Forage kochia [Bassia prostrata (L.) A. J. Scott] is competitive with annual weeds and has potential for use in reclamation of disturbed land. However, land managers are reluctant to use forage kochia in revegetation programs due to lack of understanding of its compatibility with or invasiveness in the native plant community. We conducted two greenhouse experiments, one to compare the competitive effect of forage kochia versus perennial grasses on growth of cheatgrass (Bromus tectorum L.) and one to study the effect of forage kochia on growth of native perennial grasses. In the first experiment, a single seedling of B. tectorum was grown with increasing neighbor densities (0 to 5 seedlings pot−1) of either forage kochia, crested wheatgrass [Agropyron cristatum (L.) Gaertner × A. desertorum (Fisch. ex Link) Schultes; nonnative perennial grass], or thickspike wheatgrass [Elymus lanceolatus (Scribn. & J. G. Sm.) Gould; native perennial grass]. Bromus tectorum growth was reduced moderately by all three perennial neighbors, but A. cristatum and E. lanceolatus had more effect on B. tectorum when compared with forage kochia. This experiment was repeated and similar results were observed. In the second experiment, forage kochia was grown with each of four native cool-season grass species: basin wildrye [Leymus cinereus (Scribn. & Merr.) Á. Löve], bluebunch wheatgrass [Pseudoroegneria spicata (Pursh) Á. Löve], E. lanceolatus, and western wheatgrass [Pascopyrum smithii (Rydb.) Á. Löve]. Forage kochia had no effect on height, tiller number, and aboveground biomass of native grasses. Similarly, native grasses did not show a significant effect on forage kochia seedlings. This experiment was also repeated, and forage kochia somewhat reduced the aboveground biomass of L. cinereus and P. spicata. However, all native grasses significantly reduced change in height, branching, and aboveground biomass of forage kochia. These results suggest that forage kochia interfered with B. tectorum seedling growth, but it showed little competitive effect on native grass seedlings.

The robustness of competitive attributes of cereals such as rapid and uniform seedling emergence, tillering, early biomass accumulation and canopy closure, and height advantage over weeds have not yet been tested under environmental conditions typical of no-till (NT) cropping systems. Our objective was to evaluate the effects or NT practices on growth and productivity of Avena sativa, Triticum aestivum, Hordeum vulgare, and associated weeds. The experiment was conducted on a Kamouraska clay at La Pocatière, QC, in 1994, 1995, and 1996. Avena sativa, T. aestivum, and H. vulgare were grown under tilled and NT practices. Cereal growth parameters were measured six (1994) or seven (1995) times between planting and the 11th week after planting but only once in 1996. Grain yields and yield components were determined at crop maturity. Avena sativa and H. vulgare populations were little affected by tillage, whereas T. aestivum populations were reduced by 16 to 20% in NT systems. Growth in height in NT systems was either similar or greater than in tilled systems in all three cereals. Cereal leaf area index (LAI) and biomass accumulation was also comparable between tillage systems, except for T. aestivum LAI in 1994, which was greater in tilled plots on two sampling dates. Response of annual dicots to tillage was inconsistent in all crops. Annual monocots dominated in some but not all NT systems. Perennial dicots dominated in NT systems, whereas perennial monocots were more abundant in tilled systems in all three cereals. Avena sativa and T. aestivum yields in NT plots were comparable or greater than in tilled plots, in spite of having either lower test weights (A. sativa) or lower 1,000-grain weights (T. aestivum). NT T. aestivum productivity was maintained in spite of reduced plant establishment. Hordeum vulgare yields were also similar across tillage systems, except in 1995, when yields in tilled plots were greater than in NT plots. The height advantage observed for NT H. vulgare did not result in improved yields. All three cereals, and particularly A. sativa, appeared well suited to NT systems, despite the pressure provided by different weed groups, compared to tilled systems. However, results suggest that NT production of cereals could benefit from improved attention to perennial dicot control and crop seedling establishment, particularly for T. aestivum.

Improved crop tolerance and weed suppressive ability are tactics that may reduce the negative effect of weeds on crop yield. Irrigated field experiments were conducted to compare leaf area index (LAI), intercepted photosynthetic photon flux (PPF), and relative tolerance and velvetleaf suppressive ability among two old (circa 1940) and two modern corn hybrids. Each hybrid was grown in monoculture and in mixture with velvetleaf at 1, 4, 16, and 40 plants m−1 row. Plants were periodically harvested in monoculture plots to obtain estimates of corn LAI, and PPF interception was measured. Variation in hybrid tolerance to velvetleaf competition for light was evaluated by comparing among hybrids the coefficients of a regression of corn yield loss on velvetleaf density. Velvetleaf seed capsule production in the presence of each hybrid was compared to evaluate variation in velvetleaf suppressive ability among hybrids. Maximum corn yield loss was 32% lower for the two old hybrids, and velvetleaf capsule production was reduced by 62% at low velvetleaf densities in 1995 compared to the modern hybrids. In 1996, yield loss of the modern hybrid 3394 was 74% lower than that of the other three hybrids at low velvetleaf densities, whereas maximum yield loss of the old hybrid 336 was 44% lower at high densities. Velvetleaf capsule production did not vary among hybrids at any velvetleaf density in 1996. Hybrids with greater tolerance and velvetleaf suppressive ability also had greater LAI and PPF interception, suggesting optimized corn LAI and PPF interception may be useful in an integrated weed management program.

Redstem is an important weed in California water-seeded rice fields because of its aquatic habit, wide distribution, interference with harvest, and resistance to the herbicide bensulfuron. Our objective was to understand the mechanisms of competition for light between rice and redstem, with the goal of improving redstem control. A replicated greenhouse experiment was done in 1993 and 1994. Rice was water-seeded at a rate of 400 seeds m−2, and redstem was seeded simultaneously at approximate densities of 0, 50, and 100 seeds m−2 in continuously flooded 0.77 m2 basins. Plants were harvested once at final harvest in 1993 and twice in 1994, with an additional nondestructive sampling 34 days after seeding (DAS). Despite slower early growth, redstem height exceeded rice height about 45 DAS. At the midseason harvest in 1994 (56 DAS), no effects of redstem competition on any rice response variables were detected. However, at final harvest (110 and 118 DAS, 1993 and 1994, respectively) redstem competition at both treatment densities reduced rice tiller density, panicle density, shoot drymass, and grain drymass. Redstem competition reduced rice growth only after penetrating the canopy. Shade cast by redstem through rice maturity decreased shoot and grain production and increased tiller mortality. Lodging caused by redstem further affected rice growth. Season-long competition from redstem at mean densities of 67 and 110 plants m−2 reduced rough rice yields 31 and 39%, respectively, making redstem the most competitive broadleaved rice weed yet studied. Improved understanding of rice-redstem interactions indicates that using alternative herbicides to bensulfuron is unlikely to increase yield losses to redstem, and that control may be improved by increasing rice plant densities or slightly delaying early season chemical control. Because these strategies are mechanistic, they may also be useful for controlling other rice weeds with growth patterns similar to redstem.

The photosynthetic responses of rice (C3) and three Echinochloa species (C4), barnyardgrass, early watergrass, and late watergrass, to changes in CO2 intercellular partial pressure, light intensity, and leaf temperature were investigated under laboratory conditions. The three Echinochloa species exhibited photosynthetic responses characteristic of C4 plants. The three weedy species showed higher efficiency for CO2 utilization at low CO2 intercellular partial pressure (CO2i) than rice. Compensation and saturation of CO2i for photosynthesis were lower in the weedy species than in rice. The maximum photosynthetic rates at high light intensity were 33.5, 32.7, 30.5, and 21.5 μmol CO2 m-2s-1 for barnyardgrass, early watergrass, late watergrass, and rice, respectively. Photosynthesis temperature optimum was 35 to 37 C for the three Echinochloa species and 33 C for rice. Overall, under simulated summer conditions, the four taxa showed a photosynthetic ability hierarchy with regard to gas exchange performance as follows: barnyardgrass ≥ early watergrass > late watergrass > rice.

Sweet corn is planted over a long season to temporally extend the perishable supply of ears for fresh and processing markets. Most growers' fields have weeds persisting to harvest (hereafter called residual weeds), and evidence suggests the crop's ability to endure competitive stress from residual weeds (i.e., crop tolerance) is not constant over the planting season. Field studies were conducted to characterize changes in the residual weed community over the long planting season and determine the extent to which planting date influences crop tolerance to weed interference in growth and yield traits. Total weed density at harvest was similar across five planting dates from mid-April to early-July; however, some changes in composition of species common to the midwestern United States were observed. Production of viable weed seed within the relatively short growth period of individual sweet corn plantings showed weed seedbank additions are influenced by species and planting date. Crop tolerances in growth and yield were variable in the mid-April and both May plantings, and the crop was least affected by weed interference in the mid-June and early-July planting dates. As the planting season progressed from late-May to early-July, sweet corn accounted for a great proportion of the total crop–weed biomass. Based on results from Illinois, a risk management perspective to weeds should recognize the significance of planting date on sweet corn competitive ability. This work suggests risk of yield loss from weed control failure is lower in late-season sweet corn plantings (June and July) than earlier plantings (April and May).

The inclusion of competitive crop cultivars in crop rotations is an important integrated weed management (IWM) tool. However, competitiveness is often not considered a priority for breeding or cultivar selection by growers. Field pea (Pisum sativum L.) is often considered a poor competitor with weeds, but it is not known whether competitiveness varies among semileafless cultivars. The objectives of this study were to determine if semileafless field pea cultivars vary in their ability to compete and/or withstand competition, as well as to identify aboveground trait(s) that may be associated with increased competitive ability. Field experiments were conducted in 2012 and 2013 at three locations in western Canada. Fourteen semileafless field pea cultivars were included in the study representing four different market classes. Cultivars were grown either in the presence or absence of model weeds (wheat and canola), and competitive ability of the cultivars was determined based on their ability to withstand competition (AWC) and their ability to compete (AC). Crop yield, weed biomass and weed fecundity varied among sites but not years. Cultivars exhibited inconsistent differences in competitive ability, although cv. Reward consistently exhibited the lowest AC and AWC. None of the traits measured in this study correlated highly with competitive ability. However, the highest-yielding cultivars generally were those that had the highest AC, whereas cultivars that ranked highest for AWC were associated with lower weed fecundity. Ranking the competitive ability of field pea cultivars could be an important IWM tool for growers and agronomists.

Two field experiments were conducted in Oklahoma to determine whether changing wheat row spacing, seeding rate, and cultivar would reduce interference from rye. Wheat row spacing did not affect rye seed production. Averaged over row spacing, increasing wheat seeding rate from 67 to 134 kg/ha reduced rye seed production 21 and 25% in two experiments. At one site, grain yield of rye-infested wheat was increased 27 and 23% by doubling wheat seeding rate in 10- and 20-cm rows. Doubling the seeding rate of rye-infested wheat in 30-cm rows did not increase wheat yield. At a second site when data were pooled over row spacings, increasing the wheat seeding rate from 67 to 101 kg/ha increased yield of rye-infested wheat 21%, but yield was still 36% less than for rye-free wheat. In two additional experiments, rye seeds in harvested wheat were reduced 36% by increasing the wheat seeding rate from 60 to 162 kg/ha, whereas yield of rye-infested wheat was increased 82%. Nine cultivars were compared for competitive ability against rye in four experiments. Of 32 possible cultivar location situations, the rye-induced yield loss of ‘Jagger’ wheat was less than 16 other cultivar location situations, whereas the yield loss of ‘Triumph 64’ wheat was less than 10 other cultivar location situations. Mature wheat height, wheat yield, and wheat maturity classification were each negatively correlated to rye yield in infested wheat at two of four sites. Wheat plant density, head density, sunlight interception, and fall cover capability classification were each negatively correlated to rye yield at one of four sites.

Field pea cultivars often differ in weed competition. In several crop types branching has been cited as one of the characteristics conferring competition with weeds. The objective of this study was to determine the difference in weed competition among field pea cultivars differing in basal branching and other characteristics. Ten field pea cultivars with divergent basal branching ability were seeded at 50 plants m−2 under weedy and weed-free conditions to evaluate their competition with weeds. Branching did not differ greatly between cultivars and was not associated with the weed competiveness of the field pea cultivars. The forage pea cultivars, which were leafed and had longer vines, were much more competitive than the semi-leafless grain cultivars. As a result, the forage cultivars were better able to suppress weeds and maintain their yield under weed presence. However, the absolute seed yield of the forage pea cultivars was low, making them a poor choice for seed production. Vine length and the leafy characteristic may be important genetic characteristics associated with competition in field pea cultivars.

The interaction between wheat and perennial ryegrass seed density and seedlings of different ages was studied under controlled conditions. Root length of perennial ryegrass, after sowing, was suppressed by wheat and was dependent on the density of wheat seeds. Shoot growth of perennial ryegrass, however, was unaffected by the presence of wheat. Perennial ryegrass density had no effect on the first 2 wk of wheat seedling growth. The age of wheat seedlings had no appreciable influence on either root or shoot growth of perennial ryegrass. The present study highlights the need for an unbiased two-way experimental design to identify the dominant competitor.

In a garden experiment we monitored tillering and flowering for target plants of two grass species, surrounded by one to five neighbours and compared them with control plants growing alone. We analysed the results using a non-linear regression. The species differed markedly in their seasonal patterns of growth but these differences faded when growing with two or more neighbours. The presence of neighbours had a significant depressing effect on both tillering and flowering in both species. Effects and responses were strongly asymmetrical concerning species and mode of growth. Andropogon semiberbis is in disadvantage against Trachypogon plumosus concerning vegetative growth. However, T. plumosus is strongly affected in its reproductive growth by competition not only from A. semiberbis but also from conspecific neighbours. We conclude that the complexity of interactions between effects and responses as well as vegetative and reproductive growth could have a net result of competitive equivalence or compensating competitive abilities of these two species.

‘Phalanx’ and ‘guerrilla’ phenotypes have been characterized as distinct, adaptive growth strategies exhibited by marine encrusting taxa, as well as a variety of other colonial taxa, that differ in patterns of colonial growth and areal expansion. Phalanx morphs exhibit compact growth, expanding outward concentrically and generating radially symmetric colony shapes, whereas guerrillas exhibit diffuse growth and typically elongate, asymmetric colony shapes. Several species in the colonial hydroid genus Hydractinia display inter-genotypic morphological variation in early developmental growth, although it is unclear if and how this growth form variation is tied to colony symmetry. Here I show that the phalanx versus guerrilla distinction does not adequately characterize genetic variation in Hydractinia growth form. Genotypes characterized by extreme mat and stoloniferous growth exhibited high levels of symmetry while genotypes generating growth forms intermediate between these two extremes were more asymmetric. Asymmetric growth is tied to reduced field fitness as a result of slower growth, reduced investment in future reproduction, and increased susceptibility to abiotic environmental stress. Asymmetric, guerrilla-like growth may be the morphological symptom of maladaptive growth early in colony development. This notion contrasts greatly with the traditional view of guerrilla growth as an adaptive strategy. Several hypotheses are proposed to address why asymmetric, guerrilla-like growth may be maladaptive in this and similar systems.

The objective of this study was to determine any differences in the competitive ability of rams differing in levels of sexual performance. Twenty sexually experienced Targhee-type rams were preselected from a population of 95 males for exhibiting relatively high and low ejaculation rates when individually exposed to oestrous ewes for 30 min. Each of 10 high performers (HP) were paired with each of 10 low performers (LP) on three occasions when competing for food (after food deprivation) and twice when competing for an oestrous female. Tests were 5 min in duration. Food-deprived HP and LP rams were equally aggressive when competing for a source offood that could only be accessed by one ram at a time. Time offeeding by HP and LP rams did not differ. HP rams spent more time with the oestrous ewe and attained more mounts in a greater number of pairings than LP rams even though HP and LP rams did not differ in number of successful agonistic interactions (active displacements and successful defences) when competing for the ewe. It was concluded that HP and LP rams, taken as groups, do not differ in their basic competitive ability and that the greater mating success of HP rams can be explained by their higher levels of libido.