The concepts of pollen source area and of production and dispersal biases in pollen representation are quantified by means of a simple theoretical model. Source areas and relative pollen representation are shown to depend on basin size according to functions that describe the amount of pollen remaining airborne at increasing distances from single pollen sources. The form of these functions is determined by physical processes. Standard formulas for elevated sources do not apply, but the integrated form of Sutton's equation for particle dispersal from a ground-level source gives useful approximations applicable to pollen transport over a forest canopy. Simulations using this equation yielded source areas that increased realistically with basin size, showed substantial differences between source areas for pollen grains with different deposition velocities, and predicted that lighter pollen grains should become better represented with increasing basin size. All of these predictions are qualitatively consistent with present knowledge of the characteristics of pollen assemblages in different depositional environments. The model further allows parameters that can be estimated by statistical calibration methods to be predicted from underlying physical quantities. This extension suggests procedures for testing the theory with quantitative data on surface pollen and forest composition. Preliminary results showed reasonable agreement between estimated and predicted values of dispersal indices for the most abundant taxa in pollen spectra from the northern midwestern United States.

Dissimilarity coefficients measure the difference between multivariate samples and provide a quantitative aid to the identification of modern analogs for fossil pollen samples. How eight coefficients responded to differences among modern pollen samples from eastern North America was tested. These coefficients represent three different classes: (1) unweighted coefficients that are most strongly influenced by large-valued pollen types, (2) equal-weight coefficients that weight all pollen types equally but can be too sensitive to variations among rare types, and (3) signal-to-noise coefficients that are intermediate in their weighting of pollen types. The studies with modern pollen allowed definition of critical values for each coefficient, which, when not exceeded, indicate that two pollen samples originate from the same vegetation region. Dissimilarity coefficients were used to compare modern and fossil pollen samples, and modern samples so similar to fossil samples were found that most of three late Quaternary pollen diagrams could be “reconstructed” by substituting modern samples for fossil samples. When the coefficients indicated that the fossil spectra had no modern analogs, then the reconstructed diagrams did not match all aspects of the originals. No modern analogs existed for samples from before 9300 yr B.P. at Kirchner Marsh, Minnesota, and from before 11,000 yr B.P. at Wintergreen Lake, Michigan, but modern analogs existed for almost all Holocene samples from these two sites and Brandreth Bog, New York.

Field evidence from the Atacama Desert of northern Chile indicates that the stratigraphic position of the land snail Bostryx variabilis Herm has been incorrectly assigned to a marine series of the lower Pleistocene. Isotopic dating proves that the snail is a recently extinct Holocene taxon requiring no special paleoenvironmental explanations for its presence. However, the snail appears to offer an example of stress-induced speciation in confirmation of the “punctuated equilibrium” theory of evolution. It was probably transported from a fog oasis mountain habitat to a lower, hyperarid hostile environment by a sudden mudflow.

Eight box cores from the tropical Atlantic were studied in detail with regard to foraminiferal oxygen isotopes, radiocarbon, and Globorotalia menardii abundance. A standard Atlantic oxygen-isotope signal was reconstructed for the last 20,000 yr. It is quite similar to the west-equatorial Pacific signal published previously. Deglaciation is seen to occur in two steps which are separated by a pause. Onset of deglaciation is after 15,000 yr B.P. The pause is centered between 11,000 and 12,000 yr B.P., but may be correlative with the Younger Dryas (10,500 yr B.P.) if allowance is made for a scale shift due to mixing processes on the sea floor. Step 2 is centered near 10,000 yr B.P. and is followed by a brief excursion toward light oxygen values. This excursion (the M event) may correlate with the Gulf of Mexico meltwater spike.

The well known Clovis and Plainview archaeological sites of New Mexico and Texas have yielded new data on regional late Quaternary geologic, paleoclimatic, and pedologic histories. Eolian sedimentation at the Clovis site from about 10,000 to less than 8500 yr B.P. was followed by the formation of a cumulic soil between 8500 and 5000 yr B.P. Episodic eolian and slope wash deposition then culminated in massive eolian sedimentation about 5000 yr B.P. after which a Haplustalf formed then was subsequently buried by part of a dune system within the last 1000 yr. At the Plainview site, a basal stream gravel contains Plainview cultural material (ca. 10,000 yr B.P.), which is followed by a localized early Holocene lacustrine deposit, two eolian deposits (the younger dating to about 5000 yr B.P.), and a marsh deposit which slowly accreted as an Argiustoll formed in the younger eolian unit. The data indicate that on the Southern High Plains (1) between 12,000 and 8500 yr B.P. sedimentation varied from site to site, (2) there was a regional climate change toward warming and drying in the early Holocene, (3) two episodes of severe drought apparently occurred in the middle Holocene (6500 to 4500 yr B.P.), (4) between 4500 yr B.P. and the present an essentially modern climate existed, but with several shifts toward aridity within the last 1000 yr, (5) argillic horizons have developed in late Holocene soils, (6) clay illuviation can occur in calcareous soils, and (7) long-distance correlation of Holocene stratigraphy in the region is possible, particularly with the aid of soil morphology.

A 33,000-yr pollen record from Carp Lake provides information on the vegetation history of the forest/steppe border in the southwestern Columbia Basin. The site is located in the Pinus ponderosa Zone but through much of late Quaternary time the area was probably treeless. Pollen assemblages in sediments dating from 33,000 to 23,500 yr B.P. suggest a period of temperate climate and steppe coinciding with the end of the Olympia Interglaciation. The Fraser Glaciation (ca. 25,000–10,000 yr B.P.) was a period of periglacial steppe or tundra vegetation and conditions too dry and cold to support forests at low altitudes. Aridity is also inferred from the low level of the lake between 21,000 and 8500 yr B.P., and especially after about 13,500 yr B.P. About 10,000 yr B.P. Chenopodiineae and other temperate taxa spread locally, providing palynological evidence for a shift from cold, dry to warm, dry conditions. Pine woodland developed at the site with the onset of humid conditions at 8500 yr B.P.; further cooling is suggested at 4000 yr B.P., when Pseudotsuga and Abies were established locally.

During the Quaternary glacial episodes, when sea level was considerably lower, Asia and North America were linked by large extensions of circumarctic land (Beringia), which remained unglaciated. This land mass served not only as a biogeographical bridge for plants, animals, and humans, but also supported a biome very different from present tundra or boreal coniferous forests, which was dominated by steppes and a rich mammalian megafauna. Carbon stable isotope ratios of Beringian late Pleistocene mummified remains of bison, equids, mammoth, caribou, musk-ox, moose, woolly rhino, and other undetermined species, found preserved in permafrost, indicate that these megaherbivores fed exclusively on C3 plants, and that C4 grasses were not differentially ingested by bison, as previously suggested. Paleoclimatic constraints probably prevented the formation of a warm-season (C4) guild during the later part of the growing season in the steppes of Beringia during the last glaciation.

Tephra in 31 piston cores from the western Gulf of Mexico and 7 piston cores from the equatorial Pacific were analyzed by electron microprobe. Six ash layers in the western Gulf of Mexico were easily distinguished by TiO2, FeO, and CaO contents and correlated by geochemistry in order to determine the distribution pattern for each ash layer. Correlation by geochemistry is an easier, more accurate method than biostratigraphic correlation; some of the tephras were miscorrelated by biostratigraphy. The six tephras were dated by geochemical identification in a piston core with oxygen-isotope stratigraphy and the ages are Y5 (30,000 yr B.P.), Y6 (65,000 yr B.P.), Y8 (84,000 yr B.P.), X2 (110,000 yr B.P.), W1 (136,000 yr B.P.), and W2 (185,000 yr B.P.). Data from this study corroborated correlations of the Y8 tephra in the western Gulf of Mexico with the D layer in the eastern equatorial Pacific Ocean. None of the other five layers in the Gulf of Mexico, however, were found in the Pacific Ocean. The limited distribution of the Y5, Y6, X2, and W2 ash layers close to Mexico indicates possible sources in Mexico. Tephra from the late Pleistocene La Primavera pumice in Mexico, however, does not correlate with the marine tephra.

This study tests the hypothesis that the number and distribution of some native American languages may be related to ice-margin changes of the Wisconsin glaciation. The analysis indicated that the number of languages per unit area is much greater in unglaciated areas of the last glacial maximum than in glaciated areas. The pattern of languge overlap between land areas sequentially exposed during deglaciation appears to indicate the direction of movement of populations from the periphery toward the core of the area once covered by the Wisconsin Ice Sheet. The data strongly indicate that North America was inhabited prior to the Wisconsin glacial maximum, because glacial maximum conditions apparently influenced linguistic distributions. Evidence suggests that ancestral Eskimo-Aleut and Na-Dene speakers occupied the northwestern edge of the continental ice mass, and that ancestral Algonquian speakers were south of the ice mass during the Wisconsin glacial maximum (approximately 18,000 yr ago). These three linguistic groups were the principal ones to spreas into areas exposed by the recession of the Wisconsin ice.