We introduce a modified Galton‒Watson process using the framework of an infinite system of particles labelled by (x,t), where x is the rank of the particle born at time t. The key assumption concerning the offspring numbers of different particles is that they are independent, but their distributions may depend on the particle label (x,t). For the associated system of coupled monotone Markov chains, we address the issue of pathwise duality elucidated by a remarkable graphical representation in which the trajectories of the primary Markov chains and their duals coalesce to form forest graphs on a two-dimensional grid.

In the paper we present a phenomenon occurring in population processes that start near 0 and have large carrying capacity. By the classical result of Kurtz (1970), such processes, normalized by the carrying capacity, converge on finite intervals to the solutions of ordinary differential equations, also known as the fluid limit. When the initial population is small relative to the carrying capacity, this limit is trivial. Here we show that, viewed at suitably chosen times increasing to ∞, the process converges to the fluid limit, governed by the same dynamics, but with a random initial condition. This random initial condition is related to the martingale limit of an associated linear birth-and-death process.