A flock of Cheviot ewes carrying the Thoka gene for fecundity was established in southern Scotland through the importation of frozen semen from two Icelandic rams. Ewes considered to be probable carriers of the Thoka gene were identified on the basis of measurements of ovulation rate as juveniles at 1·5 years of age and again as adults. Analysis of 3 years' records of 8926 Cheviot lambings and lambings of 351 Icelandic × Cheviot ewes considered to be probable carriers showed that fecundity increased with age in both genotypes, the number of lambs born per ewe mated being 1·07, 1·19, 1·31 and 1·41 in the Cheviots and 1·55, 1·74, 2·33 and 2·30 in the Icelandic-Cheviot crosses at first to fourth lambings respectively. The apparent effect of the crossbreeding on fecundity thus also increased with age, values at first to fourth lambings being 0·48, 0·55, 1·02 and 0·89 lambs per ewe mated, giving a weighted mean value of 0·61 (1·20 v. 1·81 lambs per ewe mated in Cheviot and Icelandic × Cheviot ewes respectively). There was a positive effect of the proportion of Icelandic ancestry on the number of lambs born per ewe mated and thus the observed increases in fecundity, although attributed principally to the effects of the Thoka gene, are also due in part to other genetic factors introduced from the Icelandic sheep in the crossbreeding programme. Lamb mortality to 6 weeks of age was only marginally higher in Icelandic × Cheviots (0·10) than in Cheviots (0·08). The increase in fecundity was achieved without an excessive proportion of large litters (<0·02 of probable carriers gave birth to quadruplets). The results demonstrate that the introduction of the Thoka gene to breeds such as the Cheviot can increase fecundity and hence the potential for more efficient lamb production.

One gilt from each of 24 littermate trios was assigned at birth to one of three treatments which were to adjust litter size to six, nine or 12 piglets. Plasma luteinizing hormone (LH) and follicle-stimulating hormone (FSH) were measured before and after stimulation with 1 ng gonadotrophin-releasing hormone (GnRH) per kg body weight at 55 and 140 days of age. At 150 days, oestrogen was administered and ovarian and uterine responses were measured at slaughter on day 160.

Litter size significantly modified pre-weaning growth rate (P < 0·01) but did not influence gonadotrophin secretion or sexual maturity at slaughter. The family of origin significantly influenced ovarian and uterine development (P < 0·05) and the LH and FSH responses to GnRH at 55 days (P < 0·05). Family size (P < 0·01) and sex ratio of the litter at birth (P < 0·05) also influenced the ovarian response to oestrogen treatment. Significant sire effects on ovarian and uterine development were established (P < 0·05); for the progeny of individual boars significant differences in the response to oestrogen were associated with significant differences in growth rate (P < 0·001).

Variations in family birth and litter weaning weights were negatively correlated with ovarian and uterine development (P < 0·05), whereas all measures of growth from weaning to slaughter were significantly and positively correlated with sexual maturity at slaughter (P < 0·05).

Basal FSH and the responses of both LH and FSH to GnRH at 55 days were correlated with growth rate at day 55 (P < 0·05). Basal LH and FSH at day 140 were correlated with ovarian and uterine development at day 160 (P < 0·05).

The effects of two levels of energy intake, ad libitum and 80% of ad libitum, of diethylstilboestrol implantation and no such treatment, and of two slaughter weights (‘450’ and ‘490’ kg), on the performance of Israeli-Friesian intact male cattle were studied.

Average daily gain was 1130 g and 959 g and daily carcass gain was 630 g and 553 g for animals on ad libitum and 80% of ad libitum energy intake, respectively. Average daily gain was 1038 g and 944 g and daily carcass gain was 595 g and 562 g, for diethylstilboestrol-treated and untreated animals, respectively. Slaughter weight had little effect on rate of gain.

The differences in conversion ratio of ME into live weight between treatments were small. Diethylstilboestrol-treated animals were slightly more efficient.

The non-implanted animals on the restricted plane of nutrition and which had been slaughtered at ‘490’ kg had a significantly higher dressing percentage, a higher percentage of fat trim and less bone than animals slaughtered at ‘450’ kg body weight on both levels of nutrition.

Among the diethylstilboestrol-implanted male calves, however, the animals fed at 80% of ad libitum feed and killed at ‘450’ or ‘490’ kg live weight had significantly more fat trim in the carcass and less bone than the animals fed ad libitum.

Annual surveys of fertilizer practice have been conducted since 1942. Since 1985, nitrogen application rates to grass have progressively declined and phosphorus application has been reduced since the mid 1970s. Increasing environmental issues and the present interest in organic farming and low input systems indicate that these trends will continue. Present overall fertilizer use for grazing on dairy farms is about 170 kg N, 10 kg P and 20 kg К per ha. Higher rates are used for intensive silage production, 200 kg N, 15 kg P and 50 kg К per ha. In contrast, recommended applications are 340 kg N, 18 kg P and 25 kg К per ha for grazing and 380 kg N, 40 kg P and 260 kg К per ha for intensive silage. Herbage yield is controlled by the amount and timing of nitrogen fertilizer applications. The major mineral content depends on herbage maturity within the growth cycle; both phosphorus and potassium contents fall with declining crude protein concentrations.

Nitrogen fertilizer generally increases both magnesium and sodium concentration whilst potassium application decreases both. The overall effect of combined applications is to minimize changes. In the critical spring and autumn periods, herbage dry-matter intakes may be as important as magnesium content and availability when assessing the risk of hypomagnesaemia in the grazing cow.

Fertilizers, including phosphorus, even in the long term, have little effect on the calcium and phosphorus contents of herbage. Nitrogen and phosphorus concentrations are significantly ocrrelated in grazed herbage, silage and hay. If environmental or economic policies were to result in a reduction in the crude protein content of grazed herbage from ca. 250 to ca. 200 g/kg dry matter, the phosphorus intake of a cow giving 30 kg milk per day would fall by 0·5 g/kg dry-matter intake. This, together with any desirable additional magnesium, should preferably be given as an oral mineral supplement rather than by attempting to modify the mineral content of herbage.

The food conversion and meat quality of 10 Aberdeen Angus and 10 Charolais crossbred heifers, about 18 to 20 months of age, was measured. The animals were finished using diet of hay and concentrates for up to 10 weeks prior to slaughter at either fat class or 4L using the Meat and Livestock Commission scheme. There were no significant breed differences in food conversion. Post slaughter, samples of muscle from the L. dorsi over the 11 to 12th rib from all heifers were used for sensory evaluation using a trained taste panel. The muscle from heifers in fat class 4L showed an increased flavour intensity compared with those in fat class 3 (P < 0·05). No other differences in sensory characteristics were found. The measurement of muscle fibre areas in the L. dorsi revealed no significant differences, suggesting a similarity in the texture of the meat from the breeds when slaughtered at the same age.

Three wilted silages (dry matter concentration of approximately 300 g/kg) were prepared from early-bloom lucerne which received no additive (MG-0), or was treated with a mixture of carboxylic salts (Maxgrass) at either 4 (MG-4) or 8 (MG-8) l/t fresh crop. Silages were stored in tower silos. Resulting silages were offered ad libitum to growing Holstein steers without supplementation. Untreated silage (MG-0) exhibited an extensive, predominantly lactic acid fermentation. The nitrogen (N) fraction was highly soluble, relative to the crop at ensiling. Silage fermentation and protein solublization were restricted by Maxgrass application. Maxgrass application reduced aerobic stability of silage removed from the upper third of silos but not of silage from the lower portion of silos. Apparent digestibility showed a quadratic response to level of Maxgrass application (P < 0·05). Voluntary intake was not affected by Maxgrass addition (P > 0·005) but intake of all silages was high (30 g/kg live weight (LW)). There was a positive linear response (P < 0·05) in LW gain to Maxgrass application with gains of 0·74, 0·86 and 0·87 kg/day being achieved in steers given MG-0, MG-4 and MG-8 silages, respectively. Degradability of silage N determined in nylon bags in situ was unaffected by Maxgrass application. However, the immediately degradable N fraction was reduced by Maxgrass application (linear effect, P < 0·001; quadratic effect, P < 0·05). Benefits in animal performance due to Maxgrass application were attributed to improved N composition while restricted carbohydrate fermentation during ensiling was considered to be of secondary importance.

The use of an artificial rearing system to provide controlled environment conditions for performance testing of male lambs was investigated in two trials. A sample of lambs from 5 fat-lamb sire breeds was included.

Performance during the initial restricted milk-feeding period was uniform but large differences in solid food intake gave variable performance in the adjustment period immediately following weaning. Lambs were evaluated from 40 days onwards in terms of growth rate and efficiency of food conversion. Some breed differences were significant and the variation within groups was sufficiently high to suggest that performance might be improved by within-group selection.

Estimates from Trial 2 show that food conversion efficiency was related to growth rate, but early and late performance in the evaluation period were not correlated (r = 0·12, 0·16 and 0·14 for growth rate, food intake and efficiency, respectively). Some refinements of the method are suggested, and it is concluded that the system could provide suitable conditions for testing male lambs at an early age without the variable influence of the dam.

Eight-hundred-and-fifty-two pigs were weaned at 5 weeks of age, and at an average weight of 8·5 kg were allocated in pens of 14 or 15 to diets having 13·6 or 14·6 MJ digestible energy (DE)/kg combined with 11·0 or 11·8 g lysine per kg.

Over a 5-week experimental period, pigs receiving the diet with the higher DE content had a lower daily feed intake, improved feed conversion efficiency (FCE) but a similar rate of gain to those getting less DE/kg. Efficiency of utilization of energy was poorer when DE content was higher.

Increasing lysine level increased daily feed intake and rate of gain while improving FCE. There were significant energy x lysine interactions. At the lower energy density, increasing the lysine level resulted in increased daily feed intakes whereas the reverse was the case at the higher energy density. Daily gain increased when lysine level was increased at the lower energy density but was unaffected at the higher density. Efficiency of energy utilization was improved by increasing lysine at the higher but not at the lower energy level.

Growth from two years to three years and carcass yield and composition were evaluated in nine types of steers produced by mating Angus, Boran and Red Poll bulls to Ankole, Boran and local Zebu cows. One hundred and nine steers were slaughtered at the average age of 3 years and 1 month.

At three years of age, steers by Angus and Red Poll sires were 7% heavier than steers by Boran sires, while progeny of Boran and Ankole dams were 14% heavier than progeny of Zebu dams. Significant hybrid vigour effects on weight-for-age were found.

After adjustment for weight and age at slaughter, linear carcass measurements supported the impression that steers by Angus sires were more compact and blocky than steers by Red Poll and Boran sires and that progeny of Ankole dams were longer in the leg and body than progeny of Boran and Zebu dams. Steers by Red Poll sires appeared to have a higher fat content than steers by Angus and Boran sires on the basis of dissection of a tenth rib sample joint. Progeny of Boran dams had a higher dressing percentage than progeny of Ankole and Zebu dams. Hybrid vigour appeared to have little effect on carcass traits.

Fleece and skin samples were taken at intervals from birth to 86 days from Scottish Blackface lambs born into one line selected for hairiness, another for fineness and an unselected control line. The birthcoats in the Hairy line were longer than those in the Fine line, and had a predominance of Plateauo and Pi fibre type arrays, whereas P2 and P3 arrays predominated in the Fine line. The Hairy group had significantly more halo hairs and hairy-tip curly-tip fibres, but significantly fewer histerotrichs.

Most secondary follicles had fibres by 56 days, and development appeared to be faster in the Control than in the other two groups. The Hairy and Control groups had consistently high percentages of primary medullation, whereas the Fine group had a peak at 14 days. The Fine group had consistently low secondary medullation, whereas it increased in the Hairy and Control lambs, apart from a set-back at 28 days.

Up to 56 days the Fine group had no fibre shedding in the skin, and the Hairy group had most, although the incidence in any sample was less than 2%. There was more follicle inactivity at 86 days, and the Fine group then had most.

The objective of this study was to determine the effect of stage of lactation on the concentration of polyamines (spermine, spermidine, and putrescine) in sow colostrum and milk. Ten, third parity sows (Landrace×Yorkshire) were sampled throughout a 28-day lactation. Litter size for all sows was standardized at 10 sucking piglets immediately after farrowing. Colostrum collection began at the birth of the first piglet (0 h), and colostrum was subsequently collected 6, 12, 18, and 24 h after farrowing. Sow milk was sampled on days 3, 7, 10, 14, 17, 21, 24 and 28 of lactation. Spermine concentration in sow colostrum was relatively constant for the first 12 h after farrowing and then rapidly increased to 10·26 μmol/l 24 h after farrowing (linear and quadratic effect; P<0·01). In contrast, the concentrations of spermidine and putrescine showed little change during the first 24 h with spermidine concentration in colostrum ranging from 3·76 to 4·54 μmol/l, and putrescine concentration ranging from 1·15 to 1·52 μmol/l. The highest concentration of spermine in sow milk was 18·82 μmol/l, observed on day 7, with levels gradually declining to 4·28 μmol/l by day 28 of lactation (linear and quadratic effect; P<0·01). In contrast, the concentration of spermidine increased during lactation ranging from a low of 4·32 μmol/l on day 7 to a high of 9·01 μmol/l on day 28 of lactation (linear effect P<0·01; quadratic effect P=0·03). Putrescine concentration ranged from 1·53 μmol/l on day 3 to 1·84 μmol/l on day 28 (linear effect P=0·07; quadratic effect P>0·05). The results of this study show sow colostrum and milk provide polyamines for sucking piglets before weaning, which could potentially modulate intestinal maturation.

In a multibreed experiment, 292 heifers from 25 British cattle breeds were fed a standard pelleted diet ad libitum from 12 to 72 weeks of age. Inter-age correlations involving body weight, weight gain, average daily food intake and cumulated food intake were measured across traits at the same age and within and across traits at ages separated by an interval of 12, 24, 36 and 48 weeks. Within-breed correlations were phenotypic but between-breed correlations were genetic.

The between-breed inter-age correlations involving body weight, cumulated intake and average daily intake were all very high (range 104 to 0·94) and declined only gradually as the intervening age interval increased from 12 to 48 weeks. Inter-age correlations involving weight gain were lower but usually above 0·7. The within-breed correlations by contrast were invariably lower and declined much more rapidly as the age interval increased. Although the between-breed and within-breed correlations differed in magnitude there was a similarity in their overall pattern.

For breed samples, body weight at young ages provided very accurate rankings at later ages for body weight, cumulated intake and average daily intake. Body weight was almost as reliable as food intake itself for predicting cumulated intake.

For individuals within breeds predicted rankings at later ages were not very accurate for body weight or cumulated intake and they became worse as the age interval increased. Neither weight gain nor average daily intake could be predicted with any reliability.

Inter-age correlations while dependant on age interval did not depend on age itself in the case of body weight and cumulated intake, although they were age-dependant for weight gain and to a lesser extent for average daily intake. Reciprocal correlations between body weight and cumulated intake were very symmetrical whereas those involving weight gain and daily intake were asymmetrical.

Predictions of two models were compared. The models relate ovulation rate (OR) and prenatal survival (PS) to litter size (LS): the uterine capacity model (UCM), where maximum LS is limited by uterine capacity (UC), and the threshold model (TM) whereby PS is modelled as a binary threshold trait. Records were simulated according to both models using statistics from Trench Large White gilts. Both models were able to reproduce closely the observed curvilinear relationship between OR and LS, with LS reaching a plateau at high OR. Several genetic correlations (ρg) fulfilling the conditions h20R = 0·34 and h2LS = 0·12 (the residual maximum likelihood estimates in the population) were studied by means of stochastic computer simulation. The genetic correlation between OR and LS was very sensitive to changes in h2uc, whereas ρgLS,PS was always positive, and ρgOR,PS was always negative. The correlation between PS and UC was larger than 0·90, except for very small h2UC This suggests that genes affecting PS have a strong influence on UC and that PS can be a good indirect criterion to select for UC. Both models predicted that the advantage of an index combining OR and LS with respect to direct selection on LS diminishes in successive generations of selection and that the size of the experiment needed to detect significant differences is very large. Records were also simulated by halving the mean and variance of UC, so as to mimic unilateral hysterectomy-ovariectomy (UHO). If the UHO treatment results in halving UC, LS of UHO females should behave very much as half the UC of intact females.

Two experiments were undertaken to investigate the effect of herbage allowance and concentrate food level offered to twin-bearing ewes in the last 6 weeks of pregnancy on ewe and lamb performance. In each study, 96 twin-bearing Greyface (Border Leicester × Scottish Blackface), Rouge × Greyface and Texel × Greyface ewes were used. In experiment 1, the ewes were allocated to eight treatments consisting of two herbage allowances (1·3 and 2·6 kg herbage dry matter per ewe per day) and four concentrate food levels ranging from zero to 1000 g per day. In experiment 2, ewes were offered four herbage allowances (1·3, 1·75, 2·2 and 2·6 kg herbage dry matter per ewe per day) and zero or 500 g concentrates per day. In experiment 1, herbage allowance and concentrate food level had no significant effect on lamb birth weight, lamb mortality or lamb performance up to weaning. Herbage dry matter intake decreased linearly with increasing concentrate food level with a substitution rate of 18 g of herbage dry matter per 100 g concentrate food level. In experiment 2, lamb birth weight increased with increasing herbage allowance (P < 0·01) and with increasing concentrate food level (P < 0·01). However, lamb performance to weaning was unaffected by late pregnancy nutrition. The satisfactory levels of performance obtained with ewes offered grass-only diets in late pregnancy indicates that grazed grass is a high nutritive value food for twin-bearing ewes in late pregnancy.

Post-weaning growth, dry-matter intake, carcass characteristics, chemical composition of the 9th to 11th rib section and food conversion efficiency were evaluated for Simmental bulls (15) and heifers (18) and Hereford bulls (17) and heifers (14) from approx. 240 days of age to 470 days of age. Within each breed × sex sub-class, animals were assigned to one of three energy intake levels: (1) ad libitum, (2) 795 kJ metabolizable energy (ME) per kg0·75 per day and (3) 544 kj ME per kg0·75 per day.

Simmentals tended to exhibit higher post-weaning growth rate and consumed more dry matter than Herefords. At ad libitum intakes, efficiency estimates (gain/food) were greater initially but decreased more rapidly for Herefords than for Simmentals. No differences were observed in efficiency estimates between Herefords and Simmentals fed at 795 kJ ME per kg0·75 per day and both breeds fed at this level improved in efficiency as time on test increased.

Bulls tended to have greater growth rate, and consume more dry matter than heifers. At ad libitum food intake, bulls were more efficient than heifers and the decrease in efficiency of the two sexes was parallel. Heifers fed at 795 kJ ME per kg0·75 per day had a constant efficiency throughout the test while bulls improved in efficiency.

Production and endoparasitism of first grazing season Holstein heifers and steers were investigated over two grazing seasons. Studies were conducted on low-lying peaty soil. In year 2000, 40 animals were included in a 2×2 factorial, replicated experiment with two sexes (steers v. heifers) and two stocking rates (SR): normal v. low (840 v. 420 kg live weight per ha at turn-out) in a set stocking grazing system. Mean grass heights over the entire season were 6·3±4·8 cm (mean±s.d.) at normal SR and 9·8±6·1 cm at low SR. Mean daily live-weight gain during grazing was significantly (P<0·001) lower at normal SR (256±147 g) compared with low (468±142 g) but sex did not significantly affect the daily gain. Serum pepsinogen levels, indicating uptake of gastric nematodes, were higher at normal SR and consistently higher in steers compared with heifers, although not significantly. In 2001 the experiment included 80 animals in a 2×2 factorial experiment with two sexes (steers v. heifers), two grazing systems (set stocking and two-paddock rotation) and four replicates. Pasture quality was low, 570 g digestible organic matter per kg organic matter and 139 g crude protein per kg dry matter on average, independent of grazing system. Mean daily live-weight gain was not significantly affected by grazing system. A tendency (P=0·07) to lower daily gain for the heifers than for steers was observed (427±161 g v. 474±138 g). Lower levels of pasture contamination with parasites were observed in the paddocks without grazing up to mid July but otherwise rotation did not prevent parasite infections. It is concluded that first grazing season steers and heifers have the same potential for growth when grazing marginal areas with low pasture quality. Set stocking or the two-paddock rotation scheme did not affect productivity or level of parasite infection at the end of season. Parasite infections became a problem at high SR. SR is an important factor for both daily live-weight gain per animal and total production per ha. However, due to the very heterogeneous structure of marginal areas there is a need for other indicators than kg live weight per ha at turn-out in order to define a clear relationship between stocking rate and production.

Based on the Agricultural Research Council's feeding system, equations were developed that allow the calculation of the metabolizable energy requirements for maintenance and weight gain by cattle, separately or combined. A general equation was developed for the straight-forward calculation of the total metabolizable energy requirements of growing and fattening cattle for any combination of body weight, rate of weight gain, age, level of muscular work, and metabolizable energy concentration of the diet. The estimates of energy requirement made by the use of this equation are in excellent agreement with those made by the Agricultural Research Council using an iterative method.

In addition to avoiding the awkward iterative process, the equations proposed are readily adaptable to computer use.

Serum thyroxine levels were observed in 15 Soay ewes and seven rams for 1 year. From June to November the ewes had thyroxine concentrations 40% higher than the rams, and the level in both sexes increased from December to May, demonstrating a seasonal cycle in thyroxine level that could be associated with the annual wool growth cycle. To hasten the onset of wool follicle activity, nine Wiltshire sheep were given thyroxine implants in autumn when the follicles normally become inactive, but only a slight increase in follicle activity occurred. Eighteen Soay sheep were given thiouracil orally for 1 month in spring in an attempt to block a possible seasonal increase in thyroxine secretion that might be associated with the normal regrowth of wool, but no retardation of the spring regrowth was obtained.

The inter-breed relationship of birth weight and maternal and paternal weight in cattle was examined using published data on over 200 different breeds from a total of over 500 experiments. This collection of data was far from optimal for estimating genetic between-breed regressions and should have contained fewer breeds and more experiments on the same breed. Nevertheless, it was established that the mean mature weights of bulls and cows maintained a constant ratio throughout the range of breed size but mean calf weight declined relative to mean sire and mean dam weight as breed size increased.

When considered in conjunction with other results on litter weight at birth, the best relationship to use appeared to be that of proportionality to dam weight to the power 0·83, which could uniformly be applied to both breeds and species. The constant of proportionality found for cattle was 0·197 which can be compared with estimates of 0·183 for sheep, 0·185 for dogs and 0·165 on average for mammalian species.

The effect of energy supply between 45 and 100 kg body weight (BW) on the contribution of lean tissue (muscle plus intermuscular adipose tissue) to total protein mass was studied in Large White castrated males (cLW), crossbred Piétrain × Large White castrated males (cPPx) and boars (bPPx). The pigs were allocated to four energy levels (0·70, 0·80, 0·90 or 1·00 ad libitum) and kept in metabolism cages in experiment 1 or given food ad libitum and kept in individual pens in experiment 2. Daily protein supplies were calculated to be non-limiting for growth and identical for all pigs in experiment 1. Temperature was 23°C in both experiments. The pigs were slaughtered at 100 kg BW and physically dissected; the body tissues were chemically analysed. Taking into account housing conditions, the food intake of pigs in experiment 2 corresponded to 1·20 of ad libitum intake of pigs in experiment 1; data of both experiments were combined. The energy supply and the type of pig influenced significantly the protein content of empty BW (eBW) (170 g/kg on average), of lean (184g/kg on average) and non-lean compartment (eBW minus lean, 152 g/kg on average), the proportion of total protein deposited in lean (604 g/kg of total protein, on average) but not the protein content in fat-free eBW (209 g/kg on average). The fat-free eBW can be predicted as 4·8 times the body protein mass.