The Shyok Suture Zone is an oceanic remnant of the Neo-Tethyan ocean sandwiched between the Ladakh Batholiths to the south and Karakoram Batholith to the north. The Tirit granitoids in this suture are dark-coloured, relatively rich in ferromagnesian minerals and range from granodiorite–tonalite to gabbro–diorite in composition. Mafic igneous enclaves are quite common and they are intruded by NW–SE parallel doleritic and aplitic dykes. The Tirit granitoids have a wide range of major oxide compositions (SiO2 = 52.1–72.11 wt %, TiO2 = 0.21–1.23 wt %, Al2O3 = 11.42–13.52 wt %, MgO = 1.69–10.69 wt % and CaO = 3.24–9.31 wt %) and show calc-alkaline, metaluminous, I-type characteristics, transitional between primitive and mature arc continental plutons. Rare earth elements (REE) show considerable enrichment in light REE (LREE) as compared to the heavy REE (HREE). Late Cretaceous U/Pb dates (74–68 Ma) show that they formed during the pre-collision northward movement of India. The Tirit dykes are only slightly younger and probably part of the same episode.

Fossils are more and more used in phylogenetic evolutionary studies either for clade calibration, or as terminals in a dataset including morphological characters. The strength of these methodological advances relies however on the quality and completeness of the fossil record. For crickets (Insecta, Orthoptera, Gryllidea), few ancient (pre-Cenozoic) well-preserved fossils are known, except for isolated wings often classified in purely fossil groups and a few fossils found in Cretaceous amber. Here, we present two remarkable fossils from mid-Cretaceous amber of France, that were imaged using X-ray synchrotron microtomography and exhibit an exquisite preservation allowing description with a precision similar to that of extant taxa. Palaeonemobius occidentalis Laurent and Desutter-Grandcolas, gen. nov., sp. nov. and Picogryllus carentonensis Josse and Desutter-Grandcolas, gen. nov., sp. nov. are the oldest representatives of the Nemobiinae and Podoscirtinae subfamilies of the Trigonidiidae and Oecanthidae families respectively. P. carentonensis Josse and Desutter-Grandcolas, gen. nov., sp. nov. is also the smallest adult male with a full stridulatory apparatus ever documented in crickets (body length 3.3 mm), and the first taxon of the cricket clade for which male genitalia can be partly described. We discuss the significance of Cretaceous fossils of crickets for future evolutionary studies of this clade.

The Almogholagh–Dehgolan region is in the North Sanandaj–Sirjan zone of NW Iran. The granites of the region are metaluminous and display geochemical and textural characteristics of transitional granites between ferroan (A-type) and I-type granites. In geotectonic discrimination diagrams, the Almogholagh–Dehgolan granites plot mainly in the fields of within-plate granites and volcanic arc granites. With the exception of the Qalaylan granites, parts of other granites resemble A2-type granites. Granites of the Qalaylan intrusive body have petrographic and geochemical features close to I-type granites and are not A-type. Primary mantle and chondrite-normalized spider diagrams show enrichments in light rare earth elements relative to heavy rare earth elements. For an age of 150 Ma, the initial 87Sr/86Sr and 143Nd/144Nd ratios vary from 0.702769 to 0.706545 and from 0.512431 to 0.512558, respectively. Epsilon Nd values vary in a relatively limited range between −0.3 and +2.2, which corresponds to a mixed mantle–crustal source. On the basis of new geochemical and isotopic data, we suggest a geodynamic model involving partial melting of lower crustal rocks with the contribution of mantle magmas in a weakly extensional tectonic setting for the generation of the A-type granites of the region. The occurrence of ferroan (A-type) granites in this region of the Sanandaj–Sirjan zone indicates the existence of a partly extensional tectonic environment in a mainly compressional subduction-related regime in Late Jurassic time.

La Mina is one of three sites, along with Cueva Millán and La Ermita, located in the middle course of the Arlanza river. La Mina was excavated for the first time in 2006 and three test pits were carried out. In one of them, evidence of two Palaeolithic occupations was identified and several remains of woolly rhinoceros were recovered. Amino acid racemisation dating yielded an age of 52.5 ka BP, the earliest Upper Pleistocene date for Coelodonta antiquitatis on the Iberian Peninsula. This new record may have several implications for understanding the access routes to the Castilian Plateau, together with the definition of a new migratory wave of this species at the end of the Pleistocene. The location of La Mina on the Castilian Plateau may help researchers to complete the movements of this species through the Middle and Upper Palaeolithic on the Iberian Peninsula.

The Precambrian Southern Granulite Terrane (SGT) of south India is well-known for the preservation of high- to ultrahigh-temperature (HT-UHT) granulites, prominently exposed in its central part forming a linear belt referred to as the Kambam UHT belt. This belt also hosts widespread occurrences of mafic granulites that are observed in close spatial association with the HT-UHT granulites. This study presents detailed petrology, geochemistry and geochronology of representative mafic granulites from the area to understand their petrogenesis and tectonic setting. The results demonstrate that mafic granulites are low- to medium-K tholeiites, with continental arc affinity, formed by the partial melting of a subduction-modified enriched mantle source. The composition of the parent mantle source is modelled with a spinel/garnet lherzolite contribution ratio between 100/0 and 70/30, suggesting the mixing of spinel and garnet bearing melts during asthenosphere upwelling. Zircon U–Pb geochronology of mafic granulites constrains their emplacement between 612 Ma and 625 Ma, that subsequently underwent metamorphism between 581 Ma and 531 Ma. This overlaps with the timing of HT-UHT metamorphism in the Kambam UHT belt bracketed between 593 and 532 Ma. Zircon Hf isotopic studies reveal parent magma generation from reworked melting sources involving Archean and Proterozoic components. These results propose an alternative heat source for the formation of HT-UHT granulites in the Kambam UHT belt which can be designated as a major terrane boundary within the SGT.

We present U–Pb dates from peridotitic pyrope-rich garnet from four mantle xenoliths entrained in a kimberlite from Bultfontein, South Africa. Garnet dates magmatic emplacement due to the high mantle residence temperatures of the source material prior to eruption, which were most likely above the closure temperature for the pyrope U–Pb system. We determine a U–Pb date of 84.0 ± 8.1 Ma for the emplacement of the Bultfontein kimberlite from garnet in our four xenolith samples. The date reproduces previous dates obtained from other mineral-isotope systems (chiefly Rb–Sr in phlogopite). Garnet can be dated despite extremely low concentrations of U (median ∼0.05 μg/g), because concentrations of common Pb are often low or non-detectable. This means that sub-concordant garnets can be dated with moderate precision using very large laser-ablation spots (130 μm) measured by quadrupole inductively coupled plasma – mass spectrometry (LA-Q-ICP-MS). Our strategy demonstrates successful U–Pb dating of a U-poor mineral due to high initial ratios of U to common Pb in some grains, and the wide spread of isotopic compositions of grains on a concordia diagram. In addition, the analytical protocol is not complex and uses widely available analytical methods and strategies. This new methodology has some advantages and disadvantages for dating kimberlite emplacement versus established methods (U-based decay systems in perovskite and zircon, or Rb- or K-based systems in phlogopite). However, this method has unique promise for its potential application to detrital diamond prospecting and, more speculatively, to the dating of pyrope inclusions in diamond.

This paper reports geochemical characteristics of mafic gneisses and granulites collected from four localities in central Dronning Maud Land to evaluate the tectonic setting of their precursor rocks. Precursor rocks for the mafic gneisses and granulites in central Dronning Maud Land were formed in different geological backgrounds and tectonic settings. The mafic gneisses and granulites in the Schirmacher Hills were derived from basaltic rocks in a back-arc setting. Published U–Pb zircon ages and the geochemical variation of the basement rocks indicate their emplacement between 800 and 650 Ma, close to peak metamorphism. While similar protoliths were recognized in the inland nunataks of Hochlinfjellet, in Filchnerfjella the mafic gneisses/granulites were derived from basaltic rocks formed in marginal continental arcs or island arcs. Highly disturbed trace-element patterns indicate that the metamorphic process influenced the geochemical composition during the prograde metamorphic stage. Our results imply that the outcrops in central Dronning Maud Land with different metamorphic ages contain mafic gneisses/granulites from precursor rocks formed under different tectonic settings.

This paper focuses on one orogenic belt that formed during the Rinconada phase on the final stage of the Famatinian orogeny, between 445 and 410 Ma, which is well exposed at Sierra de Ramaditas and neighbouring ranges in western Argentina. The Ramaditas Complex is formed by metasedimentary and meta-ultrabasic rocks and amphibolites. This complex forms the upper nappe of a thrust stack resulting from westward thrusting. Deformation consists of an early high-temperature S1 foliation (stromatic migmatites), coeval with thrusting and metamorphism. Metamorphism attained peak P–T conditions of 6.0–6.9 kbar and 795–810 °C, at c. 440 Ma, i.e. coincident with the Rinconada orogenic phase. The lower unit and intermediate nappes crop out in the nearby sierras of Maz and Espinal and underwent low- to medium-grade Silurian metamorphism, respectively, together with the upper nappe, defining an inverted Barrovian-type metamorphism with T decreasing and P increasing downwards across the thrust stack (i.e. westward). We argue that the Rinconada orogenic phase developed near the continental margin of SW Gondwana, during a magmatic lull following accretion of the Precordillera terrane to the continental margin at c. 470 Ma. The active margin jumped to the west after accretion, and flat-slab subduction resumed in the early Silurian, provoking thrusting and imbrication of nappe stack under the still hot root (800–900 °C) of the older Famatinian magmatic arc. This ‘hot-iron’ process explains both the inverted Barrovian-type metamorphism and the missing overburden of 21 to 24 km implied by the P estimate.

Loose limestone blocks of a newly recognized hydrocarbon-seep deposit from the lower Oligocene Jansen Creek Member of the Makah Formation were collected on a beach terrace close to the mouth of Bullman Creek in Washington State, USA. The limestone consists largely of authigenic carbonate phases, including 13C-depleted fibrous cement forming banded and botryoidal crystal aggregates with δ13C values as low as –23.5 ‰. Lipids extracted from the limestone yielded molecular fossils of anaerobic methanotrophic archaea (ANME), dominated by compounds of an ANME-2/DSS consortium with δ13C values as low as −106 ‰, indicating formation at an ancient methane seep. The fossil inventory of the seep deposit is remarkable, consisting almost solely of echinoid remains, whereas typical seep biota are absent. Varying preservation of the echinoid fossils indicates parautochthonous deposition, corroborated by evidence for high fluid flow at the ancient seep, possibly responsible for displacement of echinoids after death. Although a full taxonomic description of the echinoids cannot be given, almost all fossils were assigned to one taxon of irregular spatangoids, except for a single regular echinoid. Abundance and lifestyle of the irregular spatangoids in the Bullman Creek echinoid seep deposit resemble those of the fossil Tithonia oxfordiana from an upper Jurassic seep deposit in France and extant Sarsiaster griegii from modern seeps in the Gulf of Mexico. The Bullman Creek echinoid deposit probably represents a fossil analogue of the Gulf of Mexico Sarsiaster mass occurrence, indicating that the adaptation of spatangoid echinoids to chemosynthesis-based ecosystems ranges back at least to the earliest Oligocene.

Methodologies that analyse the colouration and external appearance of extant species are very useful tools when facing one of the greatest challenges in the palaeoartistic reconstructions of extinct fauna: inferring the colour patterns. Earlier works have applied this methodology, for example, in the reconstruction of the Miocene bovid Tethytragus, proving that the maximum likelihood (ML) analysis to infer ancestral states has promising potential. This study offers a proposal for the reconstruction of the external appearance of Heteroprox moralesi Azanza 1989, an early cervid of the Middle Miocene present in several fossil sites of Central Spain. For the reconstruction of the external appearance, the colour patterns of all the extant species of the family Cervidae were studied with the method of ML analysis, as well as recent works about their phylogeny. The results show the most probable basal colour pattern of the cervids: dark shades on the limbs, dorsal section, and head of the animal, and, in contrast, lighter colours on the neck and perianal region. This basal pattern can be used as a basis for reconstructing colouration and to hypothesise about the external appearance of extinct taxa. Furthermore, the inferred forest habitat of H. moralesi has been taken into consideration in order to adjust the colour pattern, comparing the final results of the analysis performed in this study with that of the pattern observed in extant forest deer as well as with previous works employing this methodology.

Many studies have improved our understanding of the mode of preservation at the Crato fossil Lagerstätte. The high degree of preservation of the Crato mineralized insects is thought to be a consequence of the diffusion of ions through carcasses and envelopment by bacteria that, in turn, created microenvironmental conditions that led to mineralization, mainly pyritization. Pyritized insects have been oxidized by in situ weathering to more stable oxide/hydroxy minerals during Quaternary time. This transformation is essential to maintain the palaeontological information acquired during microbially induced pyritization in an oxidizing atmosphere. However, intense weathering can diminish or obscure the morphological fidelity, and little attention has been paid to the post-diagenetic processes experienced by these fossils. Here, we aim to determine the degree of alteration undergone by Crato pyritized insects using the following combination of analytical tools: scanning electron microscopy, energy dispersive X-ray spectroscopy, Fourier transform infrared and Raman spectroscopy. Our results show that well-preserved insects are preferentially replaced by haematite and poorly preserved fossils are replaced by goethite. In addition, we recorded three types of post-diagenetic alteration: insects with iron-oxide overgrowths; insects associated with black coatings, sometimes with the formation of dendrites; and insects preserved as an impression, where only the outline of the body remains. All of these alterations have the potential to distort or tarnish palaeontological information. Here, we measured the effects of such telodiagenetic alterations at macro and micro scales. Therefore, this taphonomic approach has wide applicability wherever fine-grained deposits bearing mineralized insects are found.

In the last decade, the identification of bone fragments by peptide mass fingerprinting or zooarchaeology by mass spectrometry is developing as a powerful tool in Quaternary palaeontology. The sequence of amino acids that make up the bone collagen molecule shows slight variations between taxa, which can be studied by mass spectrometry for taxonomic purposes. This requires reference databases that allow peptide identification. Although the cave bear (Ursus spelaeus Rosenmüller, 1794) is a common component in many European Pleistocene cave sites, no peptide fingerprint taxonomic study has paid special attention to this species up to now. For peptide markers in Ursidae, the most recent proposal is based on collagen obtained from a modern brown bear sample. In this work we attempt to cover this gap by studying bone collagen of cave and brown bear samples from different origins and different chronology, applying matrix-assisted laser desorption/ionisation time-of-flight mass spectrometry (MALDI TOF). We also performed an in-silico study of ursid bone collagen sequences published in databases. In our results we detected some discrepancies between the peptides obtained from both in silico and MALDI TOF analysis of fossil collagen and those published in the literature, in which we conclude that there are some misidentified peptides. The identification of skeletal remains by means of their peptide fingerprint is proving to be a powerful tool in palaeontology, which will bear greater fruit once the limitations of a technique that is in its initial stages have been overcome.

Materials collected on the territory of the southeastern White Sea area, including diversely preserved body imprints, combined body-trace fossils, specimens with signs of intravital damage and regeneration, and extended ontogenetic series, make it possible to significantly widen the data on the body plan and biology of Dickinsonia, the oldest known mobile animal, included in the Late Precambrian taxon of high rank, Proarticulata. A number of reconstructed anatomical features were added to the obvious directly observed features of Dickinsonia, such as a consistent body shape lacking lateral appendages and temporary outgrowths, transverse differentiation, and anterior–posterior polarity. These reconstructed features include dorsoventral polarity, ciliated mucus-secreting epithelium underlain by a basal lamina, two rows of blind food-gathering pockets, absence of a through-gut, nervous system of diffusive type, axial support band and muscle fibres. Such a set of features indicates the affinity of Dickinsonia and Proarticulata as a whole (the only known Ediacaran Metazoa) to Urbilateria, a hypothetical ancestor of bilaterally symmetrical animals.

Permian fishes and their isolated microremains are known from many localities in the Zechstein Basin. However, up to date the vertebrates have never been revealed in the southeasternmost part of this ancient sea. The new material consists of euselachian-type dermal denticles,?Listracanthus sp. dermal denticle,?Omanoselache sp. tooth, actinopterygian scales and actinopterygian teeth. Here, the detailed study of euselachian and actinopterygian remains, their stratigraphic distribution and geographical contexts is presented. Based on the qualitative analysis of teeth shapes several ecomorphotypes were described as well as the probable dietary preferences of fishes were reconstructed. These finds confirmed existence of small predators who fed on soft bodied prey as well as durophagous forms which were feeding on small shelly crustaceans or molluscs. The analysis of stratigraphic distribution of microremains, and their comparison with neighbouring sections revealed a spatially correlatable trend in increasing abundance of fishes in the more clayey parts of sections, interpreted to be positively associated with a sea level transgression event.