The Arda River marine succession (Italy) is an excellent site to apply an integrated approach to paleoenvironmental reconstructions, combining the results of sedimentology, body fossil paleontology, and ichnology to unravel the sedimentary evolution of a complex marine setting in the frame of early Pleistocene climate change and tectonic activity. The succession represents a subaqueous extension of a fluvial system, originated during phases of advance of fan deltas affected by high-density flows triggered by river floods, and overlain by continental conglomerates, indicating a relative sea level fall and the establishment of a continental environment. An overall regressive trend is observed through the section, from prodelta to delta front and intertidal settings. The hydrodynamic energy and the sedimentation rate are not constant through the section, but they are influenced by hyperpycnal flows, whose sediments were mainly supplied by an increase in Apennine uplift and erosion, especially after 1.80 Ma. The Arda section documents the same evolutionary history of coeval successions in the Paleo-Adriatic region, as well as the climatic changes of the early Pleistocene. The different approaches used complement quite well one another, giving strength and robustness to the obtained results.

The hypothesis that tetrapods evolved from elpistostegids during the Frasnian, in a predominantly aquatic context, has been challenged by the discovery of Middle Devonian tetrapod trackways predating the earliest body fossils of both elpistostegids and tetrapods. Here I present a new hypothesis based on an overview of the trace fossil and body fossil evidence. The trace fossils demonstrate that tetrapods were capable of performing subaerial lateral sequence walks before the end of the Middle Devonian. The derived morphological characters of elpistostegids and Devonian tetrapods are related to substrate locomotion, weight support and aerial vision, and thus to terrestrial competence, but the retention of lateral-line canals, gills and fin rays shows that they remained closely tied to the water. Elpistostegids and tetrapods both evolved no later than the beginning of the Middle Devonian. The earliest tetrapod records come from inland river basins, sabkha plains and ephemeral coastal lakes that preserve few, if any, body fossils; contemporary elpistostegids occur in deltas and the lower reaches of permanent rivers where body fossils are preserved. During the Frasnian, elpistostegids disappear and these riverine-deltaic environments are colonised by tetrapods. This replacement has, in the past, been misinterpreted as the origin of tetrapods.