We report the first Hettangian theropod tracksite (~200 Ma) yielding a rich accumulation of plant remains from the Bleymard Strait (southern France). It constitutes an excellent opportunity to reconstruct lowermost Jurassic ecosystems hosting dinosaurs and which are still poorly documented in this area. Two morphotypes of tridactyl tracks are distinguished. They share similarities with Grallator and Kayentapus. Plant-bearing beds yield abundant leafy axes (Pagiophyllum peregrinum), male cones (Classostrobus sp.), wood (Brachyoxylon sp.) and pollen of conifers (Classopollis classoides). Sedimentological, petrological and mineralogical analyses demonstrated that, in the Dolomitic Formation from Bleymard, the palaeoenvironment progressively evolved from (1) a shoreface to a foreshore domain; to (2) a shallow environment that is restricted or occasionally open to the sea; then to (3) an intertidal to supratidal zone. The Hettangian theropod ecosystem of the Bleymard Strait was composed of tidal flats that were periodically emerged and bordered paralic environments inhabited by a littoral conifer-dominated forest in which Cheirolepidiaceae were the main component. The paucity of the palaeobotanical assemblage, as well as the xerophytic characteristics of Pagiophyllum, show that flora from Bleymard was adapted to withstand intense sunlight and coastal environments exposed to desiccant conditions coupled with salty sea spray, and dry conditions. These features are those of a conifer-dominated flora under a tropical to subtropical climate. The flora as well as the clay mineral analyses suggest contrasting seasons (cyclically dry then humid). This study supports that theropods were abundant and particularly adapted to this type of littoral environment bordering Cheirolepidiaceae-dominated forests.

A recent excavation yielded 118 large tridactyl footprints in the Lower Jurassic Dolomitic Formation of the Causses Basin, at Mongisty in southern France. Most of the tracks are ascribed to Eubrontes giganteus Hitchcock, 1845. They are preserved on a surface of 53 m2 and form parallel rows with a preferential orientation towards the north. Such an abundance and density of E. giganteus is observed for the first time in the Early Jurassic from the Causses Basin. Sedimentological and ichnotaphonomical analyses show that the footprints were made at different time intervals, thus excluding the passage of a large group. In contrast to all other tracksites from the Dolomitic Formation, where tracks are preserved in fine-grained sediments corresponding to low-energy depositional palaeoenvironments, the tracks from Mongisty are preserved in coarse-grained sediment which is a matrix- to clast-supported breccia. Clasts consist of angular to sub-rounded, millimetric to centimetric-scale (up to 2 cm), poorly sorted, randomly oriented, homogeneous dolostone intraclasts floating in a dolomudstone matrix. Sedimentological analysis shows that the depositional environments of Mongisty varied from subtidal to intertidal/supratidal settings in a large and protected flat marsh. The lithology of the track-bearing surfaces indicates that the mudflat of the Causses Basin was sporadically affected by large mud flows that reworked and redeposited mudstone intraclasts coming from the erosion of upstream, dry and partially lithified mud beds. Throughout the world, this type of preservation of dinosaur tracks in tidal matrix- to clast-supported breccias remains rare.