Studies were conducted from 1989 to 1993 in continuous no-tillage and moldboard plow corn fields to describe rates of velvetleaf seed predation with time and with seed density, and to identify principal seed predators. Rates of seed loss from the soil surface averaged 1 to 57% day−1 and were equivalent in the two tillage systems. Predator populations were the same in no-tillage and moldboard plow fields. The predation rate was generally low in winter months, increased in mid-summer, and declined in late summer. In 2 of the 4 yr, predation increased in October and November. The predation rate was described by an exponential decay function of seed density, with high rates of seed loss at low densities and leveling off to a nearly constant level at densities above 600 seeds m−2. Predation was highest where seed access was not restricted, and exclosures of 6.5 and 1.6 cm2 reduced predation up to 15 and 52%, respectively. Mice were important predators in the field. In laboratory feeding studies, the carabid beetle Amara cupreolata, the slugs Arion subfuscus and Deroceras reticulatum, and cutworms (Agrotis ipsilon) consumed imbibed velvetleaf seeds. Amara cupreolata and A. subfuscus were the only predators to damage unimbibed velvetleaf seeds.

Understanding patterns of weed seedling emergence within a growing season and over years is important to develop models to predict optimum timing of weed management practices. A study was conducted in a field with no previous velvetleaf infestations to describe emergence patterns following seed burial at three depths in two tillage systems. Freshly harvested velvetleaf seeds were planted 0, 2, and 6 cm deep in moldboard plowed (MP) and no-tillage (NT) corn stubble in October 1990. Velvetleaf seedling emergence was monitored over the following 4 yr in continuous corn. Emergence was higher in NT than in MP plots throughout 4 yr of observation. The first growing season following seeding, emerged seedlings represented 9.3 to 15.8% of the seeds sown in NT, compared with 0.1 to 0.8% of seeds sown in MP. After four growing seasons, emerged seedlings were 12.5 to 25% of seeds sown in NT but only 6 to 7.4 % of seeds sown in MP. Emergence was consistently higher from the 0-cm depth than from the 6-cm depth in NT, but seeding depth did not influence emergence in MP due to mixing of the soil during tillage. Velvetleaf emergence was related to growing degree days (base 7.5 C), with greater consistency in NT than in MP. Averaged over years and planting depths, 50% velvetleaf emergence occurred within 8 and 13 d of the predicted date in NT and MP systems, respectively.