The loop configuration, structure and growth are studied in a series of megathiridoid dorsal valves ranging in length from 0·7 to 6 mm. The structure of the posterior and anterior sectors of the descending lamellae and their relationships with the socket ridges, dorsal septa and dorsal valve floor, are studied, illustrated and described. Detailed studies show that internal structures display great variability. In the megathiridoids the lamellae of more typical terebratulide loops are replaced by partially developed loops in early developmental stages, resulting in a mix of free sections of loop together with sections where the lophophore rests directly on the epithelium of the valve floor. Comments on the ecology of the megathiridoid species are included and relate their unusually wide gapes, which position their lophophores fully accessible to the open sea, with their strong lophophore attachments and loss of typical free loops. The new genus Joania (type species Terebratula cordata Risso, 1826) is erected for those Argyrotheca which, although having a typical megathiridoid brachidium, differ in their adult crural development, their narrow hinge line, their prominent cardinal process, their characteristic dorsal median septum and their tuberculate radial ridges terminating anteriorly in tubercles.

Although sophisticated geochemical studies tell us that tonolite-trondhjemite-granodiorite (TTG) plutonic complexes must be formed by partial melting of metabasaltic source material, they cannot tell us the tectonic regime in which this crust was formed, nor how large volumes of TTG magma can be generated. This study suggests that a solution to TTG arc crust formation requires a strongly interdisciplinary approach, to resolve the tectonic setting (slab melt verses mafic lowermost crust sources), the time and length scales for melting and extraction, and the role of melt segregation mechanisms in the formation of both Archean TTGs and more recent adakite-like magmas. The aim of this paper is to present an experimental approach which, when coupled with numerical models, allows some of these issues to be addressed. The experiments are designed to reproduce the local changes in bulk composition that are predicted to occur in response to buoyancy-driven melt segregation along grain edges and associated compaction of the solid residue. The preliminary study presented here documents the changes we observe in the melt composition and melt and solid phase modes between earlier direct partial melting and the new segregation equilibration experiments on metabasalt bulk compositions. The results suggest that if dynamic melt segregation and equilibrium processes are active, they may modify the normally robust geochemical indicators, such as Mg-numbers, which are typically used to develop models of TTG petrogenesis.

The postcranial skeleton of Ariekanerpeton sigalovi (Seymouriamorpha: Discosauriscidae; Lower Permian, Tadzhikistan) differs from that of like-sized Discosauriscus specimens in showing: wider interclavicle anterior plate with smaller, rhomboidal sculptured field on its ventral surface not reaching plate posteromedial margins; broader interclavicle plate-stem junction; slightly narrower interclavicle posterior stem; anteroposteriorly narrower clavicle ventral plate with convex posterior margin; shorter, more robust humerus; four phalanges on fourth manus digit (five phalanges in Discosauriscus); ilium more elongate dorsoventrally with anteroposteriorly narrower neck; posterior process of iliac blade oriented distinctly posterodorsally rather than horizontally; more gracile atlantal rib; broadened distal end of second presacral rib. A revised cladistic analysis of the best known seymouriamorph species retrieves Ariekanerpeton either as sister group to both species of Discosauriscus, or to D. austriacus only.

Cell culture systems have been developed to assess the potential pathogenicity of mineral dusts. The in vitro cytotoxicities of a variety of dusts towards mouse peritoneal macrophages, Chinese hamster lung cells (V79 cell line) and human alveolar type II cells (A549 cell line) were investigated.

Non-pathogenic dusts were found to be inert in vitro. Fibrogenic non-fibrous dusts such as silica were only cytotoxic towards macrophages. Fibrous dusts which are both fibrogenic and carcinogenic in vivo are cytotoxic towards all three cell types, their cytotoxicity being dependent on fibre size. The size range important for the observed biological effect is longer than about 8 μm and thinner than about 1·5 μm.

Loch Fleet is an upland loch in SW Scotland, distant from significant local pollution, and unlikely to be affected by changes during the five-year project. The underlying geology is granitic. The 111 ha catchment is 90% moorland, dominated by heather and flying bent; 10% is planted with conifers.

Rainfall is approximately 2200 mm per year, predominantly from W and SW. More than half of rain events are pH>4·7; mean pH is 4·84 (1985). Marine salts are the major solutes. Deposition of acidity is about 0·1 g H+ m−2 per year, about twice that of eastern Scotland; deposition of non-seasalt sulphur is 1·6 g S m−2 per year. Soils are shallow and comprise organic peats, peaty gleys, and peaty rankers; base saturation is 3–19%. Soils (pH 2·5 to 4·6) are most acid in the forested area.

The loch is more acid (×5) than bulk deposition; additional acidity is mobilised from soils by the predominantly (>90%) superficial drainage. Moorland drainage dominates; that from the forest is more acid, but very variable in quantity and composition.

Catchment interventions in 1986 and 1987 are designed to alter the chemistry of drainage waters and the loch, so that the fishery can be restored. In addition to the work reported, extensive soils, sediment and biological investigations are underway.

The Miocene-Pliocene Macusani ash-flow tuffs and glasses from SE Peru are a rare example of two-mica felsic peraluminous volcanic rocks. They outcrop in three separate tectonic basins of the Cordillera Oriental in the Central Andes. In the Macusani field, the rocks are characterised by andalusite and muscovite phenocrysts. Compositions are both very felsic and peraluminous, similar to two-mica granites. Field relations, age differences and isotopic heterogeneities suggest that several distinct magma batches were involved. Two separate magmatic stages are recognised: (1) partial melting and evolution at or near the source region, and (2) shallow-level crystallisation and eruption. Magma genesis involved partial melting of metapelitic materials, with internally controlled. High heat flux, rapid heating, elevated temperatures and F-rich compositions were essential for the production of these mobile, H2O-undersaturated magmas. Chemical variations between the erupted products can be attributed to different degrees either of partial melting in somewhat variable source materials and (or) of fractional crystallisation at shallow levels. We discuss some important differences between the magmatic evolution of the Macusani Volcanics and of Hercynian and Himalayan two-mica granites.

The Vittarieæ, as described by Christensen, comprises five genera, viz. Vittaria, Monogramma, Antrophyum, Hecistopteris, and Anetium, all of which are epiphytic forms growing in the damp forests of the Old and New World Tropics. All of them possess creeping rhizomes on which the fronds are arranged more or less definitely in two rows on the dorsal surface. The fronds are simple in outline, with the exception of those of Hecistopteris which are dichotomously branched. The venation of the fronds is reticulate, except in Hecistopteris where there is an open, dichotomous system of veins, and in Monogramma, in some species of which the venation consists simply of a mid-rib. An interesting feature, which has proved to be valuable as a diagnostic character, is the presence of “spicule cells” in the epidermis of the fronds. These spicule cells are elongated cells containing spicules of silica, and their presence appears to be universal in the Vittarieæ. The sporangia are fairly constant in form through-out the group, but their distribution is extremely varied. The roots are characteristically provided with very numerous reddish-brown root hairs, a character shared with other epiphytic Ferns. The Gametophyte is divergent from the common cordate type in all cases where it has been investigated. Such are the main external characteristics of the group under consideration.

The opportunities for observation were afforded by the voyage and subsequent drift of the S.Y. Endurance. During December 1914 and January 1915 for a period of six weeks she successfully bored her way through pack-ice of every description—drift-ice, open-pack, and very frequently even close-pack. Continually fighting, she penetrated from 59° to 72° S. lat., and finally reached the land water off Coats Land on the latter parallel. As the crow flies, therefore, she was navigated through ice for nearly 800 geographical miles on this voyage; her actual course among the ice-fields and floes was computed to exceed 2000 miles, an achievement without parallel in the Antarctic. The principle adopted was to keep to the east, where presumably there is less pack than in the west; if the Endurance experience is a normal one, however, the meridian of 20° W. long., which was followed, is certainly not far enough east.

The present study explores the macroevolutionary dynamics of shape changes in the humeri of all major grades and clades of early tetrapods and their fish-like forerunners. Coordinate point eigenshape analysis applied to humeral outlines in extensor view reveals that fish humeri are more disparate than those of most early tetrapod groups and significantly separate from the latter. Our findings indicate sustained changes in humeral shape in the deepest portions of the tetrapod stem group and certain portions of the crown. In the first half of sampled tetrapod history, subclades show larger than expected humeral disparity, suggesting rapid diffusion into morphospace. Later in tetrapod evolution, subclades occupy smaller and non-overlapping morphospace regions. This pattern may reflect in part increasing specialisations in later tetrapod lineages. Bayesian shifts in rates of evolutionary change are distributed discontinuously across the phylogeny, and most of them occur within rather than between major groups. Most shifts with the highest Bayesian posterior probabilities are observed in lepospondyls. Similarly, maximum likelihood analyses of shifts support marked rate accelerations in lepospondyls and in various subclades within that group. In other tetrapod groups, rates either tend to slow down or experience only small increases. Somewhat surprisingly, no shifts are concurrent with structural, functional, or ecological innovations in tetrapod evolution, including the origin of digits, the water–land transition and increasing terrestrialisation. Although counterintuitive, these results are consistent with a model of continual phenotypic innovation that, although decoupled from key evolutionary changes, is possibly triggered by niche segregation in divergent clades and grades of early tetrapods.

Since the preliminary announcement of the discovery of the prothalli of Tmesipteris and Psilotum communicated to the Royal Society of Edinburgh in June 1916, a considerable amount of additional material has been found, and it seems therefore desirable to place on record a more detailed account of these interesting structures. The preliminary description, as published, was based upon but a few specimens, and these were not sufficient to reveal all of the vegetative and reproductive features, nor enough to justify any broad general conclusions as to the phylogenetic position of the Psilotaceæ. The discoyery, however, has filled in an important and interesting gap in our knowledge; for the Psilotaceæ are the very last of the known Pteridophytes to surrender their gametophytes to the light of science.

If such tremendous agents did in reality exert their influence in the Alps, it is not conceivable that other countries, situated lower, and composed of materials much more frail, could have been spared. We hear in fact of similar traces, more or less distinct, in all quarters of the globe. From the bones of animals, natives of the hottest climates, which are found over a frozen region of vast extent in the Russian empire, Pallas concludes, that an enormous torrent had transported their carcases across the ridge of Tartary.

Calyptaulax brongniartii (Portlock) from the Caradocian Bardahessiagh Formation, Pomeroy, Northern Ireland is redescribed. Equivalent material from Scottish horizons (Balclatchie and Ardwell groups) is regarded as belonging to separate populations. Campbell's system of notation is used to describe variation in the arrangement and number of lenses in the eye. The most closely related species is C. strasburgensis (Ulrich & Delo) from the Edinburg Limestone of Virginia, U.S.A.

Granites and their associated extrusive rocks are formed in large volumes whenever the continental crust is heated by rising hot mantle, or thickened by collision processes. The complexity of rocks of the granite family is related to the complexity of the continental crust itself and the complexity of processes which lead to thermal perturbations. The light continental crust acts as a density filter which screens out heavy mantle magmas and leads to complex underplating and magma mixing processes. Perhaps the primary cause of crustal melting is the deep recycling of volatiles which are fixed in the oceanic crust before subduction. Modern studies of subduction and collision processes show the large scale and complexity of processes which modify old continental crust.

Although sophisticated geochemical studies tell us that tonolite–trondhjemite–granodiorite (TTG) plutonic complexes must be formed by partial melting of metabasaltic source material, they cannot tell us the tectonic regime in which this crust was formed, nor how large volumes of TTG magma can be generated. This study suggests that a solution to TTG arc crust formation requires a strongly interdisciplinary approach, to resolve the tectonic setting (slab melt verses mafic lowermost crust sources), the time and length scales for melting and extraction, and the role of melt segregation mechanisms in the formation of both Archean TTGs and more recent adakite-like magmas. The aim of this paper is to present an experimental approach which, when coupled with numerical models, allows some of these issues to be addressed. The experiments are designed to reproduce the local changes in bulk composition that are predicted to occur in response to buoyancy-driven melt segregation along grain edges and associated compaction of the solid residue. The preliminary study presented here documents the changes we observe in the melt composition and melt and solid phase modes between earlier direct partial melting and the new segregation equilibration experiments on metabasalt bulk compositions. The results suggest that if dynamic melt segregation and equilibrium processes are active, they may modify the normally robust geochemical indicators, such as Mg-numbers, which are typically used to develop models of TTG petrogenesis.

Since the ‘Grain to Green Program' was launched in 1999 in the Loess Plateau, China, the processes and patterns of ecological change have become important. Through the use of the fractional vegetation cover (FVC) index, this study examines the spatial distribution and temporal change of vegetation cover in the Loess Plateau during 2000–2014. Over this period more than 60% of the Loess Plateau has remained with little vegetation cover (FVC <30%). The spatial distribution pattern shows an overall increase from the NW to SE. Temporally, the vegetation cover exhibits a general trend of improvement. In 2000–2005, 2000–2010 and 2000–2014, the percentages of restored vegetation (vegetation with significantly increased FVC, P<0.05) were 2, 21 and 52%, respectively. The rate of vegetation cover restoration was highly variable among different bioclimatic zones. The expansion of restored vegetation was greater in the N of the Loess Plateau than in the south. Both human restoration activities and climatic fluctuation influenced the vegetation cover change. The ‘Grain to Green Program' emphasised vegetation restoration. Regional precipitation also had clear effects on vegetation cover. The results of this study reveal that vegetation change shows a non-linear process in response to climate and ecological restoration measures, and that the change gradually emerges over time. This study highlights the importance of considering the spatiotemporal variability in vegetation cover during the implementation of restoration programs, which could aid decision-making for the effective and sustainable management of large-scale restoration programs.

The Late Ordovician–Early Silurian succession in Jämtland includes the marine Kogsta Siltstone, which is unconformably overlain by the shallow-water Ede Quartzite that grades into the open-marine Berge Limestone. A Hirnantia shelly fauna dates the uppermost Kogsta Siltstone as Hirnantian, and shelly fossils indicate an Aeronian age for the Berge Limestone. Biostratigraphically highly diagnostic conodonts of the early-middle Aeronian Pranognathus tenuis Zone provide the first firm date of the Upper Ede Quartzite and the lowermost Berge Limestone. The Lower Ede Quartzite has not yielded fossils, but sedimentological data suggest it to be of Hirnantian age and reflect the glacio-eustatic low-stand. The contact between the Lower and Upper Ede Quartzite, here taken to be the Ordovician–Silurian boundary, appears to be an unconformity associated with a stratigraphic gap that at least includes the Rhuddanian Stage. The biostratigraphically important conodonts Pranognathus tenuis, Kockelella? manitoulinensis, and Pranognathus siluricus are recorded from Sweden for the first time, and these and other conodonts are used for correlations with coeval units in Europe and North America. In a regional review of Aeronian conodont faunas, three intergrading, apparently depth-related, conodont biofacies are recognised, the Jämtland conodonts representing the one characteristic of the shallowest water.

It is possible, that the following observations may contain little that is new to those who are familiarly acquainted with the details of the science of Hydrostatics. But as I have not met with any remarks on the subject, in the course of my limited reading, the experiments which were performed, and the conclusions to which they lead, are here submitted to the consideration of the Royal Society.

An onychophoran-like fossil animal, Paucipodia inermis gen. et sp. nov., is described from the Lower Cambrian Chengjiang lagerstatte in Yunnan, China. The animal was soft-bodied, and possessed six pairs of unjointed legs, each having two claws distally. The body was finely annulated, lacked trunk plates, and apparently terminated posteriorly with the last leg pair, without preserved evidence of a posterior trunk extension. Paucipodia widens the morphological range hitherto known in Cambrian lobopodians, notably regarding leg number and presence or absence of plates. Its morphology also provides support for the recently suggested anteroposterior orientation of Hallucigenia and Microdictyon.

Two regional-scale, recumbent folds control the structure of the Beinn Udlaidh area, Tyndrum, Perthshire. They reached their maximum development during D2, following the regional metamorphic peak, and are part of a stack of larger SE-facing recumbent folds formed during the ∼470 Ma Grampian Orogeny. The rocks belong to the Neoproterozoic–Lower Ordovician Dalradian Supergroup, and preserve a sedimentary transition between the Grampian Group and the overlying Appin Group. The latter occupies the core of the S-facing, recumbent Beinn Udlaidh Syncline (D2) which, with the underlying complementary Glen Lochy Anticline, is gently folded by a regional-scale structure, the Orchy Dome. The recumbent folds postdate an early fabric (S1), which is generally obliterated by the D2 imprint, but preserved as inclusion trails in regional metamorphic garnets, that are highly oblique to, and wrapped by, S2. It is concluded that the Dalradian rocks described here from below the Iltay Boundary Slide are in structural continuity with those of the Tay Nappe above, and that the Slide represents a structurally-modified disconformity between the Leven Schist (Appin Group) and the overlying Ben Lui Schist (Argyll Group). The Orchy Dome probably influenced the spatial distribution of minor intrusions and explosion vents of the lamprophyre suite.

Establishing the magnitude and frequency of floods within upland catchments on the basis of short-term gauged runoff records is crucially dependent upon the extent to which the record is truly representative. In the case of the River Dee, upstream of Crathie in Aberdeenshire, gauged discharge records are limited in length. Although the middle Dee has been gauged since 1929, the gauge within the upper catchment has only ten years of record. Thus, reliable estimates of the return intervals of extreme floods for this part of the Dee can only be obtained by using a variety of historical sources to extend the flood series.

Long-term rainfall records, where available, provide a valuable independent check on the reconstructed flood series. Such rainfall records, when analysed in terms of the magnitude, frequency and duration of major events, should, in general terms, correspond with the flood series. In this paper, the recurrence interval of extreme rainfalls of varying magnitude and duration in upper Deeside is estimated by extreme value analysis of the annual maximum series. The frequency of rainfall events above varying thresholds is also assessed. The existence of climatic fluctuations giving highly variable recurrence intervals for rainfall events of the same magnitude is demonstrated. Finally, the seasonality of frequent flood-producing storms is analysed. Patterns observed within the rainfall record are compared with those previously established within the historic flood series to substantiate and augment the flood record.