Phyllosilicates are hypothesized to be primarily of pedogenic origin in shallowly buried paleosols (≤3 km depth), regardless of the age of the paleosol. To test this hypothesis, this work evaluates the possible presence of detrital and diagenetic phyllosilicates in middle and upper Pennsylvanian paleosols, collected from three drill cores along a north–south transect in the Illinois Basin. The abundances of 2M1 muscovite, quartz, and K-feldspar are greater in a morphologically immature Protosol from the southernmost core; 1Md illite and interstratified illite-smectite with R1 and R0 stacking orders are more abundant in the more mature Vertisols of the central and northern cores. K-Ar age values of multiple clay-size fractions from each paleosol averaged ~260 Ma in the northern core, 270 Ma in the central core, and 295 Ma in the southern core. While considering the complex tectonic and thermal history of the Illinois Basin, detrital minerals are more abundant in immature paleosols that experienced relatively greater maximum burial depths and thus greater sediment supply whereas illitization in more mature paleosols was probably initiated primarily during protracted burial diagenesis. As the present study found evidence for diagenetic and detrital minerals in clay-size fractions of shallowly buried, deep-time paleosols, caution is advised when using paleosol minerals for ancient climate and environment reconstructions.

Restudy of Campyloprion annectans Eastman, 1902 from North America demonstrated that neither specimen included is diagnostic at the species level; thus, the species name is a nomen dubium. Since this species was designated as the type species of the genus, this requires suppression of the generic name also. Another species earlier assigned to Campyloprion, Campyloprion ivanovi Karpinsky, 1924 is used as a type for a newly established genus Karpinskiprion Lebedev et Itano gen. nov. The composition of the family Helicoprionidae Karpinsky, 1911 is reviewed, and a new family Helicampodontidae Itano et Lebedev fam. nov. is erected. A new specimen of Karpinskiprion ivanovi (Karpinsky, 1924) recently discovered in the Volgograd Region of Russia is the most complete Karpinskiprion specimen ever found. It unambiguously demonstrates the coiled nature of these tooth whorls and presents information on their developmental stages. During organogeny, cutting blades of the crown became reshaped, and basal spurs progressively elongated, forming a grater. Whorl growth occurred by addition of new crowns to the earlier mineralised base followed by later spur growth. In contrast to consistently uniform cutting blades, spurs are often malformed and bear traces of growth interruption. Both sides of the outer coil of the tooth whorl bear lifetime wear facets. The youngest (lingual) crowns are as yet unaffected by wear. The best-preserved facets show parallel radially directed scratch marks. The upper jaw dentition of Karpinskiprion is unknown, but we suggest that the faceted areas resulted from interaction with the antagonistic dental structures here. Three possible hypotheses for this interaction are suggested: (a) two opposing whorls acted as scissor blades, moving alternately from one side to another; (b) the lower tooth whorl fitted between paired parasymphyseal tooth whorls of the opposing jaw; or (c) the lower tooth whorl fitted into a dental pavement in the upper jaw.

During the late Palaeozoic Era, a series of related marine strata dominated by multi-layer limestones were deposited in the southern North China Craton. In order to gain new insights into the systematic geochemistry of the carbonate succession of the representative formation (Taiyuan Formation), we examined 59 limestone samples collected from the Huaibei Coal Basin (HCB), with a view towards quantitatively determining the major and trace elements and stable isotope compositions. The data obtained can provide essential evidence for reconstruction of the depositional palaeo-environment and tectonic setting of the Taiyuan Formation. Both X-ray diffraction analyses and palaeoredox proxies (e.g. V/Cr, V/(V + Ni) and authigenic U) indicated that the limestone layers were deposited in an oxic–dysoxic zone, with calcite as the main component. Moreover, palaeomagnetic evidence provided support for the conclusion that these limestones were laid down within an epicontinental sea depositional environment under a warm or hot palaeoclimate during the transition between late Carboniferous and early Permian time. Additionally, evidence obtained from our analyses of trace and rare earth elements revealed that the tectonic setting of the Taiyuan Formation (L1–L5) in the HCB transited from an open ocean to a passive continental margin, thereby indicating that this transformation stemmed from the subduction closure of the northeastern Palaeotethys Ocean. The findings of this study would be of interest to those working on the upper Palaeozoic marine strata in the southern North China Craton.

The Carboniferous lungfish genus Sagenodus is reviewed from all available British specimens and described in detail for the first time. We identify two species exclusive to the UK: Sagenodusinaequalis, the type species, deriving from the late Carboniferous (=Pennsylvanian); and Sagenodus quinquecostatus derived from the early Carboniferous (=Mississippian). The genus is probably the most widespread of the known Carboniferous lungfish genera, but the British species have not been formally described since their discovery in the mid–late 19th Century. This work will provide data to help resolve existing questions about the position of Sagenodus in the phylogeny of Palaeozoic lungfishes, and provide a template for the recognition of isolated elements in museum collections and the finds from recent and future field work. The early Carboniferous species, S. quinquecostatus, shows a so far unique functional mechanism in which the lower tooth plates appear to rotate relative to the upper plates during jaw closure, implying a kinetic function at the symphysis or jaw joint.

Specimens of trematopid amphibians from the Asturian (Upper Carboniferous) of Nýřany, Czech Republic, are redescribed as two taxa, namely Mordex calliprepes Steen and Mattauschia (gen. nov) laticeps Fritsch. Mordex calliprepes is represented by a single post-metamorphic specimen and has the diagnostic trematopid characters of the nasal region. Mattauschia laticeps is represented by one adult partial skull and mandible plus some fragments and two small post-metamorphic specimens including the species name-bearer. It has the trematopid-type modified lacrimal and a large but oval naris and appears to be the most primitive trematopid yet described. The stratigraphically sequential large trematopids Mattauschia, Fedexia, Ecolsonia and Acheloma show progressive acquisition of the derived features that characterise the terminal form Acheloma.

Mordex has a combination of primitive and derived characters and its position within the family is less clear. The many ‘branchiosaurs' in the Nýřany assemblage include specimens that could be larvae of both Mordex and Mattauschia but certain attribution is not possible and they are assigned to Olsoniformes incertae sedis. Mordex and Mattauschia appear to be terrestrial exotic elements in the Nýřany tetrapod assemblage, but with possible larvae in the lake assemblage. Representatives of at least four Palaeozoic dissorophoid families were present in late Middle Pennsylvanian/Asturian strata implying diversification of the Dissorophoidea prior to this time.

Silicified beyrichiocopid and podocopid ostracods from limestone nodules derived from the middle part of the Ichinotani Formation within the Hida Gaien Terrane of central Honshu Island, Japan, are associated with fusulinid foraminifera that indicate strata of the middle Moscovian (Pennsylvanian, Carboniferous). This is a rare record of ostracods from the Palaeozoic of Japan and the first systematic description of ostracods from the Carboniferous of the Hida Gaien Terrane. The fauna comprises six ostracod species (two new) assigned to the genera Amphissites, Kirkbya, Bairdia, Aechmina and Healdia, and additional material of possible cavellinids. The numerical dominance of ornamented beyrichiocopids such as Kirkbya and Amphissites, along with smaller numbers of smooth podocopids such as Bairdia, indicates an ‘Eifelian mega-assemblage’ ecotype (sensu G. Becker), that is typical of mid Palaeozoic shallow marine, high-energy environments in a fore-reef ecosystem.

Vegetation diversity and pattern changes, and their relation to tectono-sedimentary histories are compared between selected Euramerican Late Palaeozoic coalfields, to understand better the controls on the dynamics of the Pennsylvanian terrestrial ecosystems and to demonstrate the problems with comparing data from various basins. The analysis is based on data from the following basins of different geotectonic and palaeogeographical positions: the cratonic Pennines Basin, the foreland South Wales and Upper Silesia basins, and the fault-related Intra Sudetic and Central and Western Bohemia basins. The analysis indicates that complex factors are responsible for changes in plant diversity and vegetation patterns. These are related to climate, tectonics, preservation potential, sampling biases and the current state of revision of the flora in each basin. Plant diversity patterns in the basins differ because of local controls and/or the character and detail of the available data. Maximum diversity varies among the basins within the Langsettian and Duckmantian substages. Two apparent step-like drops in diversity were detected within coal-bearing strata of most basins: at the Duckmantian/Bolsovian boundary and at the Bolsovian/Asturian boundary. Further and more prominent falls are related to transitions from coal-bearing to non-coal-bearing (mostly red bed) strata or vice versa during Stephanian times. Interpretation of climatic signals recorded in the sedimentary successions indicates that Westphalian and middle Stephanian times were wet intervals, whereas early and late Stephanian times were drier.

The fossil ricinuleids are described, and their systematic relationships revised, based on a study of all British and American specimens. Phylogenetic analysis, utilising a new character set, indicates a probable basal dichotomy between the fossil and extant forms. The order is divided into two new suborders: Neoricinulei subordo, nov., for the extant genera, and Palaeoricinulei subordo. nov., for the fossil forms. Two new genera are erected: Amarixys gen. nov. for Curculioides gracilis (Petrunkevitch, 1945) and A. stellaris sp. nov., and Terpsicroton gen. nov. for Poliochera alticeps Pocock, 1911. Three new species are erected in the genus Curculioides: C. mcluckiei sp. nov., C. pococki sp. nov., and C. gigas sp. nov., and one in the genus Poliochera: P. gibbsi, sp. nov.