide and associated orogens. It is possible to correlate the chitinozoan occurrences at Girvan with those of other parts of Laurentia (Quebec area), other palaeocontinents such as Baltoscandia (Sweden, Estonia), Avalonia (Wales, Brabant Massif) and some parts of Northern Gondwana (Saudi Arabia). For the lower Silurian, it is possible to correlate with the global chitinozoan biozonation, and a calibration between the chitinozoan occurrences and the graptolite biozonation of Girvan is established. Several levels within the Balclatchie, South Shore, Three Mile, Shalloch, Wood Burn, Lauchlan, Drumyork Flags and other formations are accurately dated.

Twenty six of the more important species of monograptids (s.l.), retiolitids and Cyrtograptus (from a total fauna of 52 species) are described from Wenlock strata of the Cape Phillips Formation, Canadian Arctic Archipelago. Of this fauna, eight new species or subspecies, Monograptus firmus festinolatus, M. instrenuus, M. opimus, M. testis incomptus, Cyrtograptus falcatus, C. hamatus brevis, C. kolobus and C. pseudomancki, are described and illustrated.

Wenlock biostratigraphic zones comprise the Cyrtograptus centrifugus-C. insectus Zone (earliest Wenlock), M. instrenuus-C. kolobus Zone, tentatively divisible into lower and upper subzones, C. perneri-M. opimus Zone possibly divisible into lower and upper subzones, C. lundgreni-M. testis Zone divisible into a lower M. testis incomptus Subzone and an upper M. testis testis Subzone, and the Pristiograptus ludensis Zone (latest Wenlock).

The history of investigation of the Silurian rocks of the Dunquin inlier, Dingle Peninsula, County Kerry, Ireland is briefly reviewed. The Dunquin Group of upper Wenlock (Homerian) to middle Ludlow (Ludfordian) sedimentary and volcanic rocks is divided into five formations: the Ferriters Cove, Clogher Head, Mill Cove, Drom Point and Croaghmarhin Formations, the last of which reaches the Ludlow and passes transitionally into the succeeding continental Dingle Group, probably largely Přídolí in age. The Dunquin group is of varied shallow marine sediments with a continental episode probably represented in the Mill Cove Formation. The biostratigraphy of the other four formations is considered in relation to a full faunal list. Details of representative sections and exposures are provided. There is brief comment on the palaeogeographical setting of the inlier.