Is Sinhala caste simply a weak regional variant of Hindu caste or is it something else entirely? This essay argues that Sinhala caste as found in the territory of the former Kandyan Kingdom has had a distinctive ontology and retains its unique character. The essay begins with an overview of textual, genetic, and archaeological evidence for the origins of caste on the subcontinent. It then turns to the island and the fourth century CE bifurcation of Sinhala society into “high” and “low”; this duality’s persistence into the second millennium CE; its elaboration in the Kandyan Kingdom’s bureaucratic political economy; and the dissonance between this Sinhala “cartwheel” model of collective inequality and the Brahmanical “ladder” of colonial powers and the Sinhala elite. The essay concludes by examining how the ongoing discordance between these two models of Sinhala caste plays out in people’s lives through a case study of a non-elite caste community.